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<mods:title id="5E584EE97DBE528BC8204D348B89A78F">Review of Rhinotora (Diptera, Heleomyzidae) with description of a new species and key to all known heleomyzid species from Brazil</mods:title>
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<mods:namePart id="4A09F514D59C298B70C2DCC58208365B">Lima, Felipe Varussa De Oliveira</mods:namePart>
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Key to the known species of
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occurring in
<collectingCountry id="ADFA9ADFFFADFF80247BFD4A2BB4FD49" box="[849,925,733,759]" name="Brazil" pageId="17" pageNumber="530">Brazil</collectingCountry>
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1. Face without subantennal pits (
<figureCitation id="4DD6C6CAFFADFF802539FCB42A66FC87" box="[531,591,803,825]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
C, 7F, 8F). Compound eyes large and prominent. Vertex slightly to deeply excavated. Head brown to yellow.......................................................................Rhinotorini... 2
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</paragraph>
<paragraph id="D552DA4FFFADFF8027BDFCC92DB2FC2F" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFCC92DB2FC2F" pageId="17" pageNumber="530">
1. Face with pair of deep, oval subantennal pits (
<figureCitation id="4DD6C6CAFFADFF8025B3FCC92AC1FCCA" box="[665,744,862,884]" captionStart="FIGURE 9. A C" captionStartId="16.[151,250,1637,1659]" captionTargetBox="[151,1436,181,1614]" captionTargetId="figure@16.[151,1436,181,1615]" captionTargetPageId="16" captionText="FIGURE 9. A C, Diacia diadema (♂ holotype, NMW). A, dorsal habitus; B, ventral habitus; C, head, frontal view (arrow indicates subantennal pit). D F, Dichromyia sanguiniceps (♂, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view (arrow indicates subantennal pit)." httpUri="https://zenodo.org/record/259184/files/figure.png" pageId="17" pageNumber="530">Figs 9C</figureCitation>
,F). Compound eyes reduced. Vertex horizontal or slightly convex. Head at least partially bright orange.................................................................. Diaciini...13
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2(1). Postvertical setae absent (
<figureCitation id="4DD6C6CAFFADFF8026F2FC0F2A24FC10" box="[472,525,920,942]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Fig. 1</figureCitation>
C). Scutellum with longitudinal excavation or with protuberances (
<figureCitation id="4DD6C6CAFFADFF8023A5FC0F2CE3FC10" box="[1167,1226,920,942]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
B, 2B). Vertex deeply excavated (
<figureCitation id="4DD6C6CAFFADFF802667FC2129AEFC72" box="[333,391,950,972]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
C, 7F)................................................................................ 3
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</paragraph>
<paragraph id="D552DA4FFFADFF8027BDFC442DB5FBB8" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFC442DB5FBB8" pageId="17" pageNumber="530">
2. Postvertical setae weak, convergent (
<figureCitation id="4DD6C6CAFFADFF802563FC442AAAFC57" box="[585,643,979,1001]" captionStart="FIGURE 8. A C" captionStartId="15.[151,250,1870,1892]" captionTargetBox="[151,1436,414,1848]" captionTargetId="figure@15.[151,1436,414,1848]" captionTargetPageId="15" captionText="FIGURE 8. A C, Neorhinotora mutica (♂, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view. D F, Rhinotoroides bifurcata (♂, INPA). D, lateral habitus (arrow indicates postocellar setae); E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259183/files/figure.png" pageId="17" pageNumber="530">Fig. 8</figureCitation>
D). Scutellum without longitudinal excavation or protuberances (
<figureCitation id="4DD6C6CAFFADFF8023D5FC442D10FC57" box="[1279,1337,979,1001]" captionStart="FIGURE 8. A C" captionStartId="15.[151,250,1870,1892]" captionTargetBox="[151,1436,414,1848]" captionTargetId="figure@15.[151,1436,414,1848]" captionTargetPageId="15" captionText="FIGURE 8. A C, Neorhinotora mutica (♂, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view. D F, Rhinotoroides bifurcata (♂, INPA). D, lateral habitus (arrow indicates postocellar setae); E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259183/files/figure.png" pageId="17" pageNumber="530">Fig. 8</figureCitation>
E). Vertex slightly excavated (
<figureCitation id="4DD6C6CAFFADFF8026B6FC6729FAFBB8" box="[412,467,1008,1030]" captionStart="FIGURE 8. A C" captionStartId="15.[151,250,1870,1892]" captionTargetBox="[151,1436,414,1848]" captionTargetId="figure@15.[151,1436,414,1848]" captionTargetPageId="15" captionText="FIGURE 8. A C, Neorhinotora mutica (♂, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view. D F, Rhinotoroides bifurcata (♂, INPA). D, lateral habitus (arrow indicates postocellar setae); E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259183/files/figure.png" pageId="17" pageNumber="530">Fig. 8</figureCitation>
F)................................................
