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<document id="A229B8FFE3E802A670AB8FC9D322AF1E" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFA0F76F01F310DE4C00C9A0" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada camdenensis Köhler 2010, n.sp." docType="treatment" docVersion="3" lastPageNumber="268" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="266" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="49FC0AE05A0D07EE1D9CDB368EB8363B">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:name id="84DFABEA158AF25E648262069BBA0F9E" type="personal">
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<mods:namePart id="ACD984C61688EA755D4AE34C56C9A9E9">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="420AB79ADF1BB53945AA39A8AAC572D8">text</mods:typeOfResource>
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<mods:title id="AD7051BAE59D367184CC8010F0088494">Records of the Australian Museum</mods:title>
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<mods:part id="F509EF5F8D00BF4543CC70277700F1DB">
<mods:date id="5FA5F43903B26E1BDAC266C2B456B8F6">2010</mods:date>
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<mods:number id="252F028A3F6CE082B647A0D9AF621195">2010-11-24</mods:number>
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<mods:number id="9CF75ABC2DF377CFC1CFBBE5A005F579">62</mods:number>
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<mods:number id="411981A535B6886AD74E2A7BF4D1BA69">2</mods:number>
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<mods:url id="F3CF7F369BB5F36C0A59A4B7305C9BDE">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<mods:classification id="686838E0E368A06489A5E7A37103240C">journal article</mods:classification>
<mods:identifier id="2479D95CCBE4272A46B4E99FA9554E17" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="319D9441157A30D76D7D035FED75BF6A" type="ISSN">2201-4349</mods:identifier>
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<treatment id="03F987C5FFA0F76F01F310DE4C00C9A0" ID-DOI="http://doi.org/10.5281/zenodo.5239051" ID-Zenodo-Dep="5239051" LSID="urn:lsid:plazi:treatment:03F987C5FFA0F76F01F310DE4C00C9A0" httpUri="http://treatment.plazi.org/id/03F987C5FFA0F76F01F310DE4C00C9A0" lastPageId="51" lastPageNumber="268" pageId="49" pageNumber="266">
<subSubSection id="C34A6558FFA0F76D01F310DE4834CEC0" box="[902,1313,154,181]" pageId="49" pageNumber="266" type="nomenclature">
<paragraph id="8BEF36D3FFA0F76D01F310DE4834CEC0" blockId="49.[902,1313,154,181]" box="[902,1313,154,181]" pageId="49" pageNumber="266">
<heading id="D0A781BFFFA0F76D01F310DE4834CEC0" box="[902,1313,154,181]" centered="true" fontSize="11" level="2" pageId="49" pageNumber="266" reason="2">
<emphasis id="B924EAC1FFA0F76D01F310DE4834CEC0" bold="true" box="[902,1313,154,181]" pageId="49" pageNumber="266">
<taxonomicName id="4C504D50FFA0F76D01F310DE49C8CEC1" ID-CoL="D7QC" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[902,1245,155,181]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="266" phylum="Mollusca" rank="species" species="camdenensis" status="sp. nov.">
<emphasis id="B924EAC1FFA0F76D01F310DE49C8CEC1" bold="true" box="[902,1245,155,181]" italics="true" pageId="49" pageNumber="266">Amplirhagada camdenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA0F76D069010DF4834CEC0" box="[1253,1313,154,180]" pageId="49" pageNumber="266" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFA0F76D016F10924998CC5C" pageId="49" pageNumber="266" type="materials_examined">
<paragraph id="8BEF36D3FFA0F76D016F109249CCCF17" blockId="49.[794,1421,215,355]" pageId="49" pageNumber="266">
