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<document id="6093C48556B9E51A9CBB715E0126D4D2" ID-DOI="10.3897/zookeys.1212.122222" ID-ZooBank="9996A35A-9485-4906-AD8B-52BEABE11B16" ID-publisher-id="122222" URI-arpha="F99FC822-5424-5947-A61C-743994BB09FE" URI-zoobank="http://zoobank.org/9996A35A-9485-4906-AD8B-52BEABE11B16" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1726498494866" checkinUser="pensoft" docAuthor="Hou, Yin Meng, Zheng, Pu Yang, Yu, Hao Qi, Wang, Bin, Chen, Xiao Hong &amp; Xie, Feng" docDate="2024" docId="52D41E833A41504F88014FBD39AA1316" docLanguage="en" docName="ZooKeys 1212: 109-127" docOrigin="ZooKeys 1212" docSource="https://zookeys.pensoft.net/article/122222/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Oreolalax yanyuanensis Hou, Zheng, Yu, Wang, Chen &amp; Xie, 2024, sp. nov." docType="treatment" docUuid="D770076A-DB34-4045-AEF9-FFFBC2FBA5D8" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="127" masterDocId="F99FC82254245947A61C743994BB09FE" masterDocTitle="A new toad of Oreolalax Myers &amp; Leviton, 1962 (Anura, Megophryidae) from Sichuan Province, southwest China" masterLastPageNumber="127" masterPageNumber="109" pageNumber="109" updateTime="1726502524705" updateUser="admin">
<mods:mods id="5E34B6F80F548EC66FFB44CF23E2CEB7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="62E532C079ACDFC27AE2C69E6359E773">A new toad of Oreolalax Myers &amp; Leviton, 1962 (Anura, Megophryidae) from Sichuan Province, southwest China</mods:title>
</mods:titleInfo>
<mods:name id="30A03794629C66D946AD85593ACB4F40" type="personal">
<mods:role id="D6001597827CE5712376342767C89EBC">
<mods:roleTerm id="FCF5B4EA6353C7078A256B77B5C9880F">Author</mods:roleTerm>
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<mods:namePart id="2A692CF6FA2018AD99E71F9719CE72B0">Hou, Yin Meng</mods:namePart>
<mods:nameIdentifier id="21DAFED26F9BBF5C37DC9EA5200E49A9" type="ORCID">0000-0002-4216-9678</mods:nameIdentifier>
<mods:affiliation id="43B5A85B73C5CAA39EFC3B6BF6C72B91">College of Life Sciences, Henan Normal University, Xinxiang 453007, China &amp; Department of Herpetology, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; The Observation and Research Field Station of Taihang Mountain Forest Ecosystems of Henan Province, Xinxiang 453007, China</mods:affiliation>
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<mods:namePart id="F3116E8FA51B66FFB39E8A2F93166056">Zheng, Pu Yang</mods:namePart>
<mods:affiliation id="ECD2C765BABD5E1028B26C4C02A47286">Department of Herpetology, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China &amp; College of Life Sciences, Sichuan University, Chengdu 610065, Sichuan, China</mods:affiliation>
</mods:name>
<mods:name id="FA35BF1409073F2441A46CDE1FB6F5F8" type="personal">
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<mods:namePart id="37FB98CCF601010093FB920CEFCDB19A">Yu, Hao Qi</mods:namePart>
<mods:affiliation id="3D924B0A5CF5BFE1938CF2D56E888D4D">Department of Herpetology, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China &amp; College of Life Sciences, Sichuan University, Chengdu 610065, Sichuan, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="9197001F2F59B3C0685B19C32EFBC9FE">Wang, Bin</mods:namePart>
<mods:nameIdentifier id="DB121148494BD690BE5471C03B07A401" type="ORCID">0000-0001-6036-5579</mods:nameIdentifier>
<mods:affiliation id="5A1639E7859358C6A730A67713AB743C">Department of Herpetology, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
</mods:name>
<mods:name id="F86D4E464DF4EB9C7E7ADA468B999FFE" type="personal">
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<mods:namePart id="46DF2C0B47D2B20B079B6739433E1441">Chen, Xiao Hong</mods:namePart>
<mods:affiliation id="5A3662D1790DEF973540D9EC894F27A4">College of Life Sciences, Henan Normal University, Xinxiang 453007, China &amp; The Observation and Research Field Station of Taihang Mountain Forest Ecosystems of Henan Province, Xinxiang 453007, China</mods:affiliation>
</mods:name>
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<mods:namePart id="56B710C640E7D5CDAAC033536BC7C33C">Xie, Feng</mods:namePart>
<mods:nameIdentifier id="626381308B59BF579949B3D618EF05B1" type="ORCID">0000-0001-9532-9586</mods:nameIdentifier>
<mods:affiliation id="53D2345F71FC1ED5F383BB10896EACCB">Department of Herpetology, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
</mods:name>
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<mods:date id="F773366047656FE7B7978EED0C96A952">2024</mods:date>
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<mods:number id="85DA5321CD795B0496EA24A4E288CB14">2024-09-16</mods:number>
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<mods:number id="A8400EE5571CC1A4F3E96A011A914655">1212</mods:number>
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<mods:start id="B3669AE06AB420DBEAF49A2CBCA856FA">109</mods:start>
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<mods:identifier id="0ABA474C195777FFA2246F498E784832" type="DOI">10.3897/zookeys.1212.122222</mods:identifier>
<mods:identifier id="63FE1DD5EECC138ED43E9D4CB0FB758A" type="ZooBank">9996A35A-9485-4906-AD8B-52BEABE11B16</mods:identifier>
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<treatment id="52D41E833A41504F88014FBD39AA1316" ID-arpha="52D41E83-3A41-504F-8801-4FBD39AA1316" ID-zoobank="https://zoobank.org/D770076A-DB34-4045-AEF9-FFFBC2FBA5D8" LSID="urn:lsid:zoobank.org:act:D770076A-DB34-4045-AEF9-FFFBC2FBA5D8" LSID-ZBK="urn:lsid:zoobank.org:act:D770076A-DB34-4045-AEF9-FFFBC2FBA5D8" httpUri="http://treatment.plazi.org/id/52D41E833A41504F88014FBD39AA1316">
<subSubSection id="C80ECC75D9CFE068F1049448FC85BA70" type="nomenclature">
<paragraph id="2638C02AC0804AD7FEE5A81C29D9661A">
<taxonomicName id="E3048C32733892B55FCCA41A6970B573" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="7E0267DFD31A3D6492F18120C097B48C" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EDE91725ADBF8606870DD56CB8D29BA" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="81624A4076DED98B9134F4E1DF2BFD17" type="description">
<paragraph id="1630142E88C92EB504635FECCD61E4A7">
<figureCitation id="116C099D21D7579BC271CC7033451FD8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photos of the holotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. A dorsal view B ventral view; the white arrow indicates the position of the pectoral gland C lateral view D ventral view of hand E ventral view of foot." figureDoi="10.3897/zookeys.1212.122222.figure3" httpUri="https://binary.pensoft.net/fig/1134579">Figs 3</figureCitation>
,
<figureCitation id="CB09E5825D97F07EA38E209E8E4AD6B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. in preservative A dorsal view B ventral view C lateral view D dorsal view of the left hand E ventral view of the left foot." figureDoi="10.3897/zookeys.1212.122222.figure4" httpUri="https://binary.pensoft.net/fig/1134580">4</figureCitation>
,
<figureCitation id="F53C2DF90408B7B39673C993B9F653B9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos for the tadpole CIBSH 20230603 kd 01 in preservative of Oreolalax yanyuanensis sp. nov. A dorsal view B ventral view C lateral view D ventral view of head." figureDoi="10.3897/zookeys.1212.122222.figure5" httpUri="https://binary.pensoft.net/fig/1134581">5</figureCitation>
,
<figureCitation id="21CF50EFFD6939E052D60F1ADC7F0EC3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Specimen photos of Oreolalax yanyuanensis sp. nov. and its relative species A, B dorsal and ventral view of the holotype CIB 25333 of O. rugosus C, D dorsal and ventral view of the topotype CIB 24678 of O. liangbeiensis E, F dorsal and ventral view of the topotype CIB 24695 of O. major G, H dorsal and ventral view of the topotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. Scale bar: equal to 10 mm." figureDoi="10.3897/zookeys.1212.122222.figure6" httpUri="https://binary.pensoft.net/fig/1134582">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E56AG" type="type material">
<paragraph id="1DB8DA2A38125B5A89B92C98DCCF33F8">
<heading id="E1648CAA7EE8C074242ECE3A8C0D7E6C" reason="title">Type material.</heading>
</paragraph>
<paragraph id="F8C6CFD5FE547DFAECA20E3B44DC785A">
<materialsCitation id="6F5D165B5F0A0AC2F7BE0FF3C41A4336" collectionCode="CIBSH" collectorName="F. Xie" country="China" elevation="3108" latitude="27.473442" location="Shuhe town" longLatPrecision="1" longitude="101.78963" municipality="Yanyuan county" specimenCode="CIBSH 20230603020" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan Province" typeStatus="holotype">
<emphasis id="477D12342C72DE45CC291F68C1839804" bold="true">
<emphasis id="8BF1C0B378BC8597DBD6B08B63DD6C26" bold="true" italics="true">
<typeStatus id="6814918FCCF336EE35BD1C25461E20B4" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="1AA18687926F7A5FEF01AB1C129E19A6" collectionCode="CIBSH">CIBSH 20230603020</specimenCode>
(Fig.
<figureCitation id="34E03664907B777995C8F28C3FB7DC07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photos of the holotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. A dorsal view B ventral view; the white arrow indicates the position of the pectoral gland C lateral view D ventral view of hand E ventral view of foot." figureDoi="10.3897/zookeys.1212.122222.figure3" httpUri="https://binary.pensoft.net/fig/1134579">3</figureCitation>
),
<specimenCount id="1B90CF93C94133F53BE4427E5E0D7A03" count="1" type="male">adult male</specimenCount>
, collected by
<collectorName id="833722079AE48CDC4D9A27624FDB06E6">F. Xie</collectorName>
from
<location id="75C5104ED52B796DE37404EA2F005178" LSID="urn:lsid:plazi:treatment:52D41E833A41504F88014FBD39AA1316:75C5104ED52B796DE37404EA2F005178" country="China" latitude="27.473442" longLatPrecision="1" longitude="101.78963" municipality="Yanyuan county" name="Shuhe town" stateProvince="Sichuan Province">Shuhe town</location>
,
<collectingMunicipality id="FDC1E5CC43181C975A1E266D0D265CDE">Yanyuan county</collectingMunicipality>
,
<collectingRegion id="F061D6D233EE8E371A607EDF7EE002FD" country="China" name="Sichuan">Sichuan Province</collectingRegion>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.789624,27.473443]}">
<geoCoordinate id="43CD31B75EFB1F126AE217C8ABEE8055" degrees="27.473443" direction="north" orientation="latitude" precision="1" value="27.473442">27.473443 ° N</geoCoordinate>
,
<geoCoordinate id="3F73CDE59FA9572E35990CFA8CDDDAE0" degrees="101.789624" direction="east" orientation="longitude" precision="1" value="101.78963">101.789624 ° E</geoCoordinate>
</named-content>
,
<elevation id="1AE83DF14707CBA3D5CE88B2B9B1AA7A" metricMagnitude="3" metricUnit="m" metricValue="3.108" unit="m" value="3108.0">
<quantity id="D516EF6D0703CF3FA3F9B08FFBFF9E02" metricMagnitude="3" metricUnit="m" metricValue="3.108" unit="m" value="3108.0">3108 m</quantity>
a. s. l.
</elevation>
)
<collectingCountry id="325493D0782C9B37AEBA6A0AC1A1FB68" name="China">China</collectingCountry>
</materialsCitation>
.