<taxonomicName id="12EDA1CCFFADFF80231EFC672DB0FBB8" ID-CoL="78TC2" authority="Lopes, 1934" authorityName="Lopes" authorityYear="1934" box="[1076,1433,1008,1030]" class="Insecta" family="Heleomyzidae" genus="Rhinotoroides" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis id="E799065DFFADFF80231EFC672D31FBB8" box="[1076,1304,1008,1030]" italics="true" pageId="17" pageNumber="530">Rhinotoroides bifurcata</emphasis>
<bibRefCitation id="B17CA7BEFFADFF802235FC662DB0FBB8" author="Lopes" box="[1311,1433,1008,1030]" pageId="17" pageNumber="530" refString="Lopes, H. S. (1934) Nota previa sobre alguns Rhopalomeridae (Dipt.). Revista de Entomologia, 4, 517." type="journal article" year="1934">Lopes, 1934</bibRefCitation>
</taxonomicName>
.
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3(2). Scutellum flattened, not excavated, with spiniform or lobed protuberances (
<figureCitation id="4DD6C6CAFFADFF802499FB992BC7FB9A" box="[947,1006,1038,1060]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
B, 2B). Pruinescence of scutum fine, present around setal sockets (
<figureCitation id="4DD6C6CAFFADFF802684FBBC29C1FBFF" box="[430,488,1067,1089]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
B,D)...............................................................
<emphasis id="E799065DFFADFF80223CFBBC2DA3FBFF" box="[1302,1418,1067,1089]" italics="true" pageId="17" pageNumber="530">
<taxonomicName id="12EDA1CCFFADFF80223CFBBC2D53FBFF" ID-CoL="78TT" box="[1302,1402,1067,1089]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="genus">Rhinotora</taxonomicName>
...
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4
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3. Scutellum inflated and longitudinally excavated (
<figureCitation id="4DD6C6CAFFADFF8025EFFBDF2B3DFBE0" box="[709,788,1096,1118]" captionStart="FIGURE 7. A C" captionStartId="14.[151,250,1649,1671]" captionTargetBox="[151,1436,193,1626]" captionTargetId="figure@14.[151,1436,193,1627]" captionTargetPageId="14" captionText="FIGURE 7. A C, Neorhinotora amapaensis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, wing. D F, Neorhinotora aristalis (♂ holotype, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259182/files/figure.png" pageId="17" pageNumber="530">Fig. 7B</figureCitation>
). Pruinescence of scutum thick, absent around setal sockets (
<figureCitation id="4DD6C6CAFFADFF80225BFBDF2934FBC2" captionStart="FIGURE 7. A C" captionStartId="14.[151,250,1649,1671]" captionTargetBox="[151,1436,193,1626]" captionTargetId="figure@14.[151,1436,193,1627]" captionTargetPageId="14" captionText="FIGURE 7. A C, Neorhinotora amapaensis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, wing. D F, Neorhinotora aristalis (♂ holotype, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259182/files/figure.png" pageId="17" pageNumber="530">Figs 7AB</figureCitation>
,DE).............................................................................
