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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1039.64740" ID-GBIF-Dataset="8fe0041e-40a6-4fd0-a674-948f3f919703" ID-PMC="PMC8159918" ID-Pensoft-Pub="1313-2970-1039-1" ID-Pensoft-UUID="FA2622A854585C59A7E8182A4E66C6DF" ID-PubMed="34084064" ID-ZooBank="3487C3570FAC4907A5C0C0A0C801C6E3" ModsDocID="1313-2970-1039-1" checkinTime="1621541808324" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2021" docId="F3A641DFB6F4528BBAB1E012F1B717C3" docLanguage="en" docName="ZooKeys 1039: 1-108" docOrigin="ZooKeys 1039" docPubDate="2021-05-20" docSource="http://dx.doi.org/10.3897/zookeys.1039.64740" docTitle="Pyrrhalta semifulva species group" docType="treatment" docVersion="4" id="FA2622A854585C59A7E8182A4E66C6DF" lastPageNumber="1" masterDocId="FA2622A854585C59A7E8182A4E66C6DF" masterDocTitle="Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei" masterLastPageNumber="108" masterPageNumber="1" pageNumber="1" updateTime="1668152740594" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4358-7211</mods:nameIdentifier>
<mods:affiliation>Mendel University in Brno, Department of Zoology, Fisheries, Hydrobiology and Apiculture, Zemĕdĕlska 1, 613 00, Brno, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-20</mods:number>
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<mods:detail type="volume">
<mods:number>1039</mods:number>
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<mods:start>1</mods:start>
<mods:end>108</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1039.64740</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1039.64740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1039-1</mods:identifier>
<mods:identifier type="ZooBank">3487C3570FAC4907A5C0C0A0C801C6E3</mods:identifier>
<mods:identifier type="Pensoft-UUID">FA2622A854585C59A7E8182A4E66C6DF</mods:identifier>
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<treatment ID-GBIF-Taxon="182248265" LSID="urn:lsid:plazi:treatment:F3A641DFB6F4528BBAB1E012F1B717C3" httpUri="http://treatment.plazi.org/id/F3A641DFB6F4528BBAB1E012F1B717C3" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="F3A641DF-B6F4-528B-BAB1-E012F1B717C3" authority="species group" authorityName="species group" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">Pyrrhalta semifulva species group</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="included species">
<paragraph pageId="0" pageNumber="1">Included species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gressitt &amp; Kimoto" authorityYear="1963" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta maculata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta maculata</emphasis>
</taxonomicName>
Gressitt &amp; Kimoto, 1963;
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
&amp; Lee, 2019;
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov.;
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. semifulva" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">P. semifulva</emphasis>
</taxonomicName>
(Jacoby, 1885);
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. discalis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="discalis">
<emphasis italics="true" pageId="0" pageNumber="1">P. discalis</emphasis>
</taxonomicName>
Gressitt &amp; Kimoto, 1963;
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
Kimoto, 1976; and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Small sized species (3.3-5.6 mm). Antenna stout, antennomeres VIII-X stout (1.4-2.0x longer than wide), III-VI slender. Body convex. Elytra relatively wider, 1.4-1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Aedeagus asymmetric; ostium covered by a membrane or lacking cover; endophallic sclerites composed of two slender sclerites, with several teeth on apex of primary sclerite and with one additional tooth near apex of secondary sclerite except
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. with only primary sclerite (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28D-F</figureCitation>
),
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
lacking teeth on sclerites (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24C, D</figureCitation>
),
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. semifulva" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">P. semifulva</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29C, D</figureCitation>
) and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. discalis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="discalis">
<emphasis italics="true" pageId="0" pageNumber="1">P. discalis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32C, D</figureCitation>
) lacking additional tooth on secondary sclerite. The ventrite VIII in females apically sclerotized, with dense short and long setae mixed along apical margin; spiculum long (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24F</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25F</figureCitation>
,
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28H</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29G</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32E</figureCitation>
,
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35F</figureCitation>
,
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36E</figureCitation>
). Gonocoxae apically sclerotized and with variable number of setae; both gonocoxae small and connected, with two long setae on each gonocoxa in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24H</figureCitation>
) and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25J</figureCitation>
), both gonocoxae longitudinally oriented and connected, with a number of setae near apices in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. discalis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="discalis">
<emphasis italics="true" pageId="0" pageNumber="1">P. discalis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32I</figureCitation>
), gonocoxae longitudinal and with dense, long setae in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35K</figureCitation>
), both gonocoxae separated, transverse, and disc with a number of scattered short setae in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36I</figureCitation>
). Both gonocoxae separated, small and with dense short setae in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28J</figureCitation>
) and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. semifulva" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">P. semifulva</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29K</figureCitation>