<materialsCitation id="0639503430190A4F094C5F2A5CC3EABD" collectedFrom="small stream" collectingDate="2023-06-03" collectionCode="CIBSH" collectorName="P. Y. Zheng &amp; H. Q. Yu &amp; F. Xie" country="China" elevation="3127" latitude="27.475204" location="Shuhe town" longLatPrecision="1" longitude="101.78911" specimenCode="CIBSH 20230603016-19, CIBSH 20230603021, CIBSH 20230603023-24" specimenCount="7" specimenCount-adult="7" specimenCount-male="7" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="F1181EA339A57FB3EF2397D23C708AA5" bold="true">
<emphasis id="A85ECEAEBD2E7C01D80AA86B4AAC86A9" bold="true" italics="true">
<typeStatus id="FE02C07BC05C123F087F204FC23800B1" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="476B63E9F8DC34ED7FCB0F010089014B" count="7" type="male">Seven adult males</specimenCount>
collected from a
<collectedFrom id="CAB0CD323A958DB9948AD524B5D44165">small stream</collectedFrom>
of
<location id="DE8E545D045BD0F6D8CCB05C6289F06A" LSID="urn:lsid:plazi:treatment:52D41E833A41504F88014FBD39AA1316:DE8E545D045BD0F6D8CCB05C6289F06A" country="China" latitude="27.475204" longLatPrecision="1" longitude="101.78911" name="Shuhe town" stateProvince="Sichuan">Shuhe town</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3ABG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.789108,27.475205]}">
<geoCoordinate id="00229A8B65A1A230B7DB8C817204EA08" degrees="27.475205" direction="north" orientation="latitude" precision="1" value="27.475204">27.475205 ° N</geoCoordinate>
,
<geoCoordinate id="8F2D2332AB20148AE4DCA5BD8A51C000" degrees="101.789108" direction="east" orientation="longitude" precision="1" value="101.78911">101.789108 ° E</geoCoordinate>
</named-content>
,
<elevation id="AE6BDC3309C4E41D3619F8EFDEEBD9C8" metricMagnitude="3" metricUnit="m" metricValue="3.127" unit="m" value="3127.0">
<quantity id="DDDFC8DFC4EB1911F6E0E2C835CBB346" metricMagnitude="3" metricUnit="m" metricValue="3.127" unit="m" value="3127.0">3127 m</quantity>
a. s. l.
</elevation>
); on
<date id="02A5521E07942334B68703FC675FE9CC" value="2023-06-03">
<collectingDate id="90357D79BD1EC562E578522CBCEDEFDC" value="2023-06-03">June 3, 2023</collectingDate>
</date>
(
<specimenCode id="B440A824816D920CA1F7BBE653BE96EC" collectionCode="CIBSH">CIBSH 2023060301619</specimenCode>
,
<specimenCode id="D496870CFE4F7B2DD8C7EF9A205D9126" collectionCode="CIBSH">CIBSH 20230603021</specimenCode>
,
<specimenCode id="C244F24F1FC72737F64DD9E6EB4C4547" collectionCode="CIBSH">CIBSH 2023060302324</specimenCode>
) by
<collectorName id="014CA6928E9D16904FF674D01E0E5979">P. Y. Zheng</collectorName>
,
<collectorName id="0587539B855F2292027443EB1AEF550B">H. Q. Yu</collectorName>
and
<collectorName id="B5E8202FD414C3C6F905C233AFF9EDF0">F. Xie</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="44236872B19C107671659672C38C50EF" ID-DOI="10.3897/zookeys.1212.122222.figure3" ID-arpha="223BEE09-394B-5A86-A03E-8744E19E4081" httpUri="https://binary.pensoft.net/fig/1134579" startId="F3">
<paragraph id="3A49E3E923B0AD7794234CDD418673FF">
<label id="FAABFA9C208A414153F7D34D511434A0">Figure 3.</label>
</paragraph>
<paragraph id="5766DC9881B9F52AE23D28697BB752D9">
Photos of the holotype CIBSH 20230603020 of
<taxonomicName id="95ACDCD73F522CCCDBC5BC43460EFC13" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="6FC3674C749EEC08970AE284586AC75A" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98287CA806658BB2107988A38650C1E2" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E161F48AA5CB50C9C9D880F51359662B" bold="true">A</emphasis>
dorsal view
<emphasis id="B800732CC7F42D6E927B87B1BFD36FBF" bold="true">B</emphasis>
ventral view; the white arrow indicates the position of the pectoral gland
<emphasis id="C38D21AA1E88A1F7FA7567755F487142" bold="true">C</emphasis>
lateral view
<emphasis id="7BEE00AA7EEB437113E8F9A85F8648B4" bold="true">D</emphasis>
ventral view of hand
<emphasis id="E2CB09F89D8A50F4BF17DB44684C0B7A" bold="true">E</emphasis>
ventral view of foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDCBG" type="Other specimens">
<paragraph id="C2ECF5C9AADCBC518C2498FCAE8B2D89">
<heading id="07C98F15ADE43536F6EBD4C84CA54267" reason="title">Other specimens.</heading>
</paragraph>
<paragraph id="F90411F45F28038752FB4C2D061DA458">
<materialsCitation id="6A70109DDF0DC09F2295C9BC3F56841F" collectedFrom="small stream" collectingDate="2023-06-03" collectionCode="CIBSH" collectorName="P. Y. Zheng &amp; H. Q. Yu &amp; F. Xie" elevation="3032" latitude="27.466242" location="Shuhe town" longLatPrecision="1" longitude="101.786934" municipality="Shuhe town" specimenCode="CIBSH 20230603" specimenCount="7">
<specimenCount id="13A4DB0E880F2EAF3C33C1C15AC8D71B" count="7" type="generic">Seven tadpoles</specimenCount>
(
<specimenCode id="A214A5EEFFDA70A44BF1F8B87FC64E39" collectionCode="CIBSH">CIBSH 20230603</specimenCode>
kd 0107) collected from a
<collectedFrom id="9AFC1526DBD696C9DAE55A605CF63114">small stream</collectedFrom>
of
<location id="4BC3156F2B8CC9582331888EAF049DA5" LSID="urn:lsid:plazi:treatment:52D41E833A41504F88014FBD39AA1316:4BC3156F2B8CC9582331888EAF049DA5" latitude="27.466242" longLatPrecision="1" longitude="101.786934" municipality="Shuhe town" name="Shuhe town">Shuhe town</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMCBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.786933,27.466241]}">
<geoCoordinate id="FAF1FDE4E913467D1C985EE91B6AD629" degrees="27.466241" direction="north" orientation="latitude" precision="1" value="27.466242">27.466241 ° N</geoCoordinate>
,
<geoCoordinate id="CC396B7BD6D06D8B01DACA25CC5B8232" degrees="101.786933" direction="east" orientation="longitude" precision="1" value="101.786934">101.786933 ° E</geoCoordinate>
</named-content>
,
<elevation id="02E292960E816A0C455F81F3FEF2B68B" metricMagnitude="3" metricUnit="m" metricValue="3.032" unit="m" value="3032.0">
<quantity id="B719838A0CC1C70A5D271D71AB931F3F" metricMagnitude="3" metricUnit="m" metricValue="3.032" unit="m" value="3032.0">3032 m</quantity>
a. s. l.
</elevation>
); on
<date id="7AD6F9685CF0878BB3BA45BEECFD0532" value="2023-06-03">
<collectingDate id="518FA66CDCB90CA895CF9A980BBDBF9C" value="2023-06-03">June 3, 2023</collectingDate>
</date>
; by
<collectorName id="9E5AE0CC372EB98740155BCC934B5E8D">P. Y. Zheng</collectorName>
,
<collectorName id="A2DB08EEE8D8313437C6448BE73CE2DC">H. Q. Yu</collectorName>
and
<collectorName id="37629106FF137D6022D8A63704EB3501">F. Xie</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERCBG" type="diagnosis">
<paragraph id="27A1F7D2376C4E98F7DE20FCA203048F">
<heading id="8C7AF03138B59B58557061DC87CB9CF0" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="C57BF1FC6FA521C81E725765056B9A06">
The new species is assigned to genus
<taxonomicName id="E8F89F710646101D80F5B3C7C13910AC" authorityName="Myers &amp; Leviton" authorityYear="1962" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="8A026587847DB66F4AACFFD2328354BE" italics="true">Oreolalax</emphasis>
</taxonomicName>
based on following characters: maxillary teeth prominent; back rough, scattered with large warts, covered with oval black spots; pupil vertical; tongue moderately broad, notched behind; femoral glands prominent; pectoral and axillary gland present in males.
</paragraph>
<paragraph id="A5802AB5C8CEFA7B7FD91489F631E06D">
The new species differs from its congeners by a combination of the following characters: body size moderate
<quantity id="1BB8DF25FA10A6530121630EDDA20D2C" metricMagnitude="-2" metricUnit="m" metricValue="4.63" metricValueMax="5.279999999999999" metricValueMin="3.9799999999999995" unit="mm" value="46.3" valueMax="52.8" valueMin="39.8">39.852.8 mm</quantity>
in male; head broad; tympanum hidden; distinct black spots present on flanks; toes 1 / 3 webbed, with broad lateral fringes; dorsal body deep brown or yellowish-brown; belly smooth, middle yellow, scattered fine mottling, abdominal margin more spotted; skin on dorsum rough with dense and varied size granules, warts are covered with dark spots or no spots; iris light orange or light yellow above, creamy silver white bellow; and middle pectoral glands are evident in males.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6CBG" type="description">
<paragraph id="D267481DD8EF1C015C08A5C6B287D113">
<heading id="7472AFA3EE177C395030528F16AF7AE6" reason="title">
<typeStatus id="61CC7CEA8C5B4F55CFA652EB95CB3D38" type="holotype">Holotype</typeStatus>
description
</heading>
</paragraph>
<paragraph id="A946D64EF6256CE0F63D087D7CA73FB8">
<emphasis id="70FABC200B37CD5A788C9E697F7BC5D1" bold="true">
(Fig.
<figureCitation id="0213B1E068B56194041F9D5E42980963" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photos of the holotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. A dorsal view B ventral view; the white arrow indicates the position of the pectoral gland C lateral view D ventral view of hand E ventral view of foot." figureDoi="10.3897/zookeys.1212.122222.figure3" httpUri="https://binary.pensoft.net/fig/1134579">3</figureCitation>
).
</emphasis>
Body size moderate (
<abbrev id="ABBRID0EMDBG" xlink_title="snoutvent length">
<collectionCode id="DF5EFE5F527497563C1694DFB6688179">SVL</collectionCode>
</abbrev>
<quantity id="BA24C2108FC4698EB24ABCA02ECDBEE8" metricMagnitude="-2" metricUnit="m" metricValue="4.74" unit="mm" value="47.4">47.4 mm</quantity>
). Head width greater than length (
<abbrev id="ABBRID0EQDBG" xlink_title="head width">
<collectionCode id="E7F9473E108B7958CB22DB54AD732EA7">HW</collectionCode>
</abbrev>
<quantity id="B254F33BD330AE321E107C4619780A13" metricMagnitude="-2" metricUnit="m" metricValue="1.76" unit="mm" value="17.6">17.6 mm</quantity>
,
<abbrev id="ABBRID0EUDBG" xlink_title="head length">HL</abbrev>
<quantity id="278616B91FAD8C695FF23882886E5706" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" unit="mm" value="16.2">16.2 mm</quantity>
); maxillary teeth developed, without vomerine teeth or acoustic sac; snout bluntly rounded in dorsal view, slightly projecting over lower jaw, longer than eye diameter (
<abbrev id="ABBRID0EYDBG" xlink_title="snout length">SL</abbrev>
<quantity id="D2708DE12921E5203451C819E1DC0C6B" metricMagnitude="-3" metricUnit="m" metricValue="6.3" unit="mm" value="6.3">6.3 mm</quantity>
,
<abbrev id="ABBRID0E3DBG" xlink_title="eye diameter">
<collectionCode id="62B6B2DD0CEA68F02D68D4E9C8701002">ED</collectionCode>
</abbrev>
<quantity id="338B9E009E5F0C26516F70A6560D2BA4" metricMagnitude="-3" metricUnit="m" metricValue="5.9" unit="mm" value="5.9">5.9 mm</quantity>
); canthus rostralis indistinct, interorbital distance (
<abbrev id="ABBRID0EAEBG" xlink_title="interorbital distance">
<collectionCode id="CAE8700A4B1C4F4693AD3CFADEFCCCE1">IOD</collectionCode>
</abbrev>
5.0 mm) wider than internarial distance (
<abbrev id="ABBRID0EEEBG" xlink_title="internarial distance">
<collectionCode id="9289E36AD633506A6303678BBC49DE25" country="USA" lsid="urn:lsid:biocol.org:col:15485" name="Indiana University" type="Herbarium">IND</collectionCode>
</abbrev>
<quantity id="5B5885740DECA34DA4802D673D0F0484" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
), distinctly larger than upper eyelid (
<abbrev id="ABBRID0EIEBG" xlink_title="upper eyelid width">
<collectionCode id="F8EC7B8D0860B3A4A7111768E071B63A">UEW</collectionCode>
</abbrev>
3.0 mm); nostrils oval, closer to tip of snout than eyes (
<abbrev id="ABBRID0EMEBG" xlink_title="eyenostril distance">
<collectionCode id="D4D0379B77E9CF2D5A7736343349D0A2">EN</collectionCode>
</abbrev>
<quantity id="B47E1D189865406203FC0EE7EDCB1DC6" metricMagnitude="-3" metricUnit="m" metricValue="3.1" unit="mm" value="3.1">3.1 mm</quantity>
,
<abbrev id="ABBRID0EQEBG" xlink_title="snoutnostril length">
<collectionCode id="3293953727ED39ECF3DD65BF03B3185B">NS</collectionCode>
</abbrev>
3.0 mm); no tympanic membrane; supratympanic fold broad; tongue moderately broad, notched behind; pupil vertical.