<taxonomicName id="12EDA1CCFFADFF8023C1FBF12D44FBC2" ID-CoL="62JB" box="[1259,1389,1126,1148]" class="Insecta" family="Heleomyzidae" genus="Neorhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="E799065DFFADFF8023C1FBF12D44FBC2" box="[1259,1389,1126,1148]" italics="true" pageId="17" pageNumber="530">Neorhinotora</emphasis>
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...11
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4(3). Scutellum with apical and lateral spiniform protuberances (
<figureCitation id="4DD6C6CAFFADFF80243CFB142B64FB27" box="[790,845,1155,1177]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Fig. 1</figureCitation>
B). Wing membrane hyaline with dark spots; cell r1 with several supernumerary crossveins, occasionally some of them branched (
<figureCitation id="4DD6C6CAFFADFF802451FB372B9DFB08" box="[891,948,1184,1206]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
D, 3D, 6D).............................. 5
</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDFB292DBCFB4F" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFB292DBCFB4F" pageId="17" pageNumber="530">
4. Scutellum with discrete apical and lateral lobed protuberances (
<figureCitation id="4DD6C6CAFFADFF80246BFB292BA5FB6A" box="[833,908,1214,1236]" captionStart="FIGURE 2. A B" captionStartId="6.[151,250,1771,1793]" captionTargetBox="[151,1436,329,1747]" captionTargetId="figure@6.[151,1436,329,1749]" captionTargetPageId="6" captionText="FIGURE 2. A B, Rhinotora fonsecai (♀ paratype, MNRJ). A, lateral habitus; B, scutellum. C D, R. lopesi (♂ holotype, MZUSP); C, lateral habitus; D, dorsal habitus (arrow indicates parallel posteromedial spots on scutum). E F, R. lopesi (♂, INPA). E, protandrium; F, male genitalia, ventral view (arrows indicate greater and smaller lobes of cercus)." httpUri="https://zenodo.org/record/259177/files/figure.png" pageId="17" pageNumber="530">Fig. 2B</figureCitation>
). Wing membrane dark with hyaline spots; cell r1 with several supernumerary crossveins, most of them branched (per
<bibRefCitation id="B17CA7BEFFADFF802415FB4C2C67FB4F" author="Almeida" box="[831,1102,1243,1265]" pageId="17" pageNumber="530" refString="Almeida, J. C. &amp; Ale-Rocha, R. (2011) Comparative morphology of the male terminalia of the subtribe Rhinotorina (Diptera, Heleomyzidae, Rhinotorini). Zootaxa, 2736, 44 - 56." type="journal article" year="2011">Almeida &amp; Ale-Rocha 2011</bibRefCitation>
)................
<taxonomicName id="12EDA1CCFFADFF802201FB4B2DBCFB4F" ID-CoL="8DMXZ" box="[1323,1429,1243,1265]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="fonsecai">
<emphasis id="E799065DFFADFF802201FB4B2DBCFB4F" box="[1323,1429,1243,1265]" italics="true" pageId="17" pageNumber="530">R. fonsecai</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDFB6F2DB5FAF3" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDFB6F2DB2FA8E" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFB6F2DB2FA8E" pageId="17" pageNumber="530">
5(4). Cell r2+3 medially with 23 supernumerary crossveins, arising independently from R2+3 (sometimes connected posteriorly) (
<figureCitation id="4DD6C6CAFFADFF8027CDFA8D2908FA8E" box="[231,289,1306,1328]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Figs 1</figureCitation>
D, 6D)........................................................................................ 6
</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDFAA02DB5FAF3" blockId="17.[151,1437,803,2015]" box="[151,1436,1335,1363]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFAA02DB5FAF3" box="[151,1436,1335,1363]" pageId="17" pageNumber="530">
5. Cell r2+3 medially with a single (either simple or branched) supernumerary crossvein (