<materialsCitation id="3B383C8EFFA0F76D016F109249C0CF17" collectingDate="2008-05-25" collectorName="Vine &amp; M. Shea" county="Kimberley" latitude="-15.337777" location="Camden Sound" longLatPrecision="21" longitude="124.52112" municipality="Bonaparte Archipelago" pageId="49" pageNumber="266" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFA0F76D016F10924EADCE9B" bold="true" box="[794,952,215,239]" pageId="49" pageNumber="266">
<typeStatus id="54EB8871FFA0F76D016F10924E43CE9B" box="[794,854,215,239]" pageId="49" pageNumber="266">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFA0F76D01B810924901CE9A" box="[973,1044,215,238]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="49" pageNumber="266">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFA0F76D065A10924814CE9A" box="[1071,1281,215,238]" country="Australia" name="Western Australia" pageId="49" pageNumber="266">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFA0F76D07651092489FCE9A" box="[1296,1418,215,238]" pageId="49" pageNumber="266">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFA0F76D016F10B14903CF7F" box="[794,1046,244,267]" pageId="49" pageNumber="266">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FFA0F76D065610B149D9CF7F" LSID="urn:lsid:plazi:treatment:03F987C5FFA0F76F01F310DE4C00C9A0:8E8F6008FFA0F76D065610B149D9CF7F" box="[1059,1228,244,267]" county="Kimberley" latitude="-15.337777" longLatPrecision="21" longitude="124.52112" municipality="Bonaparte Archipelago" name="Camden Sound" pageId="49" pageNumber="266" stateProvince="Western Australia">Camden Sound</location>
,
<location id="8E8F6008FFA0F76D06A210B1489CCF7F" LSID="urn:lsid:plazi:treatment:03F987C5FFA0F76F01F310DE4C00C9A0:8E8F6008FFA0F76D06A210B1489CCF7F" box="[1239,1417,244,267]" county="Kimberley" latitude="-15.337777" longLatPrecision="21" longitude="124.52112" municipality="Bonaparte Archipelago" name="Augustus Island" pageId="49" pageNumber="266" stateProvince="Western Australia">Augustus Island</location>
, northwestern section,
<geoCoordinate id="EE645014FFA0F76D067D11574997CF5D" box="[1032,1154,274,297]" degrees="15" direction="south" minutes="20" orientation="latitude" pageId="49" pageNumber="266" precision="15" seconds="16" value="-15.337777">15°20'16&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFA0F76D06FF11574800CF5D" box="[1162,1301,274,297]" degrees="124" direction="east" minutes="31" orientation="longitude" pageId="49" pageNumber="266" precision="15" seconds="16" value="124.52112">124°31'16&quot;E</geoCoordinate>
; KIS 1-49.
<collectorName id="26A55305FFA0F76D016F116A4E59CF32" box="[794,844,303,326]" pageId="49" pageNumber="266">Vine</collectorName>
thicket amongst boulders at base of sandstone cliffs, under rocks (leg.
<collectorName id="26A55305FFA0F76D01A11109493ACF17" box="[980,1071,332,355]" pageId="49" pageNumber="266">M. Shea</collectorName>
,
<date id="FFEE1013FFA0F76D064F110949D8CF17" box="[1082,1229,332,355]" pageId="49" pageNumber="266" value="2008-05-25">
<collectingDate id="EFAAE9FBFFA0F76D064F110949D8CF17" box="[1082,1229,332,355]" pageId="49" pageNumber="266" value="2008-05-25">25 May 2008</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFA0F76D016F11C74E82CFAA" blockId="49.[794,1422,386,478]" pageId="49" pageNumber="266">
<emphasis id="B924EAC1FFA0F76D016F11C74E8ACFE1" bold="true" box="[794,927,386,405]" pageId="49" pageNumber="266">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFA0F76D01D911C64E82CFAA" accessionNumber="S34624, S37006, S37005" collectionCode="WAM" latitude="-15.349722" longLatPrecision="21" longitude="124.525" pageId="49" pageNumber="266" specimenCount="3" typeStatus="holotype">
<typeStatus id="54EB8871FFA0F76D01D911C64EE8CFE2" box="[940,1021,387,406]" pageId="49" pageNumber="266" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFA0F76D067011C6492FCFE2" box="[1029,1082,387,406]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D063411C64991CFE2" box="[1089,1156,387,406]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34624" pageId="49" pageNumber="266" type="EnaNcbi">S34624</accessionNumber>