</paragraph>
<paragraph id="3F5768FFE08DEC16EECAFF1C088AD349">
Fingers moderate, relative finger lengths: I &lt;II &lt;IV &lt;
<collectionCode id="5D3397A3FD4D948F627808C7419ED401">III</collectionCode>
; finger tips slightly dilated; subarticular tubercles absent; inner palmar tubercle large, nearly rounded, outer palmer tubercle small, oval, completely separated.
</paragraph>
<paragraph id="A1692CEE1B8D33F0DED081B00AB2D168">
Hindlimbs relatively long, length 177 % of body length; shank length subequal to thigh length, slightly shorter than foot length (
<abbrev id="ABBRID0EXEBG" xlink_title="thigh length">
<collectionCode id="268748939B002E8A904C8417B69A9562" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13138" name="Grierson Museum" type="Herbarium">THL</collectionCode>
</abbrev>
<quantity id="1D2B2616A92B9BE3B6E469C21B114A78" metricMagnitude="-2" metricUnit="m" metricValue="2.34" unit="mm" value="23.4">23.4 mm</quantity>
,
<abbrev id="ABBRID0E2EBG" xlink_title="tibia length">TL</abbrev>
<quantity id="03E6FEC673B9A47BB9C399EED32C447D" metricMagnitude="-2" metricUnit="m" metricValue="2.38" unit="mm" value="23.8">23.8 mm</quantity>
,
<abbrev id="ABBRID0E6EBG" xlink_title="foot length">FL</abbrev>
<quantity id="89C4A3502243E77ADE87EB70A5E1A8B5" metricMagnitude="-2" metricUnit="m" metricValue="2.44" unit="mm" value="24.4">24.4 mm</quantity>
); heels partially overlap when thighs are positioned at right angles to the body and tibia-tarsal articulation reaches the middle eye when leg stretched; toes 1 / 3 webbed with distinct fringes; inner metatarsal tubercle long oval, small.
</paragraph>
<paragraph id="4209D7A7918F34C19F3D23EBFB216460">In life, dorsal body and head rough, back with moderate sparse granules, relatively small warts on head; with dark-brown triangular between eyes; dorsal arms and hindlimbs with small granules and bumps; distinct warts cover the fold and posterior of snout. Ventral skin smooth; pectoral glands flat; pectoral glands evident, chest spines and finger spines not visible; femoral glands slightly swollen, distinct on posterior thigh. Small verrucous granules around the anus.</paragraph>
<paragraph id="80D63850F56B4BFE53192D886DFADD42">Large brown markings on dorsum, dark brown triangular pattern between eyes; large markings dorsum brown; ventral skin medium yellow, with scattered little dark speckling; supratympanic fold dark brown; lateral head and flanks brown with dark patches; throat mixed pink and orange yellow, margin with small beige warts; chest pink and the pectoral glands medium yellow; forelimbs covered with black irregular spots; dorsal limbs yellowish-brown, the spots and stripes of dorsal upper arms and tibiotarsal articulation black; ventral arms, thigh, tibia medium yellow with flesh marking; upper iris light orange yellow, lower iris creamy white, both parts embedded black mesh lines.</paragraph>
<paragraph id="FB2C5C674C7E76EDCB1FBD72E206B2D8">
In preservative (75 % ethanol), dorsal body and head dark grey; irregular spots in forelimbs, black longitudinal stripes on hindlimbs; ventral surface beige, throat and arms beige white; with grayish-brown speckling; mandibular margin warts white; pectoral glands and ventral of the hindlimb beige yellow, scattered black spots; hand and feet dark grey, finger tips and palms grayish-white, inner metatarsal tubercle grey; lateral grey on snout and undereye, patches black; skins beneath supratympanic fold dark grey, flanks grey, covered with creamy white warts, black spots around the edges of warts. Perianal warts and femoral gland creamy-white (Fig.
<figureCitation id="E8FE4500AB4211815244D510BCC6D87F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. in preservative A dorsal view B ventral view C lateral view D dorsal view of the left hand E ventral view of the left foot." figureDoi="10.3897/zookeys.1212.122222.figure4" httpUri="https://binary.pensoft.net/fig/1134580">4</figureCitation>
).
</paragraph>
<caption id="614E9EB9488CD63CAED07E86E6EBCC91" ID-DOI="10.3897/zookeys.1212.122222.figure4" ID-arpha="669386D1-66A2-5CFC-BC77-2AD73638BD8E" httpUri="https://binary.pensoft.net/fig/1134580" startId="F4">
<paragraph id="B9BD7E0B896ECA41721CC9BBC1597082">
<label id="AC8F0B82819AEAA6B006E9C3D6FA64BD">Figure 4.</label>
</paragraph>
<paragraph id="B4122E76B69E277D1D1956915B9E256B">
The holotype CIBSH 20230603020 of
<taxonomicName id="3957885085B166AF89F544F9B8DD9D3C" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="2BE82828E79A8AEB5B17B7946ED78BE0" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3263F7C2A1A66B6941C839D8838C964" rank="species">sp. nov.</taxonomicNameLabel>
in preservative
<emphasis id="F771FB72B43482B296C1CEFB9A73FE9F" bold="true">A</emphasis>
dorsal view
<emphasis id="9CD83BF1ACAA29E2CFE11D326ACC1444" bold="true">B</emphasis>
ventral view
<emphasis id="F3FE201508EBF151DCA6865B7AC06CF3" bold="true">C</emphasis>
lateral view
<emphasis id="39907CF1BFA51FAC48DDF34B13B308BB" bold="true">D</emphasis>
dorsal view of the left hand
<emphasis id="F03533F068449406B0A251612CBA7854" bold="true">E</emphasis>
ventral view of the left foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ENGBG" type="Tadpoles">
<paragraph id="06CA92012CC8B892F9136274D7A74481">
<heading id="82287EF21952EAAAA03300C3718BF80E" reason="title">Tadpoles.</heading>
</paragraph>
<paragraph id="7EAAD489F6F92B8047E547942A5AFC73">
Measurements see Suppl. material
<xref id="E6AA6AFB0B349E88834F3B3BE5CD94F3" ref-type="supplementary-material" rid="S4">4</xref>
. Description based on preserved tadpole
<collectionCode id="5A329D40257479B0CCCE0414AF1E2D54">CIBSH</collectionCode>
20230603 kd 01 at Gosner stage 37 (
<abbrev id="ABBRID0EXGBG" xlink_title="total length">
<collectionCode id="C13DCFEF99AC9375445627D97A13FE4B">TOL</collectionCode>
</abbrev>
<quantity id="35A1DB807ABC6D79F6D608CBBFDC8FC2" metricMagnitude="-2" metricUnit="m" metricValue="6.2299999999999995" unit="mm" value="62.3">62.3 mm</quantity>
,
<collectionCode id="14F1D645DC7F34767FE1BA403AB39E9A">BL</collectionCode>
<quantity id="3F49DD6E3ED60056ADD5A68A9BF6A04D" metricMagnitude="-2" metricUnit="m" metricValue="2.06" unit="mm" value="20.6">20.6 mm</quantity>
) (Fig.
<figureCitation id="3BF6599B37761ADD98DE2DD692FFA2E7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos for the tadpole CIBSH 20230603 kd 01 in preservative of Oreolalax yanyuanensis sp. nov. A dorsal view B ventral view C lateral view D ventral view of head." figureDoi="10.3897/zookeys.1212.122222.figure5" httpUri="https://binary.pensoft.net/fig/1134581">5</figureCitation>
). The mouth is located below the rostral end; labial tooth row formula I: (5 + 5) / (5 + 5): I; the upper lip papillae is large, the central missing length is close to 3 papillae position, the lower lip papillae is small and pointed; and there are small teeth on the auxiliary processes of the oral corner; jaw sheaths strong, serrated, the lip teeth are daggerlike; body elliptical in dorsal view, body width is 113 % of height; snout rounded, eye positioned dorsolateral;
<abbrev id="ABBRID0E6GBG" xlink_title="snout length">SL</abbrev>
29 % of BL; eyes relatively small,
<abbrev id="ABBRID0EDHBG" xlink_title="eye diameter">
<collectionCode id="1D9CC0DC4ED82F6FB1A8543359640431">ED</collectionCode>
</abbrev>
8.3 % of BL; nostrils near oval; tail long and muscled,
<abbrev id="ABBRID0EHHBG" xlink_title="tail length">TAL</abbrev>
210 % of BL;
<abbrev id="ABBRID0ELHBG" xlink_title="tail height">TAH</abbrev>
113 % of
<abbrev id="ABBRID0EPHBG" xlink_title="maximum body height">BH</abbrev>
;
<abbrev id="ABBRID0ETHBG" xlink_title="maximum width of tail base">TBW</abbrev>
44 % of
<abbrev id="ABBRID0EXHBG" xlink_title="maximum body width">BW</abbrev>
;
<abbrev id="ABBRID0E2HBG" xlink_title="snout to spiraculum">SS</abbrev>
53 % of BL. Body dark brown in the back and lateral view, creamy yellow in the ventral; tail brown; the single opening of the spiracle lateral, without a free distal tube; tail end blunt; faint brown cloud spots faintly visible on upper caudal fin, caudal fin light and broad.