<figureCitation id="4DD6C6CAFFADFF80233CFAA02C72FAF3" box="[1046,1115,1335,1357]" captionStart="FIGURE 3. A G" captionStartId="8.[151,250,1721,1743]" captionTargetBox="[151,1436,265,1697]" captionTargetId="figure@8.[151,1436,265,1699]" captionTargetPageId="8" captionText="FIGURE 3. A G, Rhinotora paschoali, sp. nov. (♂ holotype, INPA). A, lateral habitus; B, dorsal habitus (arrow indicates convergent posteromedial spots on scutum); C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view; G, male genitalia, dorsal view (arrow indicates digitiform projection of cercus)." httpUri="https://zenodo.org/record/259178/files/figure.png" pageId="17" pageNumber="530">Fig 3D</figureCitation>
)......................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDFACE2DB2FA52" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDFACE2DB5FA2E" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFACE2DB5FA2E" pageId="17" pageNumber="530">
6(5). Wing with a distinct spot around crossveins arising from R2+3, darker than remaining wing spots (
<figureCitation id="4DD6C6CAFFADFF8023ADFACE2CF9FAD1" box="[1159,1232,1369,1391]" captionStart="FIGURE 6. A F" captionStartId="13.[151,250,1975,1997]" captionTargetBox="[151,1436,519,1952]" captionTargetId="figure@13.[151,1436,519,1954]" captionTargetPageId="13" captionText="FIGURE 6. A F, Rhinotora travassosi (♂, INPA). A, lateral habitus; B, dorsal habitus; C, head; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view." httpUri="https://zenodo.org/record/259181/files/figure.png" pageId="17" pageNumber="530">Fig 6D</figureCitation>
). Scutellum centrally with a dark pruinose triangle; margin golden pruinose. Male cerci with convex apical margin (
<figureCitation id="4DD6C6CAFFADFF80234DFAED2C80FA2E" box="[1127,1193,1402,1424]" captionStart="FIGURE 6. A F" captionStartId="13.[151,250,1975,1997]" captionTargetBox="[151,1436,519,1952]" captionTargetId="figure@13.[151,1436,519,1954]" captionTargetPageId="13" captionText="FIGURE 6. A F, Rhinotora travassosi (♂, INPA). A, lateral habitus; B, dorsal habitus; C, head; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view." httpUri="https://zenodo.org/record/259181/files/figure.png" pageId="17" pageNumber="530">Fig 6F</figureCitation>
)........
<taxonomicName id="12EDA1CCFFADFF80220AFAEC2DB5FA2E" ID-CoL="78TBQ" box="[1312,1436,1403,1424]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="E799065DFFADFF80220AFAEC2DB5FA2E" box="[1312,1436,1403,1424]" italics="true" pageId="17" pageNumber="530">R. travassosi</emphasis>
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</paragraph>
<paragraph id="D552DA4FFFADFF8027BDFA0F2DB2FA52" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFA0F2DB2FA52" pageId="17" pageNumber="530">
6. Wing with two parallel, often posteriorly connected supernumerary crossveins arising from the middle of R2+3; dark spot around those crossveins not darker than the remaining wing spots (
<figureCitation id="4DD6C6CAFFADFF802445FA2E2B88FA71" box="[879,929,1465,1487]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Fig 1</figureCitation>
D). Pruinescence of scutellum golden to brown, not forming a distinct pattern. Male cerci with indented apical margin (
<figureCitation id="4DD6C6CAFFADFF80244EFA412BBDFA52" box="[868,916,1494,1516]" captionStart="FIGURE 1" captionStartId="4.[151,250,1853,1875]" captionTargetBox="[151,1436,397,1830]" captionTargetId="figure@4.[151,1436,397,1831]" captionTargetPageId="4" captionText="FIGURE 1. Rhinotora diversipennis (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view (arrow indicates posterior margin of cercus)." httpUri="https://zenodo.org/record/259176/files/figure.png" pageId="17" pageNumber="530">Fig 1</figureCitation>
F)............................
