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<document id="6BE965AE3F254A2F917C602A0DA73D40" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFF93733332FAFBE4E0D8FB1F" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium ignotum Wesener 2009, new species" docType="treatment" docVersion="4" lastPageNumber="100" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="97" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A3D85724123A29C75DEBD37BCABE5D62" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2514801F0E085182A08502CE4DEDFF8E">
<mods:title id="BE25A39FCBC9A31A5F71B0997A54DD28">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="05D5045E71E32FA15DEDB756D2600D76" type="personal">
<mods:role id="F2CD1036198492C4496FAD22114E7501">
<mods:roleTerm id="47E16891DBFCF5D1EEE21EA06CA5F819">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6BD86E2279DA6578C744E4101340773">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="4A185A7C5D561475B794275C24374A19">text</mods:typeOfResource>
<mods:relatedItem id="7F9FE32D014CBD1FDA17DEA9EEAF9C12" type="host">
<mods:titleInfo id="7935FA76FA1CFCF8FC9E8C105E1A557B">
<mods:title id="94443359738F088C6FB8E3E7D317D207">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0532F6FCFAEC4EFAE85C17FE8B977D0B">
<mods:date id="8C41CAAA54335C64CFB0D321F3AA3A97">2009</mods:date>
<mods:detail id="F860F9ED58E8E9BCD93B33F04D17E245" type="pubDate">
<mods:number id="53C2B5CCEA3A27B06DF9A62C3B41A4AF">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="85351866A1AC7ADA1DF67BE43609A0C5" type="volume">
<mods:number id="C892626B7492CF78154D3214C3ADA271">2097</mods:number>
</mods:detail>
<mods:detail id="9FC93646F006634284D15EA69EE20037" type="issue">
<mods:number id="C5528ADF0C1E643A09AF5FF99B98D1CC">1</mods:number>
</mods:detail>
<mods:extent id="380F8E54D6910B7062C0083F322E45CC" unit="page">
<mods:start id="AFEAB70C5E15EB2A2D74A9621606B10F">1</mods:start>
<mods:end id="A9B9D008591BFFECF781185421B70E36">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C02AE356FBB8B435ADB878E0305F0887">
<mods:url id="36D583FA51E26F1353B4E56630BC5922">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="73A11EC83C5375C7FB495026DC1A321F">journal article</mods:classification>
<mods:identifier id="A222A7F4DFA7E11F25D94DF40042AF3B" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="E3A73155EE9F27531505D47D38EFC803" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="3E0DB3A6B9904370135058804A740BB3" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="F29A9B3E566AD3A868B322990CA48345" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFF93733332FAFBE4E0D8FB1F" ID-DOI="http://doi.org/10.5281/zenodo.5325511" ID-Zenodo-Dep="5325511" LSID="urn:lsid:plazi:treatment:03FFBE1DFF93733332FAFBE4E0D8FB1F" httpUri="http://treatment.plazi.org/id/03FFBE1DFF93733332FAFBE4E0D8FB1F" lastPageId="99" lastPageNumber="100" pageId="96" pageNumber="97">
<subSubSection id="C34C5C80FF93733032FAFBE4E156FBA1" box="[151,593,1118,1144]" pageId="96" pageNumber="97" type="nomenclature">
<paragraph id="8BE90F0BFF93733032FAFBE4E156FBA1" blockId="96.[151,593,1118,1144]" box="[151,593,1118,1144]" pageId="96" pageNumber="97">
<heading id="D0A1B867FF93733032FAFBE4E156FBA1" bold="true" box="[151,593,1118,1144]" fontSize="11" level="1" pageId="96" pageNumber="97" reason="1">
<emphasis id="B922D319FF93733032FAFBE4E156FBA1" bold="true" box="[151,593,1118,1144]" pageId="96" pageNumber="97">