</paragraph>
<caption id="B6DC8EA33119E348B79EFBBCF5AB6F2E" ID-DOI="10.3897/zookeys.1212.122222.figure5" ID-arpha="1E0177F7-4426-5BB8-9F99-0499EF3A1C16" httpUri="https://binary.pensoft.net/fig/1134581" startId="F5">
<paragraph id="770A74FC6E74CB047196B6698C6EC213">
<label id="44EF3D3EB9EC1A57D317E3971D861DB5">Figure 5.</label>
</paragraph>
<paragraph id="9BEDA4BBB3F9441A79E9DBA663109531">
Photos for the tadpole CIBSH 20230603 kd 01 in preservative of
<taxonomicName id="B5AE18D53025C363BADF80CBC17E7B69" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="1CFE316D0667FD008B171CBFE96D4F48" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF03DC70DE893B4261D2B7DD75BACE26" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7DCF015FB3A2004143212748CB5B20AD" bold="true">A</emphasis>
dorsal view
<emphasis id="B83E56FBB1FA71303D7C117FE41813B3" bold="true">B</emphasis>
ventral view
<emphasis id="777CD17BE29CE5EB1EA254D3A689B03A" bold="true">C</emphasis>
lateral view
<emphasis id="9BB83DAA3E1BD18E7D10E69FFF736893" bold="true">D</emphasis>
ventral view of head.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E6IBG" type="variation">
<paragraph id="CDC3D118769060036595C1FBBD1341B1">
<heading id="DFEAB2A62A86772AB2CFDFB0EE6BB07B" reason="title">Variation.</heading>
</paragraph>
<paragraph id="CE3C590D11F7F9166C5C746684F392CE">
Measurements variation of specimens see Suppl. material
<xref id="D2E08DF89325187F7401700CBA60327B" ref-type="supplementary-material" rid="S1">1</xref>
(unit in mm). Half of the individuals had faint markings on the abdomen (
<collectionCode id="FEB91056E7D1019227517C921779A568">CIBSH</collectionCode>
20230603016,
<collectionCode id="4722F7A308A4B56BD6923DBDFFDA07EA">CIBSH</collectionCode>
20230603018,
<collectionCode id="F50ECE2FBE6555ECDBE1965B469FE584">CIBSH</collectionCode>
20230603021,
<collectionCode id="BAFB74685BEA189753320F5FFDB827DD">CIBSH</collectionCode>
20230603023), and half had more markings on the abdomen (
<collectionCode id="7C85829AF17676469643750A35043684">CIBSH</collectionCode>
20230603017,
<collectionCode id="F2C6833A606DD19F2AE28EA7EE592279">CIBSH</collectionCode>
20230603019,
<collectionCode id="121D04FB605A477B5AB8B66AFB6E3B88">CIBSH</collectionCode>
20230603020,
<collectionCode id="0F4545268956B79628EE681F3484D0D3">CIBSH</collectionCode>
20230603022). Color of each specimen was brownish-yellow (
<collectionCode id="2147CD70A2465277F43C363BE5F810DE">CIBSH</collectionCode>
20230603018,
<collectionCode id="227BCD70D3E8E52678299411260B8C0A">CIBSH</collectionCode>
20230603023), medium brown (
<collectionCode id="0B28E65E7F02BBB818B8645526CC3BA7">CIBSH</collectionCode>
20230603016,
<collectionCode id="49C149489E55F21209EBFD03D3DCF9AC">CIBSH</collectionCode>
20230603019,
<collectionCode id="73A748EBECAEE3EAEE9C60B949E02AC9">CIBSH</collectionCode>
20230603021,
<collectionCode id="234BABB2BFD38DCB2260B6FF3D94DA46">CIBSH</collectionCode>
20230603022), or dark brown (
<collectionCode id="F68D1402D0190DC245597AEB96F80FD4">CIBSH</collectionCode>
20230603017,
<collectionCode id="CF8F302B0F7903ADB467CE4587ADC7B4">CIBSH</collectionCode>
20230603020). Dorsal markings are obvious in most individuals (
<collectionCode id="DEA3A3FFDC560A54ABC9296259CFF333">CIBSH</collectionCode>
2023060301622), except one (
<collectionCode id="4B5F28BB9B718995736BAFD689D71F33">CIBSH</collectionCode>
20230603023). Lateral markings are obvious in most individuals (
<collectionCode id="EEEBEF616D2B7F64999E0156CB8041F4">CIBSH</collectionCode>
2023060301621,
<collectionCode id="F534F969FA1047C70260C246E0AB5FFD">CIBSH</collectionCode>
20230603023), except one (
<collectionCode id="B3A0228046E20199477F6A4D470D3083">CIBSH</collectionCode>
20230603022). Color of warts covering the back was yellowish-brown (
<collectionCode id="B15227E96277BA017889E64D371D97C7">CIBSH</collectionCode>
20230603022), light brown (
<collectionCode id="D4AF09AD157C6FF2F0496B1E32D16D02">CIBSH</collectionCode>
20230603023), brown (
<collectionCode id="BC4298F08907B94A2751EED37AD1E08B">CIBSH</collectionCode>
20230603016,
<collectionCode id="FF408CF0C2EA5300BF893C2E08AD3781">CIBSH</collectionCode>
20230603020), dark brown (
<collectionCode id="89EE16926DDB8EF69442F16413A4C185">CIBSH</collectionCode>
20230603018,
<collectionCode id="470ECA5376A24C0C8F586A3DDB2B81B1">CIBSH</collectionCode>
20230603019), black (
<collectionCode id="EDE888BB254E6B96240E2E93304AB36D">CIBSH</collectionCode>
20230603017,
<collectionCode id="83A8EB310AD271131BFB8CF6FA321AFD">CIBSH</collectionCode>
20230603021). Wart size was relatively small in some individuals (
<collectionCode id="79A6EA3E129D6AE20B3BC2533350D179">CIBSH</collectionCode>
20230603017,
<collectionCode id="0A2D284931F2B9D0FFFC92A513A67882">CIBSH</collectionCode>
20230603020,
<collectionCode id="7BD994D3CBB25376C226A643245EBCAE">CIBSH</collectionCode>
20230603021,
<collectionCode id="B98A3FE207B401295AAB301AADC45C07">CIBSH</collectionCode>
20230603023) and relatively large in others (
<collectionCode id="663562E88F8257E10A8D809AF8BA014D">CIBSH</collectionCode>
20230603016,
<collectionCode id="C8A5C9FDB93BF74FBE961B97A626C506">CIBSH</collectionCode>
20230603018,
<collectionCode id="A0AAFCDAAA9E97E249371F64C040757F">CIBSH</collectionCode>
20230603022,
<collectionCode id="39AB16C79125DD3D7A39473D4D839945">CIBSH</collectionCode>
20230603023). Half of the specimens had dark temporal folds (
<collectionCode id="1C09B403F50831C9A537FCFF3AB7C015">CIBSH</collectionCode>
20230603017,
<collectionCode id="F42D0AA1F23EE8BB7437E6CF4FE6EE96">CIBSH</collectionCode>
20230603018,
<collectionCode id="EA9B9F935C1EE79679294324CB8B2988">CIBSH</collectionCode>
20230603020,
<collectionCode id="6EDE3E345270FA0DBD248D3C93D95F11">CIBSH</collectionCode>
20230603021), in others the folds were generally lighter (
<collectionCode id="E26AE5CC6E95BA5DE9CC980D6CDE4729">CIBSH</collectionCode>
20230603016,
<collectionCode id="D5CFC7338C0CB52782949E71046DAF52">CIBSH</collectionCode>
20230603019,
<collectionCode id="3E40571DC1E7CDACE0733E494B8984CC">CIBSH</collectionCode>
20230603022,
<collectionCode id="B332972B4C774B88FBFB45C55BEC0710">CIBSH</collectionCode>
20230603023).
</paragraph>
<paragraph id="900246D78A080B7E4AA6E6DDA8E2C80E">
Measurements variation of tadpoles see Suppl. material
<xref id="5D9E79C61648B9DE58BE3F63C44D6AE8" ref-type="supplementary-material" rid="S4">4</xref>
. The dorsal color of
<collectionCode id="5A5E8ECCE7F648DB044AA4A7E002E473">CIBSH</collectionCode>
20230606 kd 01,
<collectionCode id="DD0D3E2A8EC6384DF89E2B26E469B95A">CIBSH</collectionCode>
20230606 kd 0304,
<collectionCode id="DD5DB53843738627DC263F100048E4AF">CIBSH</collectionCode>
20230606 kd 0506 is nearly black, while numbers
<collectionCode id="9BF333D40CC72C4C8B105F6BE4F84247">CIBSH</collectionCode>
20230606 kd 02 and
<collectionCode id="5F962B8265941139BB0DBFA34D591632">CIBSH</collectionCode>
20230606 kd 07 are dark brown. Light brown cloud spots on the upper caudal fin, ranged from faintly visible (
<collectionCode id="58F3305CD2AF300556DE45A7BA53727A">CIBSH</collectionCode>
20230606 kd 01,
<collectionCode id="8D7A4BC6A57E7C3B96CDAA09853D88F3">CIBSH</collectionCode>
20230606 kd 03) to obvious (
<collectionCode id="70642819139F54C64836FC9F612A2533">CIBSH</collectionCode>
20230606 kd 02,
<collectionCode id="47A339906B0759C09EAE2D2319581BEA">CIBSH</collectionCode>
20230606 kd 0407).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPJBG" type="comparisons">
<paragraph id="BEC45335C58E725A416851C5BC47AD0B">
<heading id="89AA7BF6A7FFD71378B072B15A0D5AF9" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="9556382FF53ED1536DAD5B6BAFC6BA15">
In
<taxonomicName id="BCE81CA9D3029BC96A5B8AB637EB2AF6" authorityName="Myers &amp; Leviton" authorityYear="1962" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="3ABD7B05B0BF8CBE559A430A99A1426C" italics="true">Oreolalax</emphasis>
</taxonomicName>
, 19 species occur in southwest
<collectingCountry id="27E513C3B6818033E9DA3B3FC3B1B1BC" name="China">China</collectingCountry>
and northern
<collectingCountry id="134FF0AA49C684C28541272F64556189" name="Vietnam">Vietnam</collectingCountry>
.
<taxonomicName id="A5E7D70226C32621C53CECAEEC143F2E" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="223A9817144E6CAF74E4A3D8579F8366" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1F054C5D1A274FEB034F3E7F291D0264" rank="species">sp. nov.</taxonomicNameLabel>
could be easily distinguished from them by several characters (Suppl. material
<xref id="2364E4FE6D8B6A7AED52534A42AD54CC" ref-type="supplementary-material" rid="S5">5</xref>
). By having moderate body size (
<quantity id="8A1B5445C649074DB954C6F5329852F6" metricMagnitude="-2" metricUnit="m" metricValue="4.63" metricValueMax="5.279999999999999" metricValueMin="3.9799999999999995" unit="mm" value="46.3" valueMax="52.8" valueMin="39.8">39.852.8 mm</quantity>
) in males, the new species differs from
<taxonomicName id="784FC0F37DC3AEA8D4026DE055414AB0" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="082550B76B798EF0AE5EAD22D40127AC" italics="true">O. major</emphasis>
</taxonomicName>
(vs.
<quantity id="BF32E1411AFE8F5A2C894430617D39FA" metricMagnitude="-2" metricUnit="m" metricValue="6.395" metricValueMax="6.87" metricValueMin="5.92" unit="mm" value="63.95" valueMax="68.7" valueMin="59.2">59.268.7 mm</quantity>
),
<taxonomicName id="3DD9D58CA7483E85FD5092A8E08D3EA9" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="D58EE7F9F8FD0774A3D2A75EE6BBA625" italics="true">O. popei</emphasis>
</taxonomicName>
(vs. 60.069.0 mm),
<taxonomicName id="E7FA83762EF31F5AB11CC57D7F789F83" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="7F3065D767622EAFB174F92C136ACF07" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
(vs.
<quantity id="6FA9D1F7E3413355191FD4FFF605E2BD" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="7.35" metricValueMin="5.75" unit="mm" value="65.5" valueMax="73.5" valueMin="57.5">57.573.5 mm</quantity>
),
<taxonomicName id="D51DF0CD42A8D1C5E02DC0B2F0CBAB36" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="5CA6B810BEBB6B47440E669EFE297228" italics="true">O. sterlingae</emphasis>
</taxonomicName>
(vs.
<quantity id="20F39B2699F055EF0FE5C1CF01B511CB" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" unit="mm" value="36.8">36.8 mm</quantity>
), and
<taxonomicName id="BD45F9D60437C0092A6134D13D369DBB" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="6365642E26ABE7C88B4E476546B1DC95" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(vs.
<quantity id="DC79BC343D132D7506916C389436A5A0" metricMagnitude="-2" metricUnit="m" metricValue="5.82" unit="mm" value="58.2">58.2 mm</quantity>
).
</paragraph>
<paragraph id="2E141FE7F20A3E936D5DFBEB43D7AEB3">
By having head width&gt; head length, the new species differs from
<taxonomicName id="292D004F9C54E8C3D54F95082E9391D5" authorityName="Tian" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chuanbeiensis">
<emphasis id="1B13B9BCAA27BDF00346E7331B333AC6" italics="true">O. chuanbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="C29DC362068E622297A19F5C799DC0B4" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="F21E39CE6767E859A023ACBC944B782A" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
(vs. head width ≈ head length),
<taxonomicName id="B4CAD8B91BB055AA5081C1AFF0C7C86A" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="558FCEB4B14ABD4D573EE43997F610CE" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(vs. head width = head length),
<taxonomicName id="CA24AD766C210BF54A1A44EC59FBC3BD" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="2D6B86D59D7DB95D13DCCBA7387C9224" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="B23B9C6E04124A2B778C3C026A6B63AE" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="D6AE01C5B9F0DAED1B6279FFFD2F411B" italics="true">O. popei</emphasis>
</taxonomicName>
,
<taxonomicName id="ACE905863275CCC860AB679C7DF63032" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="8298CD2B9185C9D6828B16D53150B1E7" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="A4075CA21B440B43459D64297CCF6885" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="C7B9D6444744D86864D23500C8B8C6A9" italics="true">O. schmidti</emphasis>
</taxonomicName>
(vs. head width &lt;head length).
</paragraph>
<paragraph id="EF6A8F144FF2C6EF747BF691CADC577B">
By having no tympanum, the new species differs from
<taxonomicName id="27E192890CFF7E61842A9DCA4B78638E" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="8D453890E6B078BA27994B3D13DE82EF" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F055E3C585A41EE41C6C34B568F19D6D" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="CA1DDE27D511A22A0EF6B5C42B77A215" italics="true">O. major</emphasis>
</taxonomicName>
,
<taxonomicName id="D12365CE5741037099312BFF4DEA2C5C" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longmenmontis">
<emphasis id="98CDE707B02DB37C8ADC3241B09315EE" italics="true">O. longmenmontis</emphasis>
</taxonomicName>
,
<taxonomicName id="5E30F35C9AA97090824388CA9BDD1926" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="218FCA66A69EB6C9FAA48AA37B1E26A6" italics="true">O. sterlingae</emphasis>
</taxonomicName>
,
<taxonomicName id="985EA2188EB5243ED0A412777569F2CE" authorityName="Tian" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chuanbeiensis">
<emphasis id="AD4BBE026AB86B975EFBF371D0FC1B39" italics="true">O. chuanbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EA4701C62DE8AD9406D0AFB86CFD209C" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="8BF31EF1F71D98682509211C7A8A024B" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="E65E2DC5A65754434383E2029AD42F95" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="0E2FDE58FEAE619FFA154DCE5F2081B1" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="735A9C6C3A8E6D77845EC383FF8AB0C4" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="5CEC539BC27261559C44121F837C9D49" italics="true">O. pingii</emphasis>
</taxonomicName>
,
<taxonomicName id="60267A44ED8608477BC9E78EFFE5F885" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="886F752CCEEF672AA5DD27280B87B7A6" italics="true">O. popei</emphasis>
</taxonomicName>
,
<taxonomicName id="EDD58DB8C7D7B41D3E6A2F52E549382C" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puxiongensis">
<emphasis id="9BF02D755625212CA459F06168D3077F" italics="true">O. puxiongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="322924DC63D692B25E8A280DF49829A7" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="36A31CE022BAF48342745566CEAF116F" italics="true">O. schmidti</emphasis>
</taxonomicName>
,
<taxonomicName id="1336C195C7BA364149D6E9E4CCF5B625" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="2CC1F7DF19DA83082BB3DCE3B42D2189" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(vs. hidden),
<taxonomicName id="F23DF91B9B7BDB1C6BD7A4863C1A51E1" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="D28A0285E8E90AEAC0C22272B448D04A" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F17A9CDB16C67D54EA8D224EF80B4BE5" authorityName="Liu &amp; Hu" authorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="BB91F411BB3FC7931C93F203141A383A" italics="true">O. omeimontis</emphasis>
</taxonomicName>
(vs. concealed or slightly visible), and
<taxonomicName id="75BFCF513CA7ACD63B76746671E327E2" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="9DFA8F93E8913EF499DE5BE806E275DB" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
(vs. rather visible).