<taxonomicName id="12EDA1CCFFADFF80222BFA402DB2FA52" ID-CoL="795BR" box="[1281,1435,1494,1516]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="diversipennis">
<emphasis id="E799065DFFADFF80222BFA402DB2FA52" box="[1281,1435,1494,1516]" italics="true" pageId="17" pageNumber="530">R. diversipennis</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDFA632DB2F999" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDFA632DB2F9B4" blockId="17.[151,1437,803,2015]" box="[151,1435,1524,1546]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDFA632DB2F9B4" box="[151,1435,1524,1546]" pageId="17" pageNumber="530">
7(6). Scutum with complex pattern of dark pruinose spots (
<figureCitation id="4DD6C6CAFFADFF8025C9FA632B09F9B4" box="[739,800,1524,1546]" captionStart="FIGURE 2. A B" captionStartId="6.[151,250,1771,1793]" captionTargetBox="[151,1436,329,1747]" captionTargetId="figure@6.[151,1436,329,1749]" captionTargetPageId="6" captionText="FIGURE 2. A B, Rhinotora fonsecai (♀ paratype, MNRJ). A, lateral habitus; B, scutellum. C D, R. lopesi (♂ holotype, MZUSP); C, lateral habitus; D, dorsal habitus (arrow indicates parallel posteromedial spots on scutum). E F, R. lopesi (♂, INPA). E, protandrium; F, male genitalia, ventral view (arrows indicate greater and smaller lobes of cercus)." httpUri="https://zenodo.org/record/259177/files/figure.png" pageId="17" pageNumber="530">Figs 2</figureCitation>
D, 3B, 5E)........................................... 8
</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDF9862DB2F999" blockId="17.[151,1437,803,2015]" box="[151,1435,1553,1575]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF9862DB2F999" box="[151,1435,1553,1575]" pageId="17" pageNumber="530">
7. Scutum golden pruinose, without dark pruinose spots (
<figureCitation id="4DD6C6CAFFADFF8025C6F9862B03F999" box="[748,810,1553,1575]" captionStart="FIGURE 4. A C" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[151,1436,516,1949]" captionTargetId="figure@10.[151,1436,516,1950]" captionTargetPageId="10" captionText="FIGURE 4. A C, Rhinotora pluricellata (♂ syntype, NMW). A, lateral habitus; B, dorsal habitus; C, abdomen, ventral view. D F, R. pluricellata (♂, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259179/files/figure.png" pageId="17" pageNumber="530">Figs 4</figureCitation>
E, 5B)............................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDF9B92DB2F904" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDF9B92DB2F9C1" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF9B92DB2F9C1" pageId="17" pageNumber="530">
8(7). Male fore femur about three times longer than wide. Scutum with two parallel dark pruinose spots on posteromedial region (
<figureCitation id="4DD6C6CAFFADFF8027C2F9DB290BF9DC" box="[232,290,1612,1634]" captionStart="FIGURE 2. A B" captionStartId="6.[151,250,1771,1793]" captionTargetBox="[151,1436,329,1747]" captionTargetId="figure@6.[151,1436,329,1749]" captionTargetPageId="6" captionText="FIGURE 2. A B, Rhinotora fonsecai (♀ paratype, MNRJ). A, lateral habitus; B, scutellum. C D, R. lopesi (♂ holotype, MZUSP); C, lateral habitus; D, dorsal habitus (arrow indicates parallel posteromedial spots on scutum). E F, R. lopesi (♂, INPA). E, protandrium; F, male genitalia, ventral view (arrows indicate greater and smaller lobes of cercus)." httpUri="https://zenodo.org/record/259177/files/figure.png" pageId="17" pageNumber="530">Fig. 2</figureCitation>
D). Mid tibia with one distinct ventroapical seta. Cerci with a greater medial lobe, and a smaller lateral lobe (
<figureCitation id="4DD6C6CAFFADFF80221CF9DB2D46F9DC" box="[1334,1391,1612,1634]" captionStart="FIGURE 2. A B" captionStartId="6.[151,250,1771,1793]" captionTargetBox="[151,1436,329,1747]" captionTargetId="figure@6.[151,1436,329,1749]" captionTargetPageId="6" captionText="FIGURE 2. A B, Rhinotora fonsecai (♀ paratype, MNRJ). A, lateral habitus; B, scutellum. C D, R. lopesi (♂ holotype, MZUSP); C, lateral habitus; D, dorsal habitus (arrow indicates parallel posteromedial spots on scutum). E F, R. lopesi (♂, INPA). E, protandrium; F, male genitalia, ventral view (arrows indicate greater and smaller lobes of cercus)." httpUri="https://zenodo.org/record/259177/files/figure.png" pageId="17" pageNumber="530">Fig. 2</figureCitation>
F)...............................................................................................