<taxonomicName id="4C567488FF93733032FAFBE4E2B4FBA1" ID-CoL="5D7H9" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,435,1118,1144]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">
<emphasis id="B922D319FF93733032FAFBE4E2B4FBA1" bold="true" box="[151,435,1118,1144]" italics="true" pageId="96" pageNumber="97">Zoosphaerium ignotum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2116E62FF93733033AFFBE4E156FBA1" box="[450,593,1118,1144]" pageId="96" pageNumber="97" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF93733032FAFB1DE10BFB31" pageId="96" pageNumber="97" type="etymology">
<paragraph id="8BE90F0BFF93733032FAFB1DE7BAFB1B" blockId="96.[151,1436,1191,2030]" box="[151,1213,1191,1218]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032FAFB1DE287FB18" bold="true" box="[151,384,1191,1217]" pageId="96" pageNumber="97">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FF93733033E5FB12E2EFFB18" authorityName="Wesener" authorityYear="2009" box="[392,488,1192,1217]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="ignotum">
<emphasis id="B922D319FF93733033E5FB12E2EFFB18" box="[392,488,1192,1217]" italics="true" pageId="96" pageNumber="97">ignotum</emphasis>
</taxonomicName>
, adjective, refers to the unknown
<typeStatus id="54EDB1A9FF937330311BFB12E0AFFB1B" box="[886,936,1192,1218]" pageId="96" pageNumber="97">type</typeStatus>
locality of this species.
</paragraph>
<paragraph id="8BE90F0BFF93733032ABFB74E10BFB31" blockId="96.[151,1436,1191,2030]" box="[198,524,1230,1256]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032ABFB74E2A7FB31" bold="true" box="[198,416,1230,1256]" pageId="96" pageNumber="97">Studied material:</emphasis>
1 M, 2 F
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF93733032AAFB4FE051FA84" pageId="96" pageNumber="97" type="materials_examined">
<paragraph id="8BE90F0BFF93733032AAFB4FE606FAEF" blockId="96.[151,1436,1191,2030]" pageId="96" pageNumber="97">
<materialsCitation id="3B3E0556FF93733032AAFB4FE606FAEF" collectionCode="SMF" collectorName="K. L. Koch" country="Madagascar" location="East Coast" pageId="96" pageNumber="97" specimenCount="1" typeStatus="holotype">
<emphasis id="B922D319FF93733032AAFB4FE245FAD6" bold="true" box="[199,322,1269,1295]" pageId="96" pageNumber="97">
<typeStatus id="54EDB1A9FF93733032AAFB4FE23AFAD6" box="[199,317,1269,1295]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 M (
<quantity id="4CAEA2EEFF93733033C5FB4FE101FAD6" box="[424,518,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="96" pageNumber="97" unit="mm" value="30.0">30 mm</quantity>
long),
<emphasis id="B922D319FF9373303003FB4FE1ADFAD6" bold="true" box="[622,682,1269,1295]" pageId="96" pageNumber="97">
<collectionCode id="ED4797CEFF9373303003FB4FE1ADFAD6" box="[622,682,1269,1295]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="96" pageNumber="97">SMF</collectionCode>
</emphasis>
,
<emphasis id="B922D319FF93733030AEFB4FE06AFAD7" box="[707,877,1269,1294]" italics="true" pageId="96" pageNumber="97">Locus typicus</emphasis>
:
<collectingCountry id="F3414F9BFF93733031EAFB4FE71EFAD6" box="[903,1049,1269,1295]" name="Madagascar" pageId="96" pageNumber="97">Madagascar</collectingCountry>
: details unclear. Label stating &quot;Wasserfälle des Uamotna&quot; (= waterfalls of the Uamotna),
<location id="8E8959D0FF9373303127FAA6E0CEFAEF" LSID="urn:lsid:plazi:treatment:03FFBE1DFF93733332FAFBE4E0D8FB1F:8E8959D0FF9373303127FAA6E0CEFAEF" box="[842,969,1308,1334]" country="Madagascar" name="East Coast" pageId="96" pageNumber="97">East Coast</location>
, leg.