</paragraph>
<paragraph id="6966CCCFCDF75C425F73B7846A595A34">
By having 1 / 3 toe webbing, the new species differs from
<taxonomicName id="4279E2C4826AC67A8F1B909A3D1FC75A" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puxiongensis">
<emphasis id="78C0CADC68F8750EE5AA88354E8C5E5B" italics="true">O. puxiongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="BB43AB61EE49C0CF252A38804218FA50" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="89993EC8E550C548C1B55D8BA8ED9004" italics="true">O. schmidti</emphasis>
</taxonomicName>
(vs. no webbing),
<taxonomicName id="5365BCB24BBA4F0C412D9D2F47FDDD53" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longmenmontis">
<emphasis id="8AC8A87E8A60DC81A683C112DAC8EB95" italics="true">O. longmenmontis</emphasis>
</taxonomicName>
,
<taxonomicName id="56F5C6E503EA703A7081BF8F8A8B1BFA" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="412EF04BE0B09193CA09DE207DE8F7CE" italics="true">O. sterlingae</emphasis>
</taxonomicName>
,
<taxonomicName id="67BD403F3FE5964E2AF120603B9CC8A3" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="6ECEAE450B528A70C0EEDA53A33106D5" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6261F7ACFA67755A202626D36F7BFBD7" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="16688693604E9DE068415AF5A43C8C4B" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="980EB1FE47C911E4648AAAC6C59AEB52" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="7D1C3870C589C9CFBABE240169460719" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="BFE9402E51EC45504F69D29E4278D1E8" authorityName="Liu &amp; Hu" authorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="050562A57B9BE85548E55C198144E1F6" italics="true">O. omeimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="178909D730EF64567C242C09033F883E" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="A18983ACB8F8CDF9DE10165D4DFC3148" italics="true">O. pingii</emphasis>
</taxonomicName>
,
<taxonomicName id="A3906715C99A22E4213211469F3D6E70" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="7659D2B65E9EB57E2F1F70B599C110BD" italics="true">O. popei</emphasis>
</taxonomicName>
,
<taxonomicName id="4F19ABC503A6E012D5994125AA10EC93" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="0892DD91E280E7659A8B61D9E2709362" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
(vs. rudimentary),
<taxonomicName id="4AEE909002295DBE2CE75C156D5D03F1" authorityName="Fei &amp; Huang" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="FA143ECE0F5F287368F3987425A336B7" italics="true">O. xiangchengensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="30F6C5B5CDB863EF91301CEB91067704" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="5CACC2771ABD48DC1618387F1EC307B8" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(vs. well webbed).
</paragraph>
<paragraph id="A7EAE0BD4038BC1B4C45642E66629E7F">
By having triangular pattern between eyes, the new species differs from
<taxonomicName id="1D474682185C3E3DD265DB5924E40382" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="D0D6F70ECCA6A04502E9AEDEB69C3611" italics="true">O. rugosus</emphasis>
</taxonomicName>
,
<taxonomicName id="7DC5AD96BE675F058B7A4B1395369503" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="C5B1944502E2DDD065B95D4EA1CD8917" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6FBF22C6D3220AEA889A94F526F8FCDD" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="9FE7AC6CA2899F6E1F8BE20B71F760BE" italics="true">O. major</emphasis>
</taxonomicName>
,
<taxonomicName id="9085526FEB06385BC82819E46AD6C7EC" authorityName="Fei &amp; Huang" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="70A5EE21ADCF109E00B7671A05B54168" italics="true">O. xiangchengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="0A6BD8C3188E499AEE6BE4DDDDCB350C" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="CA83015A85464B76E03019BFE16A9E6D" italics="true">O. sterlingae</emphasis>
</taxonomicName>
,
<taxonomicName id="3D255A89370959ADEEE5D82777299812" authorityName="Tian" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chuanbeiensis">
<emphasis id="A7CA8B41C6F205695C6A4C132F523498" italics="true">O. chuanbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6F859C13FD66C5857CD48D8A9BB0891A" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="2E9E8F9232CC547A55DB3F298E148A28" italics="true">O. granulosus</emphasis>
</taxonomicName>
,
<taxonomicName id="0C1ADE8064F8EA883B1EC76178021B9E" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="6C2CF177A8AEBE512CD759F0D8A4335C" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="A416DDEE30BCFC297C1A1C4BB1F8AEDA" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="F1FD75D1A1093BA4AF862238C10789DE" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="86444DACE68A349D190CB9E4CFB909D1" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="02154BAC113047469EBF5883315AF267" italics="true">O. pingii</emphasis>
</taxonomicName>
,
<taxonomicName id="02D34C9C565C7387AAC681BDDE483BA7" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="2AAFEC46D8ACD7FC87E8D2A873D2A19D" italics="true">O. popei</emphasis>
</taxonomicName>
,
<taxonomicName id="A7B6AB0F2F2CA964FD45DDBEF12A8821" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="1039BE719EBAFBE33ED8ECA8F4E45C81" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="47812BA93841E0672F460012D8DEEAD2" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="45D888D3C86E3AF3CD83A3AA0D0D0D3A" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(vs. no triangular pattern).
</paragraph>
<paragraph id="C8555AFB995A2AC5E734A4098227ED3B">
By having middle spiny patches on the chest, the new species differs from
<taxonomicName id="D89415147A8999F10CE79B48DC581E6E" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="2FE8A20CCB689760A3EDB40CC60F1663" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="CA3A8DCDB88369B8C7D941E8ACE325E8" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="6C08BD7A14E60B365FE397F33B6E2047" italics="true">O. major</emphasis>
</taxonomicName>
,
<taxonomicName id="BB46B0BD30120E3F69CE547349D4AC38" authorityName="Fei &amp; Huang" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="B309D32A7FDB36301544D3B23610B89C" italics="true">O. xiangchengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="007F5AF74A452AC13C5789A136BEF352" authorityName="Tian" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chuanbeiensis">
<emphasis id="A59F805E63565C7D8176B973D5313DF6" italics="true">O. chuanbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2E489DC1508FBC21B8F8CC4212CD4B6C" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="74C65507014A9992D397659E8C0AD1C1" italics="true">O. granulosus</emphasis>
</taxonomicName>
,
<taxonomicName id="966D1501CA1FA9D246C859C8843211B5" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="7C9D5A1D65C15CCB068D18C804FAF308" italics="true">O. weigoldi</emphasis>
</taxonomicName>
,
<taxonomicName id="B9D138E2E4C2B0332B563CDDA27E1D4D" authorityName="Liu &amp; Hu" authorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="93C4CFA469978D8B1F444A57140A82F5" italics="true">O. omeimontis</emphasis>
</taxonomicName>
(vs. large patches),
<taxonomicName id="099CFD0C34C1ABEEAAE5F7253F5199EA" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="9635136DF25A99D296B5F417860C5DFF" italics="true">O. pingii</emphasis>
</taxonomicName>
,
<taxonomicName id="80BF61CB434BD07133728401905CD475" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="5C5C45D4B8771D270812174A475CAAC5" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
,
<taxonomicName id="86619B9CAAD17F561B0FDB17CD846466" authorityName="Ma, Yang &amp; Li" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="63DE8BD84523C6BC82D04F22640C5ED6" italics="true">O. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="696336E93F3C65C496DF42066304955C" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="6C7F6D7CCB372D6E1784CD55F4DA0452" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
(vs. relatively large patches),
<taxonomicName id="3982F0E7224C84C27C6784BDAB33DF42" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longmenmontis">
<emphasis id="03868205875C0A216DD91A3236119C11" italics="true">O. longmenmontis</emphasis>
</taxonomicName>
,
<taxonomicName id="93F844775974887A889C0F3D3167EDDF" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="2996982A8282470A1F9DF2250C5EAAD4" italics="true">O. sterlingae</emphasis>
</taxonomicName>
,
<taxonomicName id="20C4CABCF800A9116B569A7FBA03D2A1" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="A1118D4141AE0B6D267BD9BB7F888111" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="E3CD1ADCA9456827E4A9629FA1894941" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="675674E0C9424D4D4A4CE922E92E1184" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="F9924A1D940800B09B1B5A8106999C77" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="61B4E00FA257114EEE444E4552FACAE2" italics="true">O. popei</emphasis>
</taxonomicName>
(vs. small patches).
</paragraph>
<paragraph id="9707A3033E2E740CFD04A83628F8FC24">
By having dark bars on the limbs, the new species can differ from
<taxonomicName id="FC9C97827DA02DFB20ABCE90E0548148" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="927A760A3C06929C3FFBCF4C99237F67" italics="true">O. rugosus</emphasis>
</taxonomicName>
(vs. no or irregular),
<taxonomicName id="6DC172E5CB5C699F655A27D4FB06058B" authorityName="Fei &amp; Huang" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="286012D86D0682AF82534DD293A04F4C" italics="true">O. xiangchengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="984B577B58E97098F7C3991307B05D0F" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="DE2C2EC1243ADF40DC208D633CB78EC1" italics="true">O. pingii</emphasis>
</taxonomicName>
, and
<taxonomicName id="10CF3E258D63DF2E7434B540701E7343" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puxiongensis">
<emphasis id="35B811DEE3B7B487938DCC4047C11082" italics="true">O. puxiongensis</emphasis>
</taxonomicName>
(vs. no).
</paragraph>
<paragraph id="07C0839448A7C72E0CDAB6A4A440AFC2">
By having brown yellow or medium yellow scattered variable brown spots on the belly, the new species can differ from
<taxonomicName id="15BF232FD3E3590A279CC9F6E3CC2986" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="3A0B90F7DC2EB2AD1595037CC1957A90" italics="true">O. rugosus</emphasis>
</taxonomicName>
(creamy yellow or yellow, no spots),
<taxonomicName id="F611E5AE87467163EB0D1D28CD39695D" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="534A8D0B3A583C2CE3B18ED230685BA2" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
(creamy white without any spots),
<taxonomicName id="65D22E49D7816FDB8C9D8AF66395A2AE" authorityName="Fei &amp; Huang" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="D159168FC8EF8F91EAE2DBCE10AE05FA" italics="true">O. xiangchengensis</emphasis>
</taxonomicName>
(light brown, no spots),
<taxonomicName id="63B1135C5660F617CAB50BCE601B460D" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="C3277149664C50CF56599F4A15408544" italics="true">O. sterlingae</emphasis>
</taxonomicName>
(cream with dark marbling),
<taxonomicName id="F5BB3428A66D86C198E7E42EBA39C832" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="463DF6520B0AF9D8BACDC169242B6F5D" italics="true">O. granulosus</emphasis>
</taxonomicName>
(yellow-white or with fine light gray veins),
<taxonomicName id="9F68AB0F8AFBAF112C331A3D52D6A1E7" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="336FF5088F6FA6EA2C19F45730796706" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
(purplish with dark brown flecks),
<taxonomicName id="0AFCEEC285F24764F7E737ED93DA3971" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="249199A09B641792796A3AEB69D5CAFE" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
(grey brown, with few or without spots),
<taxonomicName id="BF3994A796F0C47D632032597E25D343" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="741D98CBB6C5F72DFB7B7066E129286A" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
(without dark spots),
<taxonomicName id="C43F1C0252819D1ACF64BFEEC81DDB51" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="pingii">
<emphasis id="D902EB0C40EDEC81355C5137D9DBD006" italics="true">O. pingii</emphasis>
</taxonomicName>
(gray-white, no spots),
<taxonomicName id="782E634341D7857A4A9DD4AF72FA6EE5" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="4B04A0FB2F6AB0FC40C31F8934598C1F" italics="true">O. popei</emphasis>
</taxonomicName>
(brown-red, fully covered with small gray-brown spots),
<taxonomicName id="B03BF147CC0C8A1FFE5B645DFBC58D6D" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puxiongensis">
<emphasis id="9539CFCC36AF9433C90CF8DA7F81FB5D" italics="true">O. puxiongensis</emphasis>
</taxonomicName>
(grayish-yellow, no spots),
<taxonomicName id="783C0917CB02A0B09345AC6D6B38C526" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="D3056B232F2410E14A67083FEB490FB4" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
(grayish-brown, no spots),
<taxonomicName id="AC107E37657F130A9F78F956C1ECE8D6" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="970B6CDEFCA168CC56B82B3B782E6248" italics="true">O. schmidti</emphasis>
</taxonomicName>
(entirely purple-yellow, no spots),
<taxonomicName id="DF82111A04D7E04BC87EC9BEB1AA8847" authorityName="Vogt" authorityYear="1924" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weigoldi">
<emphasis id="7C24D362FDC1723254EEC893202811EA" italics="true">O. weigoldi</emphasis>
</taxonomicName>
(light brown with dark cloudy spots on ventrolateral), and
<taxonomicName id="8748B88859AE7572F82F09A64A408332" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longmenmontis">
<emphasis id="9763391BF9ADE25C99509EC05EC43F81" italics="true">O. longmenmontis</emphasis>
</taxonomicName>
(flesh red and greyish-white with some black speckles).