<taxonomicName id="12EDA1CCFFADFF80226DF9FD2DB2F9C1" ID-CoL="795C3" box="[1351,1435,1641,1663]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="lopesi">
<emphasis id="E799065DFFADFF80226DF9FD2DB2F9C1" box="[1351,1435,1641,1663]" italics="true" pageId="17" pageNumber="530">R. lopesi</emphasis>
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</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDF9112DB2F904" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF9112DB2F904" pageId="17" pageNumber="530">8. Male fore femur about four times longer than wide. Pruinose pattern of scutum not as above. Mid tibia with 13 distinct ventroapical setae........................................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDF9562DBFF891" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDF9562DB2F94A" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF9562DB2F94A" pageId="17" pageNumber="530">
9(8). Scutum without dark pruinose spots on posteromedial region (
<figureCitation id="4DD6C6CAFFADFF80241DF9562B59F969" box="[823,880,1729,1751]" captionStart="FIGURE 5. A C" captionStartId="12.[151,250,1649,1671]" captionTargetBox="[151,1436,193,1626]" captionTargetId="figure@12.[151,1436,193,1627]" captionTargetPageId="12" captionText="FIGURE 5. A C, Rhinotora salesopolitana (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus (arrow indicates absence of posteromedial spots on scutum); C, head. D F, R. spiloptera (♂ holotype, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259180/files/figure.png" pageId="17" pageNumber="530">Fig. 5</figureCitation>
E). Mid tibia with two or three distinct ventroapical setae. Male cerci triangular in dorsal view (per
<bibRefCitation id="B17CA7BEFFADFF802540F9492BAFF94A" author="Guimaraes" box="[618,902,1758,1780]" pageId="17" pageNumber="530" refString="Guimaraes, J. H. &amp; Papavero, N. (1966) A contribution to the knowledge of the neotropical Rhinotoridae (Diptera, Acalyptratae). Papeis Avulsos do Departamento de Zoologia, Secretaria de Agricultura, 18, 261 - 269." type="journal article" year="1966">Guimarães &amp; Papavero, 1966</bibRefCitation>
)..................................
<taxonomicName id="12EDA1CCFFADFF802237F9482DB2F94A" ID-CoL="6X5C2" box="[1309,1435,1758,1780]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="E799065DFFADFF802237F9482DB2F94A" box="[1309,1435,1758,1780]" italics="true" pageId="17" pageNumber="530">R. spiloptera</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDF96B2DBFF891" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF96B2DBFF891" pageId="17" pageNumber="530">
9. Scutum with two converging dark pruinose spots on posteromedial region (
<figureCitation id="4DD6C6CAFFADFF802497F96B2C20F8AC" box="[957,1033,1788,1810]" captionStart="FIGURE 3. A G" captionStartId="8.[151,250,1721,1743]" captionTargetBox="[151,1436,265,1697]" captionTargetId="figure@8.[151,1436,265,1699]" captionTargetPageId="8" captionText="FIGURE 3. A G, Rhinotora paschoali, sp. nov. (♂ holotype, INPA). A, lateral habitus; B, dorsal habitus (arrow indicates convergent posteromedial spots on scutum); C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view; G, male genitalia, dorsal view (arrow indicates digitiform projection of cercus)." httpUri="https://zenodo.org/record/259178/files/figure.png" pageId="17" pageNumber="530">Fig. 3B</figureCitation>
). Mid tibia with one distinct ventroapical seta. Male cerci short, fused all along its apex, prolonged laterally into two digitiform lobes (
<figureCitation id="4DD6C6CAFFADFF802370F88E2C81F891" box="[1114,1192,1817,1839]" captionStart="FIGURE 3. A G" captionStartId="8.[151,250,1721,1743]" captionTargetBox="[151,1436,265,1697]" captionTargetId="figure@8.[151,1436,265,1699]" captionTargetPageId="8" captionText="FIGURE 3. A G, Rhinotora paschoali, sp. nov. (♂ holotype, INPA). A, lateral habitus; B, dorsal habitus (arrow indicates convergent posteromedial spots on scutum); C, head, frontal view; D, wing (arrow indicates supernumerary crossvein); E, protandrium; F, male genitalia, ventral view; G, male genitalia, dorsal view (arrow indicates digitiform projection of cercus)." httpUri="https://zenodo.org/record/259178/files/figure.png" pageId="17" pageNumber="530">Fig. 3G</figureCitation>
)..