<collectorName id="26A36ADDFF937330366AFAA6E789FAEF" box="[1031,1166,1308,1334]" pageId="96" pageNumber="97">K. L. Koch</collectorName>
, 02.1953.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFF93733032AAFAF8E051FA84" blockId="96.[151,1436,1191,2030]" box="[199,854,1346,1373]" pageId="96" pageNumber="97">
<materialsCitation id="3B3E0556FF93733032AAFAF8E055FA84" box="[199,850,1346,1373]" collectionCode="SMF" collectorName="K. L. Koch" country="Madagascar" location="East Coast" pageId="96" pageNumber="97" specimenCount="1" typeStatus="paratype">
<emphasis id="B922D319FF93733032AAFAF8E249FA85" bold="true" box="[199,334,1346,1372]" pageId="96" pageNumber="97">
<typeStatus id="54EDB1A9FF93733032AAFAF8E24EFA85" box="[199,329,1346,1372]" pageId="96" pageNumber="97" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2 F,
<emphasis id="B922D319FF93733033E5FAF8E2C3FA85" bold="true" box="[392,452,1346,1372]" pageId="96" pageNumber="97">
<collectionCode id="ED4797CEFF93733033E5FAF8E2C3FA85" box="[392,452,1346,1372]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="96" pageNumber="97">SMF</collectionCode>
</emphasis>
, same collection data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF93733032ABFAD2E7E6FA5A" box="[198,1249,1384,1411]" pageId="96" pageNumber="97" type="distribution">
<paragraph id="8BE90F0BFF93733032ABFAD2E7E6FA5A" blockId="96.[151,1436,1191,2030]" box="[198,1249,1384,1411]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032ABFAD2E2F4FA5B" bold="true" box="[198,499,1384,1410]" pageId="96" pageNumber="97">Distribution &amp; Ecology:</emphasis>
Only known from the
<typeStatus id="54EDB1A9FF9373303092FAD3E036FA5A" box="[767,817,1385,1411]" pageId="96" pageNumber="97">type</typeStatus>
locality, which could not be located.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF93733032ABFA35E004F8A3" pageId="96" pageNumber="97" type="diagnosis">
<paragraph id="8BE90F0BFF93733032ABFA35E7CEF906" blockId="96.[151,1436,1191,2030]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032ABFA35E24FFA70" bold="true" box="[198,328,1423,1449]" pageId="96" pageNumber="97">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFF93733033F2FA2AE2F4FA73" box="[415,499,1424,1450]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="96" pageNumber="97" unit="mm" value="37.0">37 mm</quantity>
long. Natural coloration unknown, legs and antenna green. Texture of tergites glabrous. Anal shield well-rounded, in male covered with dense field of minute setae. Disc of antenna with 25–32 apical cones (Fig. 67B). First sternite greatly enlarged, with well-rounded tip (Figs 66A, B). Movable finger of posterior telopods almost glabrous, broad, with well-rounded tip and four sclerotized spines (Fig. 66F); posterior side with
<emphasis id="B922D319FF93733033BEF991E109F99D" box="[467,526,1579,1604]" italics="true" pageId="96" pageNumber="97">circa</emphasis>
15 small sclerotized teeth (Fig. 66G). Fixed finger long, slender, curved, basally with single small spine, distally with sclerotized spots (Fig. 66F). Fixed finger shorter than movable finger. Single stridulation rib on each male harp (Fig. 66D) and two on each half of female washboard. Anal shield with two locking carinae, both small, of similar length. Operculum of vulva elongated into single process (Fig. 66C). Apical-most point of tip almost planar, very weakly notched (Fig. 66C).