</paragraph>
<paragraph id="3CDD68D4AFA2E48324C2B9F72F6B4337">
<taxonomicName id="44AE39D18CC7DD0B999EFBD97B464470" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis">
<emphasis id="DE8983C84C0EF1FB5BEF69B211DFE11E" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
is genetically closest to
<taxonomicName id="F80CD12A76CDE8F5F035C35D1E8916E0" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="AEF2E474B211554F03393D95A6A0FF05" italics="true">O. rugosus</emphasis>
</taxonomicName>
,
<taxonomicName id="31E4D7A67976C09C516713A5C1FB3EF3" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="3EDA3E09C154960E813681551BAD5FE2" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="E35D079EAC1CA97A729B9F4AB88E4C71" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="D1884514DC6158D70D5D028C1D68F9FE" italics="true">O. major</emphasis>
</taxonomicName>
. In addition to the morphological differences (Fig.
<figureCitation id="28623ECACDA02F2B4F9599AE2C45BB31" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Specimen photos of Oreolalax yanyuanensis sp. nov. and its relative species A, B dorsal and ventral view of the holotype CIB 25333 of O. rugosus C, D dorsal and ventral view of the topotype CIB 24678 of O. liangbeiensis E, F dorsal and ventral view of the topotype CIB 24695 of O. major G, H dorsal and ventral view of the topotype CIBSH 20230603020 of Oreolalax yanyuanensis sp. nov. Scale bar: equal to 10 mm." figureDoi="10.3897/zookeys.1212.122222.figure6" httpUri="https://binary.pensoft.net/fig/1134582">6</figureCitation>
; Suppl. material
<xref id="62CF3DD188727145E8FDBDA04EE2D36B" ref-type="supplementary-material" rid="S5">5</xref>
), the new species distinctly differs from these three in measurement proportions. The new species can differ from
<taxonomicName id="3D53A8BC4205C9E690531E97953332AE" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="F00B76BED3F9D5749A03E7401F94092A" italics="true">O. rugosus</emphasis>
</taxonomicName>
by having larger
<abbrev id="ABBRID0ECLAI" xlink_title="length of foot and tarsus">
<collectionCode id="EB670AB9FD49FCBD923D2BC4E9C6A5BC">LFT</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EGLAI" xlink_title="tibia length">TL</abbrev>
, and smaller IN,
<abbrev id="ABBRID0EKLAI" xlink_title="interorbital distance">IOD</abbrev>
,
<abbrev id="ABBRID0EOLAI" xlink_title="upper eyelid width">UEW</abbrev>
,
<abbrev id="ABBRID0ESLAI" xlink_title="tibia width">TW</abbrev>
. The new species distinctly differs from
<taxonomicName id="F5C8CDC9561A87D6D3143B9174E8E427" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="19E4C7CF51BC84EA3BF2B745C304B012" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
by having larger
<abbrev id="ABBRID0EBMAI" xlink_title="head length">HL</abbrev>
,
<abbrev id="ABBRID0EFMAI" xlink_title="head width">
<collectionCode id="AD5266094CAC184E6E3B9FE313C93B7F">HW</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EJMAI" xlink_title="eye diameter">
<collectionCode id="7683B2164C6625ECE996FA142C712CF4">ED</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0ENMAI" xlink_title="length of lower arm and hand">
<collectionCode id="4B69DC2F2277866852D4A7F438A15D0B">LAHL</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0ERMAI" xlink_title="tibia length">TL</abbrev>
,
<abbrev id="ABBRID0EVMAI" xlink_title="length of foot and tarsus">
<collectionCode id="0A08383EF45B863533FE06B2BA7A4D1B">LFT</collectionCode>
</abbrev>
and smaller
<abbrev id="ABBRID0EZMAI" xlink_title="snout length">SL</abbrev>
,
<abbrev id="ABBRID0E4MAI" xlink_title="interorbital distance">IOD</abbrev>
. The new species distinctly differs from
<taxonomicName id="0B59F8550AF6F39C4BCDE4305DAC5D78" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="82313991BB6811AAB4883B69E2FD7AE6" italics="true">O. major</emphasis>
</taxonomicName>
by having larger
<abbrev id="ABBRID0EMNAI" xlink_title="head width">
<collectionCode id="9857EB2514A08A4D0BCBAE8EE56A5B23">HW</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EQNAI" xlink_title="eye diameter">
<collectionCode id="34AAF7C643F171E60F6A5F47B010F883">ED</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EUNAI" xlink_title="tibia length">TL</abbrev>
,
<abbrev id="ABBRID0EYNAI" xlink_title="length of foot and tarsus">
<collectionCode id="A4BCBABC0A461087E06C8E6F9FC96BBC">LFT</collectionCode>
</abbrev>
and smaller
<abbrev id="ABBRID0E3NAI" xlink_title="snout length">SL</abbrev>
, IN,
<abbrev id="ABBRID0EAOAI" xlink_title="interorbital distance">IOD</abbrev>
(Table
<tableCitation id="6C613A133982D979700A65BE4750757B" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric comparisons between Oreolalax yanyuanensis sp. nov. and its relatives. Shaded values represent a ratio of body measurements to SVL that differs between these three species and the new species." httpUri="http://table.plazi.org/id/EA6808C88724DA8B88B55CB20200A618" tableUuid="EA6808C88724DA8B88B55CB20200A618">2</tableCitation>
).
</paragraph>
<caption id="EA6808C88724DA8B88B55CB20200A618" ID-Table-UUID="EA6808C88724DA8B88B55CB20200A618" httpUri="http://table.plazi.org/id/EA6808C88724DA8B88B55CB20200A618" startId="T2" targetIsTable="true">
<paragraph id="913E094237A33346A15516EB61C3F15D">
<label id="1902255FA998331075015398231ECA24">Table 2.</label>
</paragraph>
<paragraph id="4C02F1AA6B9F7680CB65D592EF035AF9">
Morphometric comparisons between
<taxonomicName id="6EBBA9820E05CB00E7856E86288A24A4" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="030BC5C1860F6E26B533110FB779F7AD" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6106F9AAB8DAC8813EC4F2E62E744304" rank="species">sp. nov.</taxonomicNameLabel>
and its relatives. Shaded values represent a ratio of body measurements to
<abbrev id="ABBRID0E3OAI" xlink_title="snoutvent length">SVL</abbrev>
that differs between these three species and the new species.
</paragraph>
</caption>
<paragraph id="D89E45BC4EA7595B92ED1A39039F02F4">
<table id="TID0ELSAI" rules="all">
<tbody id="4E0A2D186E120E2FE7B51D9A61012FE5">
<tr id="26652AB79315D9C3880D894B88BCDC6D">
<th id="11F84D10D26CDF00BE97F82B0C53BD40" colspan="1" rowspan="2">
Measurements /
<abbrev id="ABBRID0EJPAI" xlink_title="snoutvent length">SVL</abbrev>
(%)
</th>
<th id="50B68F1DDA5E05B7AC48C193D63816C7" colspan="1" rowspan="1">
<taxonomicName id="FB3D30B941694BFAD98857078CE7A4CE" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="8E7BACE889901D6E5868D148AE8925A1" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8DDDEA7A878D63C8F1230AAA344086F" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<emphasis id="F53683B68CFD2285C3309E6CE29E10F8" italics="true">
<th id="61E6C6BC1C254EC8F0EA291E4AB27C92" colspan="1" rowspan="1">
<taxonomicName id="1FB5E20E861C0E9C02A414561237A2CC" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">O. rugosus</taxonomicName>
</th>
<th id="4B387C8AAF7F2536629927C3E63F5812" colspan="1" rowspan="1">
<taxonomicName id="B006BAA0632D8D477234936BA013A01E" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">O. liangbeiensis</taxonomicName>
</th>
<th id="286131E5DBB846A7A15B674CD1FD8C3D" colspan="1" rowspan="1">
<taxonomicName id="A2E5770A67D7B260D00145BD9B58F681" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">O. major</taxonomicName>
</th>
</emphasis>
</tr>
<tr id="3BCA0999C4B9F8A71834280DD18D9A37">
<th id="AA63894522329CBA5F35659ACE7FCCFD" colspan="1" rowspan="1">8 ♂♂</th>
<th id="0DFEABA49AE416F5A7EF703526A79BDB" colspan="1" rowspan="1">10 ♂♂</th>
<th id="22CAB5B74A1788064E818475BB244773" colspan="1" rowspan="1">20 ♂♂</th>
<th id="B6AAD83AC10B13ABF68333B387D5D0B8" colspan="1" rowspan="1">6 ♂♂</th>
</tr>
<tr id="4604E15BA44D19BB8A540F1B15891FE0">
<td id="3180C9E4347C9E4DE4DA133CC9AB0440" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2RAI" xlink_title="head length">HL</abbrev>
</td>
<td id="03920D4462912B6AFFBF4796DA2E6428" colspan="1" rowspan="1">35.9 (34.237.4)</td>
<td id="70B7F23BC79B77C56CB3A73F20FE5276" colspan="1" rowspan="1">35.4</td>
<td id="316B56E7136248A471A16FF4C0FEE5BB" colspan="1" rowspan="1" style="background: #d5d6d8">33.3</td>
<td id="25B7C4A1B9CC6CE8E00E63D5B00C8FAE" colspan="1" rowspan="1">35.4</td>
</tr>
<tr id="2B929C210898E40B5B3C70253C64D5C8">
<td id="767FEF91844EED66562A6DC2783CE618" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERSAI" xlink_title="head width">HW</abbrev>
</td>
<td id="BC05DE7058909329CFF448A2F715D02C" colspan="1" rowspan="1">37.6 (36.639.5)</td>
<td id="FB82DF294ABADFEACA5095C276176053" colspan="1" rowspan="1">37.5</td>
<td id="20F53E7A529B4AA6CE80A938A6318AF2" colspan="1" rowspan="1" style="background: #d5d6d8">35.2</td>
<td id="DA6191EE7D94134E084D25B857BFEBBD" colspan="1" rowspan="1" style="background: #d5d6d8">35.8</td>
</tr>
<tr id="7B1BF457A2D28A7D0364790FD4C62B9C">
<td id="5E2139FBE876A11A1DFE0CBF94C8B4B9" colspan="1" rowspan="1">
<abbrev id="ABBRID0EITAI" xlink_title="snout length">SL</abbrev>
</td>
<td id="021D81811E43F08EDB7D2C68C4ACA872" colspan="1" rowspan="1">14.3 (13.215.3)</td>
<td id="092E2432FDD6626162DE090B46539DD3" colspan="1" rowspan="1">14.6</td>
<td id="95B31E9B26F2539FE7ACF218795BD4A2" colspan="1" rowspan="1" style="background: #d5d6d8">15.5</td>
<td id="6B0F6F498BDFC7417205839B870F2F04" colspan="1" rowspan="1" style="background: #d5d6d8">15.5</td>
</tr>
<tr id="7D86222DD86B728A51B1BB8A393286B7">
<td id="A553DA3EB5FC3D48F33A6A9EA88698F7" colspan="1" rowspan="1">
<abbrev id="ABBRID0E6TAI" xlink_title="internarial distance">IND</abbrev>
</td>
<td id="60A01E2C1329D5794E80CD918A635D6C" colspan="1" rowspan="1">9.0 (7.410.3)</td>
<td id="DB042EFC89834BF7B7CC3FAB00BE9DE8" colspan="1" rowspan="1" style="background: #d5d6d8">10.4</td>
<td id="19D3C52D401C1E048673A1BB0CE50618" colspan="1" rowspan="1">9.8</td>
<td id="BBCBD0C34CFFE405608B1E39787815AC" colspan="1" rowspan="1" style="background: #d5d6d8">10.4</td>
</tr>
<tr id="0B37F7D4BAB298C6001B34274BD3598A">
<td id="E7535D041503B0A872C3A5DEC67497C5" colspan="1" rowspan="1">
<abbrev id="ABBRID0EWUAI" xlink_title="interorbital distance">IOD</abbrev>