<taxonomicName id="12EDA1CCFFADFF8023E1F88D2D6AF891" ID-CoL="9HRRL" box="[1227,1347,1817,1839]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="paschoali" status="sp. nov.">
<emphasis id="E799065DFFADFF8023E1F88D2D6AF891" box="[1227,1347,1817,1839]" italics="true" pageId="17" pageNumber="530">R. paschoali</emphasis>
</taxonomicName>
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<emphasis id="E799065DFFADFF802263F88E2DBFF891" bold="true" box="[1353,1430,1817,1839]" pageId="17" pageNumber="530">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E19C2EAFFADFF8027BDF8A12DB2F81A" pageId="17" pageNumber="530">
<paragraph id="D552DA4FFFADFF8027BDF8A12DB5F8D4" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF8A12DB5F8D4" pageId="17" pageNumber="530">
10(7). Cell r1 with five to six supernumerary crossveins (
<figureCitation id="4DD6C6CAFFADFF8025CDF8A12B1FF8F2" box="[743,822,1846,1868]" captionStart="FIGURE 4. A C" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[151,1436,516,1949]" captionTargetId="figure@10.[151,1436,516,1950]" captionTargetPageId="10" captionText="FIGURE 4. A C, Rhinotora pluricellata (♂ syntype, NMW). A, lateral habitus; B, dorsal habitus; C, abdomen, ventral view. D F, R. pluricellata (♂, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259179/files/figure.png" pageId="17" pageNumber="530">Fig. 4B</figureCitation>
,D). Mid tibia with one ventroapical distinct seta. Male cerci abruptly narrowing from base to apex, bottle-shaped in dorsal view (per
<bibRefCitation id="B17CA7BEFFADFF8024BDF8C32C8FF8D4" author="Almeida" box="[919,1190,1876,1898]" pageId="17" pageNumber="530" refString="Almeida, J. C. &amp; Ale-Rocha, R. (2011) Comparative morphology of the male terminalia of the subtribe Rhinotorina (Diptera, Heleomyzidae, Rhinotorini). Zootaxa, 2736, 44 - 56." type="journal article" year="2011">Almeida &amp; Ale-Rocha 2011</bibRefCitation>
).......
<taxonomicName id="12EDA1CCFFADFF80223BF8C22DB5F8D4" ID-CoL="6WTC2" box="[1297,1436,1876,1898]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="pluricellata">
<emphasis id="E799065DFFADFF80223BF8C22DB5F8D4" box="[1297,1436,1876,1898]" italics="true" pageId="17" pageNumber="530">R. pluricellata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D552DA4FFFADFF8027BDF8E62DB2F81A" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF8E62DB2F81A" pageId="17" pageNumber="530">
10. Cell r1 with eight to nine supernumerary crossveins (
<figureCitation id="4DD6C6CAFFADFF8025F0F8E62B0FF839" box="[730,806,1905,1927]" captionStart="FIGURE 5. A C" captionStartId="12.[151,250,1649,1671]" captionTargetBox="[151,1436,193,1626]" captionTargetId="figure@12.[151,1436,193,1627]" captionTargetPageId="12" captionText="FIGURE 5. A C, Rhinotora salesopolitana (♂ holotype, MZUSP). A, lateral habitus; B, dorsal habitus (arrow indicates absence of posteromedial spots on scutum); C, head. D F, R. spiloptera (♂ holotype, MZUSP). D, lateral habitus; E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259180/files/figure.png" pageId="17" pageNumber="530">Fig. 5A</figureCitation>
). Mid tibia with two or three ventroapical distinct seta. Male cerci trapezoidal in dorsal view (
<bibRefCitation id="B17CA7BEFFADFF8026CFF8192B2AF81A" author="Guimaraes" box="[485,771,1934,1956]" pageId="17" pageNumber="530" refString="Guimaraes, J. H. &amp; Papavero, N. (1966) A contribution to the knowledge of the neotropical Rhinotoridae (Diptera, Acalyptratae). Papeis Avulsos do Departamento de Zoologia, Secretaria de Agricultura, 18, 261 - 269." type="journal article" year="1966">Guimarães &amp; Papavero, 1966</bibRefCitation>
)........................................