</paragraph>
<paragraph id="8BE90F0BFF93733032ABF951E004F8A3" blockId="96.[151,1436,1191,2030]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032ABF951E28CF8DC" bold="true" box="[198,395,1771,1797]" pageId="96" pageNumber="97">Similar species:</emphasis>
<emphasis id="B922D319FF93733033FEF956E11DF8DC" box="[403,538,1772,1797]" italics="true" pageId="96" pageNumber="97">
<taxonomicName id="4C567488FF93733033FEF956E112F8DC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[403,533,1772,1797]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">Z. ignotum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FF937330304FF951E162F8DC" bold="true" box="[546,613,1771,1797]" pageId="96" pageNumber="97">
<taxonomicNameLabel id="A2116E62FF937330304FF951E162F8DC" box="[546,613,1771,1797]" pageId="96" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
features a unique combination of characters, like a single stridulation rib on the harp (Fig. 66D), more than 20 apical cones on the antennae (Fig. 67B) and strongly protruding lateral plateau on the third podomere of anterior telopods (Fig. 66D). The shape of the female operculum is unique among all known
<taxonomicName id="4C567488FF93733033ACF8DAE17DF8A3" box="[449,634,1888,1914]" class="Diplopoda" kingdom="Animalia" order="Sphaerotheriida" pageId="96" pageNumber="97" phylum="Arthropoda" rank="order">Sphaerotheriida</taxonomicName>
(Fig. 66C).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF93733332ABF83CE0D8FB1F" lastPageId="99" lastPageNumber="100" pageId="96" pageNumber="97" type="description">
<paragraph id="8BE90F0BFF93733032ABF83CE202F837" blockId="96.[151,1436,1191,2030]" pageId="96" pageNumber="97">
<emphasis id="B922D319FF93733032ABF83CE266F879" bold="true" box="[198,353,1926,1952]" pageId="96" pageNumber="97">Description:</emphasis>
<emphasis id="B922D319FF9373303306F83DE2FCF879" box="[363,507,1927,1952]" italics="true" pageId="96" pageNumber="97">Body length</emphasis>
:
<typeStatus id="54EDB1A9FF9373303060F83CE17DF879" box="[525,634,1926,1952]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
male: length
<quantity id="4CAEA2EEFF937330314EF83CE07CF879" box="[803,891,1926,1952]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="96" pageNumber="97" unit="mm" value="30.0">30 mm</quantity>
, width of thoracic shield
<quantity id="4CAEA2EEFF93733036DEF83CE626F879" box="[1203,1313,1926,1952]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="96" pageNumber="97" unit="mm" value="16.7">16.7 mm</quantity>
, height of thoracic shield
<quantity id="4CAEA2EEFF9373303320F817E2ABF81E" box="[333,428,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="96" pageNumber="97" unit="mm" value="8.6">8.6 mm</quantity>
. Female
<typeStatus id="54EDB1A9FF9373303074F817E18EF81E" box="[537,649,1965,1991]" pageId="96" pageNumber="97" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D50C482FF93733030F6F817E034F81E" box="[667,819,1965,1991]" pageId="96" pageNumber="97" type="generic">2 specimens</specimenCount>
): length
<quantity id="4CAEA2EEFF93733031F6F817E0F5F81E" box="[923,1010,1965,1991]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="96" pageNumber="97" unit="mm" value="37.0">37 mm</quantity>
, width up to
<quantity id="4CAEA2EEFF93733036FFF817E7F8F81E" box="[1170,1279,1965,1991]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="96" pageNumber="97" unit="mm" value="22.4">22.4 mm</quantity>
, height up to
<quantity id="4CAEA2EEFF93733032FAF86EE206F837" box="[151,257,2004,2030]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="96" pageNumber="97" unit="mm" value="11.4">11.4 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF92733132FAFAC7E004F99B" blockId="97.[151,1436,1405,1603]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132FAFAC7E228FA4C" bold="true" box="[151,303,1405,1429]" pageId="97" pageNumber="98">FIGURE 066</emphasis>
.