</td>
<td id="818634ED2E286F26A5EE59372755CF22" colspan="1" rowspan="1">10.1 (9.210.6)</td>
<td id="62AF64224A1F46D360BB247B21AFFA3D" colspan="1" rowspan="1" style="background: #d5d6d8">11.5</td>
<td id="C06D333F23B78BDD7F3A48CD17BA346E" colspan="1" rowspan="1" style="background: #d5d6d8">10.9</td>
<td id="7C7717E48C2569D06FACA7222B3CB5A1" colspan="1" rowspan="1" style="background: #d5d6d8">10.8</td>
</tr>
<tr id="40A11B121D7D7B58AB029342B0073F6D">
<td id="EB7EF288A2630169BF695E5AF2221C80" colspan="1" rowspan="1">
<abbrev id="ABBRID0EOVAI" xlink_title="eye diameter">ED</abbrev>
</td>
<td id="CB65A58CB21D4C5A4E9E5D841FC1B02F" colspan="1" rowspan="1">11.9 (11.512.5)</td>
<td id="F17C3236F5202DFAF1FBE837AD213DF9" colspan="1" rowspan="1">12.2</td>
<td id="BADE35FB9C0E95B1BAB311D4485B3969" colspan="1" rowspan="1" style="background: #d5d6d8">10.1</td>
<td id="CAA0E1AB58D1F71BD7F5B22E072F21B1" colspan="1" rowspan="1" style="background: #d5d6d8">10.5</td>
</tr>
<tr id="F4A32DE2E679F37B2DC27DB9FF2F8404">
<td id="1C393F6263C78738B32AC1D326CD03B2" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFWAI" xlink_title="upper eyelid width">UEW</abbrev>
</td>
<td id="FBFED75B62ECAEB8C710F3FB495ACDBC" colspan="1" rowspan="1">8.4 (6.39.4)</td>
<td id="ADB42AA2D6E463F532B983E1D182E7A5" colspan="1" rowspan="1" style="background: #d5d6d8">9.9</td>
<td id="0686CE7ABB4B75BA753B16639FD7E222" colspan="1" rowspan="1">9.2</td>
<td id="65ECD9462D538E81045ACFF6B98FF486" colspan="1" rowspan="1">9.0</td>
</tr>
<tr id="53E55DE345142A9E53E138F301D95028">
<td id="61169B4294BBD34C760F87EE919B3CEB" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2WAI" xlink_title="forearm width">FAW</abbrev>
</td>
<td id="9A523814A8885A7FF603035DE65C69E3" colspan="1" rowspan="1">10.3 (8.912.4)</td>
<td id="B53A8466A53B8BC20E8CCE0AA7181ADE" colspan="1" rowspan="1">11.5</td>
<td id="CF1EA37D56EAEE289DF7F45BB1FD8F14" colspan="1" rowspan="1">10.9</td>
<td id="6F740DFF9D5EEDC17C08454DF46BA1EC" colspan="1" rowspan="1">10.8</td>
</tr>
<tr id="5023D256934BF9F6FE8DD4EFB65DB1AB">
<td id="6A6BFEB88F78D5523FB50D537B14B545" colspan="1" rowspan="1">
<abbrev id="ABBRID0EQXAI" xlink_title="length of lower arm and hand">LAHL</abbrev>
</td>
<td id="BC7DA34C2A8189C926C928C279A8E45F" colspan="1" rowspan="1">54.6 (51.258.0)</td>
<td id="F5107E032505AC74CA948875987A37BF" colspan="1" rowspan="1">51.6</td>
<td id="3C32A650F0FA1A408740EC9A188CA457" colspan="1" rowspan="1" style="background: #d5d6d8">51.0</td>
<td id="BCC056E5214D51CA06CFDECF4C19930E" colspan="1" rowspan="1">55.0</td>
</tr>
<tr id="8049C2DC016BD2A11181E891A91BA390">
<td id="10B5F1B6577F38C6E507A90C371C6A61" colspan="1" rowspan="1">
<abbrev id="ABBRID0EGYAI" xlink_title="tibia length">TL</abbrev>
</td>
<td id="E282E21C9BF8A0EA56F544D06766864A" colspan="1" rowspan="1">51.3 (49.653.2)</td>
<td id="1A54109551F155D1B954126B4183A3DB" colspan="1" rowspan="1" style="background: #d5d6d8">47.8</td>
<td id="4FE7A9E87607CADA8C0FDEF440ECEF9A" colspan="1" rowspan="1" style="background: #d5d6d8">45.0</td>
<td id="E1DA885C66C3FEC3620489688C5ECFA9" colspan="1" rowspan="1" style="background: #d5d6d8">48.0</td>
</tr>
<tr id="EFD02C29F940F1252661B8448FE2CBC1">
<td id="2FFF164766E3FE28934F93F0F611ADF4" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5YAI" xlink_title="tibia width">TW</abbrev>
</td>
<td id="33AF6CF816408E4EAC022F3B30821141" colspan="1" rowspan="1">11.3 (9.213.6)</td>
<td id="89123513892A174E06DFE4507D3BFF7B" colspan="1" rowspan="1" style="background: #d5d6d8">13.7</td>
<td id="AB45B29F9463E75F1FD38C9A22499754" colspan="1" rowspan="1">12.3</td>
<td id="81F470C94DAAA2DB8EF7441716C2FAAD" colspan="1" rowspan="1">11.1</td>
</tr>
<tr id="1E329A7BBB31BD3A8A408FB059A39976">
<td id="484A5A7F597BDC686D352CDF17C3C782" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUZAI" xlink_title="length of foot and tarsus">LFT</abbrev>
</td>
<td id="DA23585317E9241C691F95B511E21C20" colspan="1" rowspan="1">77.5 (74.380.2)</td>
<td id="2CA8FEC0E4227BF41A674B358985E03F" colspan="1" rowspan="1" style="background: #d5d6d8">67.5</td>
<td id="00B30484299506DA53674E23A6A0CDED" colspan="1" rowspan="1" style="background: #d5d6d8">69.0</td>
<td id="0E0DBD2C3D18BB02AB422A2B29304444" colspan="1" rowspan="1" style="background: #d5d6d8">73.8</td>
</tr>
<tr id="974192B0C60EC59D78AAF01C0E258FF0">
<td id="EEAFAD67574704157FA3B28BDA39C5FD" colspan="1" rowspan="1">
<abbrev id="ABBRID0EM1AI" xlink_title="foot length">FL</abbrev>
</td>
<td id="40372B7853342E25388E629C6BC48D6A" colspan="1" rowspan="1">50.8 (47.154.6)</td>
<td id="0F5EA466B9BE527FB615F6A0915C7BE3" colspan="1" rowspan="1">48.1</td>
<td id="A8B101F759729627B8DCB1D3C6945D65" colspan="1" rowspan="1">48.7</td>
<td id="F8E79015ABAC49523BA5EBBFA6CC2391" colspan="1" rowspan="1">50.9</td>
</tr>
</tbody>
</table>
</paragraph>
<caption id="E59C77A7ADBA9EF11443496D80BE42D4" ID-DOI="10.3897/zookeys.1212.122222.figure6" ID-arpha="44B60698-97CD-5D71-A6D4-82F9E8A9BDB5" httpUri="https://binary.pensoft.net/fig/1134582" startId="F6">
<paragraph id="B05E57916636C1879EB75D8DD111300D">
<label id="6996D5E5B18A156E90A4BF470F4747CE">Figure 6.</label>
</paragraph>
<paragraph id="76A3ECC10EA256891C71C122A7FD4128">
Specimen photos of
<taxonomicName id="B8109CE11432D4A31C49F7F4D6695450" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="4E68E051CF8FD2B4B1003DFB44D0B19B" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="566981EBA6CED54B2BACEA97EC9BA619" rank="species">sp. nov.</taxonomicNameLabel>
and its relative species
<emphasis id="9A35FD9C839231D94A56A0FD2907065F" bold="true">A, B</emphasis>
dorsal and ventral view of the holotype CIB 25333 of
<taxonomicName id="25D7CF7FCC2D81E45D210FD5E37FD665" baseAuthorityName="Liu" baseAuthorityYear="1943" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rugosus">
<emphasis id="5ABF3D13BC2F27867D0155A090F2437E" italics="true">O. rugosus</emphasis>
</taxonomicName>
<emphasis id="083C7C13DF32EB15291B32E7B2B7844F" bold="true">C, D</emphasis>
dorsal and ventral view of the topotype CIB 24678 of
<taxonomicName id="12B7E8024941CAF56CB08C288EC5BE94" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="liangbeiensis">
<emphasis id="8DB68CCD6F17130F43022D0FC53A501D" italics="true">O. liangbeiensis</emphasis>
</taxonomicName>
<emphasis id="4591CDDAB9268B23693A8CF6AA0C42D7" bold="true">E, F</emphasis>
dorsal and ventral view of the topotype CIB 24695 of
<taxonomicName id="D8587D2832FD92743183309B45B18639" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="564E36995D008844A2EE2E55B821EDEE" italics="true">O. major</emphasis>
</taxonomicName>
<emphasis id="8E0093CA20B929205770CB9DC964295F" bold="true">G, H</emphasis>
dorsal and ventral view of the topotype CIBSH 20230603020 of
<taxonomicName id="044AC6677257D92AC51B627F4C866C6C" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="16730A409D428DEE2BDC3DD350DB37B5" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9059B85C915FEA10CFC565263AC451" rank="species">sp. nov.</taxonomicNameLabel>
Scale bar: equal to 10 mm.
</paragraph>
</caption>
<paragraph id="4B4405505C615D0A1DE29E71ED1CB764">
In elevational distribution, the new species (occurring between
<quantity id="4396322B6A5713CECBDC67DAD0B6C01A" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" unit="m" value="3100.0" valueMax="3200.0" valueMin="3000.0">30003200 m</quantity>
) can be distinguished from some
<taxonomicName id="7DFF45D01D4F96BB04B35B73C0CE97C8">
<emphasis id="FA369848F131AA3066783769F8D45839" italics="true">Oreolalax</emphasis>
species
</taxonomicName>
occurring below
<quantity id="52CA70C879AB3B093B97B6C59BC9A8F3" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="m" value="3000.0">3000 m</quantity>
(a. s. l.) as follows:
<taxonomicName id="17315DCFCE9EC714B6F705265B23BB15" baseAuthorityName="Liu and Hu" baseAuthorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="major">
<emphasis id="8D28BEEAF9F4D7CB793C3C2A250CD108" italics="true">O. major</emphasis>
</taxonomicName>
(vs.
<quantity id="EE5F99E7032C9C88F71937FE907832CC" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">16002000 m</quantity>
),
<taxonomicName id="0936931A852E20FD64D7FF63F94F46C0" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="longmenmontis">
<emphasis id="813FF5AF538BDBDB94DF79F6AD17BD40" italics="true">O. longmenmontis</emphasis>
</taxonomicName>
(vs.
<quantity id="DF6F2750C575180D94A725B3CF3AB6DB" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">13001450 m</quantity>
),
<taxonomicName id="033A9790B5684FB87F6C02066A30FC6A" authorityName="Tian" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chuanbeiensis">
<emphasis id="01CDD4D426601C63DAED476B3A445BD3" italics="true">O. chuanbeiensis</emphasis>
</taxonomicName>
(vs.
<quantity id="0455A233470DAF69771FD4EE7A56C2B0" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">20002200 m</quantity>
),
<taxonomicName id="3EFD93E8224E2BEF5A4C765968F1C14A" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="2CB85830C9C1BAA2ECCDFB98AB89F38C" italics="true">O. granulosus</emphasis>
</taxonomicName>
(vs.
<quantity id="4AE353EB241CB2D67B016027F55DB0E4" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="2.45" metricValueMin="2.3" unit="m" value="2375.0" valueMax="2450.0" valueMin="2300.0">23002450 m</quantity>
),
<taxonomicName id="8E92300E2BB54B67F42713A95D6CBDC5" authorityName="Ma, Yang &amp; Li" authorityYear="1983" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="2390FC0E2735353305C6F8118229934D" italics="true">O. jingdongensis</emphasis>
</taxonomicName>
(vs.
<quantity id="220319A9C69E22F2B988F0A28F5D5475" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="2.45" metricValueMin="2.3" unit="m" value="2375.0" valueMax="2450.0" valueMin="2300.0">23002450 m</quantity>
),
<taxonomicName id="C06BD33BB1A1B455229950A20078B1C1" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lichuanensis">
<emphasis id="E1179C9E8B8267A9527A38E46266565F" italics="true">O. lichuanensis</emphasis>
</taxonomicName>
(vs.