). Apical margin of abdominal ventrite V truncate or slightly concave, with deeply rounded depression at middle in males(Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24J</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25I</figureCitation>
,
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28K</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29J</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32H</figureCitation>
,
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35J</figureCitation>
,
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36H</figureCitation>
); slightly concave or widely rounded in females (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24I</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25H</figureCitation>
,
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28L</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29I</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32G</figureCitation>
,
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35I</figureCitation>
,
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36G</figureCitation>
). Mesotibia with apical spine in males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24E</figureCitation>
),
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25E</figureCitation>
),
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28G</figureCitation>
), and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35E</figureCitation>
); or lacking apical spine in those of the remaining species; mesotarsi with tarsomere I modified in males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24K</figureCitation>
),
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Habitus of Pyrrhalta jungchani sp. nov. and P. shirozui Kimoto A P. jungchani sp. nov., male, dorsal view B ditto, ventral view C ditto, lateral view D P. shirozui, female, dorsal view E ditto, ventral view F ditto, lateral view G Same species, color variation H Same species, color variation I Same species, color variation." figureDoi="10.3897/zookeys.1039.64740.figure38" httpUri="https://binary.pensoft.net/fig/545568" pageId="0" pageNumber="1">38M</figureCitation>
), and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. ishiharai" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">P. ishiharai</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Included species can be subdivided into species complexes based on similar color patterns. For example,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4664.4.4" author="Bezdek, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="518 - 534" refId="B7" refString="Bezdek, J, Lee, C-F, 2019. Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664: 518 - 534, DOI: https://doi.org/10.11646/zootaxa.4664.4.4" title="Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra." url="https://doi.org/10.11646/zootaxa.4664.4.4" volume="4664" year="2019">
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
and Lee (2019)
</bibRefCitation>
treated the
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
species complex, including
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
, and five more species. They are characterized by their maculate elytra (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Habitus of Pyrrhalta maculata Gressitt &amp; Kimoto and P. tsoui Bezdek &amp; Lee A P. maculata, female, dorsal view B ditto, ventral view C ditto, lateral view D P. tsoui female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure23" httpUri="https://binary.pensoft.net/fig/545553" pageId="0" pageNumber="1">23</figureCitation>
), strongly curved aedeagi (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24C, D</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25C, D</figureCitation>
), and only two setae on each gonocoxa (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24H</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25J</figureCitation>
).
<taxonomicName class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta semifulva</emphasis>
</taxonomicName>
and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. belong to another species complex characterized by their reddish brown elytra (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">27</figureCitation>
) and small gonoxae possessing dense setae (Figs
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28J</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29K</figureCitation>
).
<taxonomicName authorityName="Kimoto" authorityYear="1976" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta ishiharai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta ishiharai</emphasis>
</taxonomicName>
is grouped with
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. wulaiensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="wulaiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. wulaiensis</emphasis>
</taxonomicName>
sp. nov. based on the longitudinal ridges of the elytra (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Habitus of Pyrrhalta ishiharai Kimoto and P. wulaiensis sp. nov. A P. ishiharai, female, typical form, dorsal view B ditto, ventral view C ditto, lateral view D P. wulaiensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure34" httpUri="https://binary.pensoft.net/fig/545564" pageId="0" pageNumber="1">34A, D</figureCitation>
) and angular apices of aedeagi (Figs
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Diagnostic characters of Pyrrhalta ishiharai Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H tarsi of middle leg, male I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure35" httpUri="https://binary.pensoft.net/fig/545565" pageId="0" pageNumber="1">35C</figureCitation>
,
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Diagnostic characters of Pyrrhalta wulaiensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure36" httpUri="https://binary.pensoft.net/fig/545566" pageId="0" pageNumber="1">36C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Anthophagous species. Larvae and adults feed on flowers of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Meliosma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Meliosma rhoifolia" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="rhoifolia">
<emphasis italics="true" pageId="0" pageNumber="1">Meliosma rhoifolia</emphasis>
</taxonomicName>
(
<taxonomicName class="Dicotyledoneae" family="Sabiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Sabiaceae</taxonomicName>
) or species of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Rosaceae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>