<taxonomicName id="12EDA1CCFFADFF8023DEF8182DB2F81A" ID-CoL="6X5BQ" box="[1268,1435,1934,1956]" class="Insecta" family="Heleomyzidae" genus="Rhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="salesopolitana">
<emphasis id="E799065DFFADFF8023DEF8182DB2F81A" box="[1268,1435,1934,1956]" italics="true" pageId="17" pageNumber="530">R. salesopolitana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="D552DA4FFFADFF8027BDF83B2DB5F861" blockId="17.[151,1437,803,2015]" pageId="17" pageNumber="530">
<keyStep id="6E19C2EAFFADFF8027BDF83B2DB5F861" pageId="17" pageNumber="530">
<keyLead id="6E1C797AFFADFF8027BDF83B2DB5F861" pageId="17" pageNumber="530">
11(3). Arista micropubescent (
<figureCitation id="4DD6C6CAFFADFF8026EFF83B2A38F87C" box="[453,529,1964,1986]" captionStart="FIGURE 8. A C" captionStartId="15.[151,250,1870,1892]" captionTargetBox="[151,1436,414,1848]" captionTargetId="figure@15.[151,1436,414,1848]" captionTargetPageId="15" captionText="FIGURE 8. A C, Neorhinotora mutica (♂, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view. D F, Rhinotoroides bifurcata (♂, INPA). D, lateral habitus (arrow indicates postocellar setae); E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259183/files/figure.png" pageId="17" pageNumber="530">Fig. 8C</figureCitation>
). Wing with two well-defined spots around dm-cu and on the distal extremities of R4+5 and M (
<figureCitation id="4DD6C6CAFFADFF8027CDF85E297AF861" box="[231,339,1993,2015]" captionStart="FIGURE 8. A C" captionStartId="15.[151,250,1870,1892]" captionTargetBox="[151,1436,414,1848]" captionTargetId="figure@15.[151,1436,414,1848]" captionTargetPageId="15" captionText="FIGURE 8. A C, Neorhinotora mutica (♂, MZUSP). A, lateral habitus; B, dorsal habitus; C, head, frontal view. D F, Rhinotoroides bifurcata (♂, INPA). D, lateral habitus (arrow indicates postocellar setae); E, dorsal habitus; F, head, frontal view." httpUri="https://zenodo.org/record/259183/files/figure.png" pageId="17" pageNumber="530">Figs 8AB</figureCitation>
).....................................................................
<taxonomicName id="12EDA1CCFFADFF80238EF85D2DB5F861" ID-CoL="46RW3" authority="Schiner, 1868" authorityName="Schiner" authorityYear="1868" box="[1188,1436,1993,2015]" class="Insecta" family="Heleomyzidae" genus="Neorhinotora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="530" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="E799065DFFADFF80238EF85D2D28F861" box="[1188,1281,1994,2015]" italics="true" pageId="17" pageNumber="530">N. mutica</emphasis>
(
<bibRefCitation id="B17CA7BEFFADFF802224F85E2DBDF861" author="Schiner" box="[1294,1428,1993,2015]" pageId="17" pageNumber="530" refString="Schiner, J. R. (1868) Diptera. In: Wullerstorf-Urbair (Ed.), Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859, unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Vol. 2, Part 1, Section B. Hof- und Staatsdruckerei, Wien, pp. 1 - 388. http: // dx. doi. org / 10.5962 / bhl. title. 7913" type="journal article" year="1868">Schiner, 1868</bibRefCitation>
)
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
</subSubSection>
</treatment>
</document>