<taxonomicName id="4C567488FF9273313353FAC4E13FFA4C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[318,568,1406,1429]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">
<emphasis id="B922D319FF9273313353FAC4E13FFA4C" box="[318,568,1406,1429]" italics="true" pageId="97" pageNumber="98">Zoosphaerium ignotum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF927331302CFAC7E178FA4C" bold="true" box="[577,639,1405,1429]" pageId="97" pageNumber="98">
<taxonomicNameLabel id="A2116E62FF927331302CFAC7E178FA4C" box="[577,639,1405,1429]" pageId="97" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, B, D–G
<typeStatus id="54EDB1A9FF9273313162FAC4E068FA4C" box="[783,879,1406,1429]" pageId="97" pageNumber="98" type="holotype">holotype</typeStatus>
, C female
<typeStatus id="54EDB1A9FF9273313185FAC4E743FA4C" box="[1000,1092,1406,1429]" pageId="97" pageNumber="98" type="paratype">paratype</typeStatus>
<emphasis id="B922D319FF9273313621FAC7E758FA4C" bold="true" box="[1100,1119,1405,1429]" pageId="97" pageNumber="98">A</emphasis>
: first left coxa and prefemur with first stigmatic plate, anterior view,
<emphasis id="B922D319FF927331302EFA1AE152FA61" bold="true" box="[579,597,1440,1464]" pageId="97" pageNumber="98">B</emphasis>
: first stigmatic plate, posterior view;
<emphasis id="B922D319FF927331318AFA1AE0FDFA61" bold="true" box="[999,1018,1440,1464]" pageId="97" pageNumber="98">C</emphasis>
: second left coxa with vulva, posterior view;
<emphasis id="B922D319FF92733132B5FA79E3ECFA02" bold="true" box="[216,235,1475,1499]" pageId="97" pageNumber="98">D</emphasis>
: left half of anterior telopods, anterior view;
<emphasis id="B922D319FF92733130A1FA79E1D9FA02" bold="true" box="[716,734,1475,1499]" pageId="97" pageNumber="98">E</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FF9273313772FA79E628FA02" bold="true" box="[1311,1327,1475,1499]" pageId="97" pageNumber="98">F</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FF92733133E2FA5FE2A4FA24" bold="true" box="[399,419,1509,1533]" pageId="97" pageNumber="98">G</emphasis>
: posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FF92733131D3FA5FE0E3FA24" bold="true" box="[958,996,1509,1533]" pageId="97" pageNumber="98">cav</emphasis>
= cavity;
<emphasis id="B922D319FF927331363DFA5FE777FA24" bold="true" box="[1104,1136,1509,1533]" pageId="97" pageNumber="98">Cx</emphasis>
= coxa;
<emphasis id="B922D319FF92733136A0FA5FE7E8FA24" bold="true" box="[1229,1263,1509,1533]" pageId="97" pageNumber="98">EP</emphasis>
= external plate of vulva;
<emphasis id="B922D319FF9273313290F9B2E210F9F9" bold="true" box="[253,279,1544,1568]" pageId="97" pageNumber="98">IP</emphasis>
= mesal plate of vulva;
<emphasis id="B922D319FF9273313074F9B2E129F9F9" bold="true" box="[537,558,1544,1568]" pageId="97" pageNumber="98">O</emphasis>
= operculum;
<emphasis id="B922D319FF92733130A6F9B2E1F4F9F9" bold="true" box="[715,755,1544,1568]" pageId="97" pageNumber="98">Pre</emphasis>
= prefemur;
<emphasis id="B922D319FF9273313112F9B2E0A6F9F9" bold="true" box="[895,929,1544,1568]" pageId="97" pageNumber="98">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FF927331361FF9B2E7A6F9F9" bold="true" box="[1138,1185,1544,1568]" pageId="97" pageNumber="98">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FF9273313717F9B2E69BF9F9" bold="true" box="[1402,1436,1544,1568]" pageId="97" pageNumber="98">SR</emphasis>
= stridulation rib;
<emphasis id="B922D319FF9273313335F991E269F99A" bold="true" box="[344,366,1579,1603]" pageId="97" pageNumber="98">St</emphasis>
= stigmatic plate. Scale bars =
<quantity id="4CAEA2EEFF92733130D3F991E1F8F99B" box="[702,767,1579,1602]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="97" pageNumber="98" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF92733132AAF9CBE1FCF952" blockId="97.[151,1436,1649,2022]" box="[199,763,1649,1675]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132AAF9CBE229F953" box="[199,302,1649,1674]" italics="true" pageId="97" pageNumber="98">Habitus:</emphasis>
Medium sized, glabrous pill millipede.