<quantity id="B6A8597AED5E448F0E1211CA63680299" metricMagnitude="3" metricUnit="m" metricValue="1.815" metricValueMax="1.8399999999999999" metricValueMin="1.7899999999999998" unit="m" value="1815.0" valueMax="1840.0" valueMin="1790.0">17901840 m</quantity>
),
<taxonomicName id="02483F9BC8AEEE3BD04C1EA15D141905" authorityName="Wu, Zhao, Inger &amp; Shaffer" authorityYear="1993" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="034A1256CD2E872F4A69B12490537232" italics="true">O. multipunctatus</emphasis>
</taxonomicName>
(vs.
<quantity id="450525EDC0E1C5156C3C851E5888412F" metricMagnitude="3" metricUnit="m" metricValue="1.86" metricValueMax="1.92" metricValueMin="1.8" unit="m" value="1860.0" valueMax="1920.0" valueMin="1800.0">18001920 m</quantity>
),
<taxonomicName id="3D29AE72A49D46AE9ABF95A449E36B75" authorityName="Fei &amp; Ye" authorityYear="1999" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nanjiangensis">
<emphasis id="8179DB984F1804A803BEE280D2DB7A28" italics="true">O. nanjiangensis</emphasis>
</taxonomicName>
(vs.
<quantity id="3D5B216C8F9539F4A11244048F38524B" metricMagnitude="3" metricUnit="m" metricValue="1.728" metricValueMax="1.8559999999999999" metricValueMin="1.6" unit="m" value="1728.0" valueMax="1856.0" valueMin="1600.0">16001856 m</quantity>
),
<taxonomicName id="655AD10BA92E3227846692EAEE74608D" authorityName="Liu &amp; Hu" authorityYear="1960" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="42F7C8831A0E5EFD4345995110C16FCE" italics="true">O. omeimontis</emphasis>
</taxonomicName>
(vs.
<quantity id="507FF9DD0F27BCED38AE25B9483A4322" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.8" metricValueMin="1.05" unit="m" value="1425.0" valueMax="1800.0" valueMin="1050.0">10501800 m</quantity>
),
<taxonomicName id="C01ECBB5AE4ECCDD1058214D986FA490" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="popei">
<emphasis id="9E0BD20EA83697EE371DDAE1F78A5940" italics="true">O. popei</emphasis>
</taxonomicName>
(vs.
<quantity id="17471D84D09749B74ED08A151E633C69" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">10002000 m</quantity>
),
<taxonomicName id="6471D3AC325D69DF0815E74D51A2093E" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="puxiongensis">
<emphasis id="3ABAA4C5FE660AA7C0A75F54715BB4CE" italics="true">O. puxiongensis</emphasis>
</taxonomicName>
(vs.
<quantity id="E9EA04A0EB22E9648F77D995E3C1C16E" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" unit="m" value="2750.0" valueMax="2900.0" valueMin="2600.0">26002900 m</quantity>
),
<taxonomicName id="C1A9C10AA31AB9D81D567AC985D363AE" authorityName="Liu, Hu &amp; Fei" authorityYear="1979" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="C3120FB2627F155074251D29D11E4BE5" italics="true">O. rhodostigmatus</emphasis>
</taxonomicName>
(vs.
<quantity id="382255949183666DA119C91A8F28DE3F" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.7899999999999998" metricValueMin="0.7" unit="m" value="1245.0" valueMax="1790.0" valueMin="700.0">7001790 m</quantity>
),
<taxonomicName id="613FFE7E45B9227205539A4594D734E8" baseAuthorityName="Liu" baseAuthorityYear="1947" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="1639067E58504603892D657C75A0EFB8" italics="true">O. schmidti</emphasis>
</taxonomicName>
(vs.
<quantity id="7BEF58465B7C583245DD66760424E00E" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" unit="m" value="2050.0" valueMax="2400.0" valueMin="1700.0">17002400 m</quantity>
), and
<taxonomicName id="2C1E6532F506302EEAB78DC92FE26777" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sterlingae">
<emphasis id="5151576A1A7E76FAF50AD8D35BCC98D6" italics="true">O. sterlingae</emphasis>
</taxonomicName>
(vs.
<quantity id="01E8F9147AC130309C1AC33CBC17AB5C" metricMagnitude="3" metricUnit="m" metricValue="2.9" unit="m" value="2900.0">2900 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELCBI" type="etymology">
<paragraph id="16D704F03A87BECA48F7CCC6579C74E3">
<heading id="2616B12D5961B3C4E36706A95222D86A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="57186B0B8ED748AA350B0C86F126FFB5">
The specific epithet “ yanyuan ” refers to the type locality of the species, Yanyuan County,
<collectingRegion id="C5385A6E466CA82F8783D5590C8ED650" country="China" name="Sichuan">Sichuan Province</collectingRegion>
. We suggested the common name as “ Yanyuan toothed toad ”, and the Chinese name as “ Yan Yuan Chi Chan (盐源齿蟾) ”.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQCBI" type="distribution">
<paragraph id="640859DB8448701B98F929AD533F5ED4">
<heading id="77289AB6044843C53781E9812E13E864" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="54A346DD777954BF28BFBF05C6F55C8C">
<taxonomicName id="46257247888AA4A3ED538FC8AD4C9711" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="52AC40D2397598D062CBD52E14845C7D" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24003FC8D25FD46844F45AB09DA46015" rank="species">sp. nov.</taxonomicNameLabel>
is currently only known from the type locality, Shuhe town, Yanyuan county,
<collectingRegion id="4BBB8085C6CCBE6245866135292C93C2" country="China" name="Sichuan">Sichuan Prov.</collectingRegion>
,
<collectingCountry id="FA9D73C68C591431D7C3173716647E21" name="China">China</collectingCountry>
at elevations of
<quantity id="6F99C0B976249ABAD6795F7986C2A73A" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" unit="m" value="3100.0" valueMax="3200.0" valueMin="3000.0">30003200 m</quantity>
. The new species inhabits subtropical alpine scrub and swamp, and was found in small montane streams (Fig.
<figureCitation id="649B7495933FE91633C1DF028A0D8D9E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitats of Oreolalax yanyuanensis sp. nov. in southeastern Hengduan Mountains region, Sichuan Province, China A, B, C adults habitats D tadpoles habitat." figureDoi="10.3897/zookeys.1212.122222.figure7" httpUri="https://binary.pensoft.net/fig/1134583">7</figureCitation>
). The breeding season is currently uncertain; it is speculated that it breeds in April or May based on the tadpole development stage. Four sympatric amphibian species (
<taxonomicName id="0B0DC39A16757284D5E577059C466D9D" authority="Boulenger, 1905" authorityName="Boulenger" authorityYear="1905" class="Amphibia" family="Bombinatoridae" genus="Bombina" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="maxima">
<emphasis id="3E0BD3423EAD1C9A3DAE4BF6A5A8B503" italics="true">Bombina maxima</emphasis>
<bibRefCitation id="9CE95DA4C41579012DE6C6A5E522AD7B" DOI="10.1080/03745480509442817" author="Boulenger GA" refId="B2" refString="Boulenger GA (1905) Description of a new batrachian of the genus Bombinator from Yunnan. Annals and Magazine of Natural History, Series 7, 15: 188 - 190. https: // doi. org / 10.1080 / 03745480509442817" year="1905">Boulenger, 1905</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="3A96D1F08249E3CE56BD9CF39778837F" DOI="10.1080/03745480509442817" author="Boulenger" firstAuthor="Boulenger" refId="B2" refString="Boulenger GA (1905) Description of a new batrachian of the genus Bombinator from Yunnan. Annals and Magazine of Natural History, Series 7, 15: 188190. https://doi.org/10.1080/03745480509442817" year="1905">Boulenger 1905</bibRefCitation>
),
<taxonomicName id="08FD5720A6D392B0A70E73B1D340965B" authority="Liu, 1946" authorityName="Liu" authorityYear="1946" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="chaochiaoensis">
<emphasis id="DB79DD3DF89EBA1531D63F21D4F087A5" italics="true">Rana chaochiaoensis</emphasis>
<bibRefCitation id="9103C0E6712DA088A6E9BF08E616933D" author="Liu CC" journalOrPublisher="Journal of the West China Border Research Society, Series B" pagination="7-14" refId="B24" refString="Liu CC (1946) A new woodfrog Rana chaochiaoensis with a discussion of its allied species, from West China. Journal of the West China Border Research Society, Series B 16: 7 - 14." title="A new woodfrog Rana chaochiaoensis with a discussion of its allied species, from West China." volume="16" year="1946">Liu, 1946</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="7BFE41C754081659EADE9809F0261B76" author="Liu" firstAuthor="Liu" journalOrPublisher="Journal of the West China Border Research Society, Series B" pagination="7-14" refId="B24" refString="Liu CC (1946) A new woodfrog Rana chaochiaoensis with a discussion of its allied species, from West China. Journal of the West China Border Research Society, Series B 16: 714." title="A new woodfrog Rana chaochiaoensis with a discussion of its allied species, from West China." volume="16" year="1946">Liu 1946</bibRefCitation>
),
<taxonomicName id="CDD7A92D09E6508958304A72F80B3A5C" authority="Ye &amp; Fei, 1995" authorityName="Ye &amp; Fei" authorityYear="1995" class="Amphibia" family="Megophryidae" genus="Panophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="1D3F5CF3D8AE6E6727ABAF29C80D94F2" italics="true">Panophrys binchuanensis</emphasis>
<bibRefCitation id="71BB3D7F3A1F2258980ACC30BB1BC457" author="Ye CY &amp; Fei L" refId="B41" refString="Ye CY, Fei L (1995) Taxonomic studies on the small type Megophrys in China including descriptions of the new species (subspecies) (Pelobatidae: genus Megophrys). Acta Herpetologica Sinica 4 - 5: 72 - 81." year="1995">Ye &amp; Fei, 1995</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E13F2755525C87C58335D2661A78DF3E" author="Ye and Fei" firstAuthor="Ye" refId="B41" refString="Ye CY, Fei L (1995) Taxonomic studies on the small type Megophrys in China including descriptions of the new species (subspecies) (Pelobatidae: genus Megophrys). Acta Herpetologica Sinica 45: 7281." year="1995">Ye and Fei 1995</bibRefCitation>
) and
<taxonomicName id="0F2C6D19F8716A245B67DB64C6E9F226" authority="Dubois, 1987" authorityName="Dubois" authorityYear="1987" class="Amphibia" family="Dicroglossidae" genus="Nanorana" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sichuanensis">
<emphasis id="62F26F1EF765D261F0FB3DDD9EA1AD76" italics="true">Nanorana sichuanensis</emphasis>
<bibRefCitation id="A3259CE55D656E132184BE5DFC0EC0D4" author="Dubois A" journalOrPublisher="Paris" pagination="7-95" refId="B5" refString="Dubois A (1987) [&quot; 1986 &quot;] Miscellanea taxinomica batrachologica (I). Alytes. Paris 5: 7 - 95." title="[“ 1986 ”] Miscellanea taxinomica batrachologica (I). Alytes." volume="5" year="1987">Dubois, 1987</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="1E999B42BE146E6482337BC4C2A1EAE3" author="Dubois" firstAuthor="Dubois" journalOrPublisher="Paris" pagination="7-95" refId="B5" refString="Dubois A (1987) [“ 1986 ”] Miscellanea taxinomica batrachologica (I). Alytes. Paris 5: 795." title="[“ 1986 ”] Miscellanea taxinomica batrachologica (I). Alytes." volume="5" year="1987">Dubois 1987</bibRefCitation>
“ 1986 ” )) were found in the same habitat.
</paragraph>
<caption id="7C683306CE7E16F0E0A0FF9D648F6D5F" ID-DOI="10.3897/zookeys.1212.122222.figure7" ID-arpha="C9BFEE5D-1C2D-5C8A-A7BF-AFF6AFF36131" httpUri="https://binary.pensoft.net/fig/1134583" startId="F7">
<paragraph id="8026F77E1BEB20DA41F5963F548E709F">
<label id="69293C65AB0BB6B3B5209D5C71937854">Figure 7.</label>
</paragraph>
<paragraph id="ECF3548717218C08C441128AAAEEE85A">
Habitats of
<taxonomicName id="6E569DB6F509470DF5D70D2B20291496" authorityName="Hou &amp; Zheng &amp; Yu &amp; Wang &amp; Chen &amp; Xie" authorityYear="2024" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yanyuanensis" status="sp. nov.">
<emphasis id="E71921AD0AF18AF8EFC0B3DD15726AB5" italics="true">Oreolalax yanyuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B08CEABDC26320212078B33205D159B" rank="species">sp. nov.</taxonomicNameLabel>
in southeastern Hengduan Mountains region, Sichuan Province, China
<emphasis id="A31EDDA7D120DD67812EA410CC42FA1C" bold="true">A, B, C</emphasis>
adults habitats
<emphasis id="D5309E608E30918E4563A5B4F606117E" bold="true">D</emphasis>
tadpoles habitat.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>