</paragraph>
<paragraph id="8BE90F0BFF92733132AAF922E2B8F927" blockId="97.[151,1436,1649,2022]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132AAF922E256F968" box="[199,337,1688,1713]" italics="true" pageId="97" pageNumber="98">Coloration:</emphasis>
Strongly faded after more than 50 years in alcohol. Head, collum, thoracic shield, tergites and anal shield faded light-green to light-brown colorized, posterior margin of all tergites thin, dark-brownish. Antennae and legs green.
</paragraph>
<paragraph id="8BE90F0BFF92733132AAF8B1E116F8FC" blockId="97.[151,1436,1649,2022]" box="[199,529,1803,1829]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132AAF8B1E217F8FD" box="[199,272,1803,1828]" italics="true" pageId="97" pageNumber="98">Head:</emphasis>
Eyes with&gt;75 ocelli.
</paragraph>
<paragraph id="8BE90F0BFF92733132AAF888E60EF840" blockId="97.[151,1436,1649,2022]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132AAF888E238F892" box="[199,319,1842,1867]" italics="true" pageId="97" pageNumber="98">Antennae:</emphasis>
Length of antennomeres: 1=2&gt;3&gt;4=5&lt;&lt;6, sixth antennomere longest, disc bearing 25–32 sensory cones (Fig. 67B). First antennomere remarkably broader than others, short, with groove and 1 or 2 rows of
<emphasis id="B922D319FF927331329AF83AE2D0F840" box="[247,471,1920,1945]" italics="true" pageId="97" pageNumber="98">sensilla basiconica</emphasis>
(Fig. 67A). Sclerotized teeth at base of antennomeres 1–5 (Fig. 67B).
</paragraph>
<paragraph id="8BE90F0BFF92733132AAF81CE0CEF83F" blockId="97.[151,1436,1649,2022]" pageId="97" pageNumber="98">
<emphasis id="B922D319FF92733132AAF81CE247F866" box="[199,320,1958,1983]" italics="true" pageId="97" pageNumber="98">Mandible:</emphasis>
Six rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 67C). Condylus with strongly developed step near apical margin (Fig. 67C).
</paragraph>
<paragraph id="8BE90F0BFF91733232AAFF22E123FF01" blockId="98.[151,1436,152,332]" pageId="98" pageNumber="99">
<emphasis id="B922D319FF91733232AAFF22E296FF68" box="[199,401,152,177]" italics="true" pageId="98" pageNumber="99">Gnathochilarium</emphasis>
: Laterally of palpi 3 or 4 sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
</paragraph>
<paragraph id="8BE90F0BFF91733232AAFF5FE1AEFF26" blockId="98.[151,1436,152,332]" box="[199,681,229,255]" pageId="98" pageNumber="99">
<emphasis id="B922D319FF91733232AAFF5FE220FF27" box="[199,295,229,254]" italics="true" pageId="98" pageNumber="99">Collum:</emphasis>
Median part of collum glabrous.
</paragraph>
<paragraph id="8BE90F0BFF91733232AAFEB6E030FEFF" blockId="98.[151,1436,152,332]" box="[199,823,268,294]" pageId="98" pageNumber="99">
<emphasis id="B922D319FF91733232AAFEB6E279FEFC" box="[199,382,268,293]" italics="true" pageId="98" pageNumber="99">Thoracic shield</emphasis>
: Thoracic shield surface like tergites.
</paragraph>
<paragraph id="8BE90F0BFF91733232AAFE88E021FE95" blockId="98.[151,1436,152,332]" box="[199,806,306,332]" pageId="98" pageNumber="99">
<emphasis id="B922D319FF91733232AAFE88E224FE92" box="[199,291,306,331]" italics="true" pageId="98" pageNumber="99">Tergites</emphasis>
: Paratergites slightly projecting posteriorly.
</paragraph>
<paragraph id="8BE90F0BFF91733232FAF83BE719F807" blockId="98.[151,1436,1921,2015]" pageId="98" pageNumber="99">
<emphasis id="B922D319FF91733232FAF83BE237F840" bold="true" box="[151,304,1921,1945]" pageId="98" pageNumber="99">FIGURE 067</emphasis>
.
<taxonomicName id="4C567488FF917332332DF838E13CF840" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[320,571,1922,1945]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">
<emphasis id="B922D319FF917332332DF838E13CF840" box="[320,571,1922,1945]" italics="true" pageId="98" pageNumber="99">Zoosphaerium ignotum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9173323029F83BE184F840" bold="true" box="[580,643,1921,1945]" pageId="98" pageNumber="99">
<taxonomicNameLabel id="A2116E62FF9173323029F83BE184F840" box="[580,643,1921,1945]" pageId="98" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FF91733230FEF838E1F5F840" box="[659,754,1922,1945]" pageId="98" pageNumber="99" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FF917332312BF83BE05EF840" bold="true" box="[838,857,1921,1945]" pageId="98" pageNumber="99">A</emphasis>
: antenna, first antennomere with
<emphasis id="B922D319FF91733236A0F838E69CF840" box="[1229,1435,1922,1945]" italics="true" pageId="98" pageNumber="99">sensilla basiconica</emphasis>
(arrow);
<emphasis id="B922D319FF9173323298F81EE200F865" bold="true" box="[245,263,1956,1980]" pageId="98" pageNumber="99">B</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FF917332306FF81EE112F865" bold="true" box="[514,533,1956,1980]" pageId="98" pageNumber="99">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FF9173323194F81EE727F865" bold="true" box="[1017,1056,1956,1980]" pageId="98" pageNumber="99">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FF91733236BFF81EE7F5F865" bold="true" box="[1234,1266,1956,1980]" pageId="98" pageNumber="99">Co</emphasis>
= condylus;
<emphasis id="B922D319FF9173323713F81EE69BF865" bold="true" box="[1406,1436,1956,1980]" pageId="98" pageNumber="99">eT</emphasis>
= external tooth;
<emphasis id="B922D319FF9173323323F87DE274F806" bold="true" box="[334,371,1991,2015]" pageId="98" pageNumber="99">mp</emphasis>
= molar plate;
<emphasis id="B922D319FF917332307BF87DE131F806" bold="true" box="[534,566,1991,2015]" pageId="98" pageNumber="99">pL</emphasis>
= pectinate lamella;
<emphasis id="B922D319FF917332317BF87DE028F806" bold="true" box="[790,815,1991,2015]" pageId="98" pageNumber="99">sb</emphasis>
=
<emphasis id="B922D319FF9173323121F87DE710F807" box="[844,1047,1991,2014]" italics="true" pageId="98" pageNumber="99">sensilla basiconica</emphasis>
.
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFF22E161FF26" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFF22E267FF68" box="[199,352,152,177]" italics="true" pageId="99" pageNumber="100">Endotergum:</emphasis>
Inner area with broad conical spines and long hair. Single row of small, circular cuticular impression, regularly distributed (Fig. 74B). Two or three rows of short marginal bristles, reaching up towards 1/3 of way to tergite margin (Fig. 74B).
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFEB6E0D3FE95" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFEB6E25CFEFC" box="[199,347,268,293]" italics="true" pageId="99" pageNumber="100">Anal shield:</emphasis>
Well-rounded, neither bell-shaped nor tapered. Completely covered with dense field of minute setae. Underside carries two same-sized, black locking carinae.
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFEE3E264FE3E" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFEE3E20FFEAB" box="[199,264,345,370]" italics="true" pageId="99" pageNumber="100">Legs:</emphasis>
Tarsi of first pair of legs with two or three, second with three or four and third pair with six to eight ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 12 or 13 ventral spines and one apical spine. In 9th leg femur 2.1 times, tarsi 5.4 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFE4EE772FDD7" blockId="99.[151,1437,152,1222]" box="[199,1141,500,526]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFE4EE28BFDD4" box="[199,396,500,525]" italics="true" pageId="99" pageNumber="100">Stigmatic plates:</emphasis>
First lobe enlarged, with well-rounded, blunt tip (Figs 66A, B).
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFDA0E2EAFD16" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFDA0E2FFFDEA" box="[199,504,538,563]" italics="true" pageId="99" pageNumber="100">Female sexual characters</emphasis>
: Subanal plate with washboard, consisting of well-developed stridulation ribs, two ribs on each half. Stridulation ribs symmetrical and strong. Vulva large, covering more than 2/3 of coxa surface (Fig. 66C). Operculum large, protruding to apical edge of coxa, with one medial process. Apical margin of operculum on tip only shallowly notched. Mesal plate long and broad, reaching 1/4 to half of operculum height (Fig. 66C).
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFD66E0F8FD2F" blockId="99.[151,1437,152,1222]" box="[199,1023,732,758]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFD66E2DDFD2C" box="[199,474,732,757]" italics="true" pageId="99" pageNumber="100">Male sexual characters</emphasis>
: Anal shield covered with dense field of setae.
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFCB8E7A3FC6E" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFCB8E29CFCC2" box="[199,411,770,795]" italics="true" pageId="99" pageNumber="100">Anterior telopods</emphasis>
(Figs 66D, E): Syncoxite on both sides with only very few, isolated hairs. First podomere broad, width greater than height, harp with one stridulation rib, rib straight, very long and welldeveloped. Second podomere process lobe-like and little curved with rounded edges, reaching up to half of third podomeres height, apically with sclerotized spots. Third podomere long, cavity mesally with numerous sclerotized spots and three sclerotized spines. One sclerotized spot located closely to tip.
</paragraph>
<paragraph id="8BE90F0BFF90733332AAFC7EE0D8FB1F" blockId="99.[151,1437,152,1222]" pageId="99" pageNumber="100">
<emphasis id="B922D319FF90733332AAFC7EE2A3FC04" box="[199,420,964,989]" italics="true" pageId="99" pageNumber="100">Posterior telopods</emphasis>
(Figs 66E, F): Movable finger of chela thick, 2.1 times longer than wide, weakly curved, apically tapering. Hollowed-out inner margin with four sclerotized spines, posterior aspect with
<emphasis id="B922D319FF907333370CFC50E69BFBDA" box="[1377,1436,1002,1027]" italics="true" pageId="99" pageNumber="100">circa</emphasis>
15 small, sclerotized teeth. Fixed finger shorter and more slender than movable finger. Fixed finger tip weakly curved, anterior side basally with single sclerotized spine and apically with numerous sclerotized spots juxtaposed. Fixed finger on both sides covered with few long hairs. Movable finger on posterior side medially with few hairs, remaining parts of posterior telopods glabrous. Inner horns of syncoxite apically curved posteriorly. Tips sharp, whole horns covered with numerous short hairs.
</paragraph>
</subSubSection>
</treatment>
</document>
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