(Pl. 2.12). AMS C463722 (3 preserved specimens, as
<typeStatus id="54EB8871FFA0F76D019C11DE4922CFDA" box="[1001,1079,411,430]" pageId="49" pageNumber="266" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FFA0F76D063111DE496DCFDA" box="[1092,1144,411,430]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D060911DE49ABCFDA" box="[1148,1214,411,430]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37006" pageId="49" pageNumber="266" type="EnaNcbi">S37006</accessionNumber>
(5 preserved specimens, as
<typeStatus id="54EB8871FFA0F76D014311F64E98CFB2" box="[822,909,435,454]" pageId="49" pageNumber="266" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FFA0F76D01D511F64EC2CFB2" box="[928,983,435,454]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D019511F64932CFB2" box="[992,1063,435,454]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37005" pageId="49" pageNumber="266" type="EnaNcbi">S37005</accessionNumber>
(3 preserved specimens,
<geoCoordinate id="EE645014FFA0F76D075611F64898CFB2" box="[1315,1421,435,454]" degrees="15" direction="south" minutes="20" orientation="latitude" pageId="49" pageNumber="266" precision="15" seconds="59" value="-15.349722">15°20'59&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFA0F76D016F118E4E9ECFAA" box="[794,907,459,478]" degrees="124" direction="east" minutes="31" orientation="longitude" pageId="49" pageNumber="266" precision="15" seconds="30" value="124.525">124°31'30&quot;E</geoCoordinate>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEF36D3FFA0F76D016F11B84998CC5C" blockId="49.[794,1421,509,552]" pageId="49" pageNumber="266">
<emphasis id="B924EAC1FFA0F76D016F11B8490ACC64" bold="true" box="[794,1055,509,528]" pageId="49" pageNumber="266">Additional, non-type material</emphasis>
.
<collectionCode id="ED41AE16FFA0F76D065211B8494FCC64" box="[1063,1114,509,528]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D062B11B849B4CC64" box="[1118,1185,509,528]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37007" pageId="49" pageNumber="266" type="EnaNcbi">S37007</accessionNumber>
8,
<collectionCode id="ED41AE16FFA0F76D06CE11B849FBCC64" box="[1211,1262,509,528]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D068711B84827CC64" box="[1266,1330,509,528]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37445" pageId="49" pageNumber="266" type="EnaNcbi">S37445</accessionNumber>
50,
<collectionCode id="ED41AE16FFA0F76D072F11B84898CC64" box="[1370,1421,509,528]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="266" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA0F76D016F12504E48CC5C" box="[794,861,533,552]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41481" pageId="49" pageNumber="266" type="EnaNcbi">S41481</accessionNumber>
,
<collectionCode id="ED41AE16FFA0F76D011312504EB1CC5C" box="[870,932,533,552]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="49" pageNumber="266" type="Museum">FMNH</collectionCode>
219063 (Augustus Island).
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFA0F76F016F120C4C00C9A0" lastPageId="51" lastPageNumber="268" pageId="49" pageNumber="266" type="etymology">
<paragraph id="8BEF36D3FFA0F76D016F120C4955CC0A" blockId="49.[794,1421,585,638]" pageId="49" pageNumber="266">
<emphasis id="B924EAC1FFA0F76D016F120C4E8ECC15" bold="true" box="[794,923,585,609]" pageId="49" pageNumber="266">Etymology</emphasis>
. In reference to Camden Sound, in which Augustus Island is situated.
</paragraph>
<paragraph id="8BEF36D3FFA0F76D016F12DA495FCCC2" blockId="49.[794,1098,671,694]" box="[794,1098,671,694]" pageId="49" pageNumber="266">Sealing strategy. Free sealer.</paragraph>
<paragraph id="8BEF36D3FFA0F76E016F12924C7ACBD9" blockId="49.[794,1422,727,985]" lastBlockId="50.[166,794,1195,1453]" lastPageId="50" lastPageNumber="267" pageId="49" pageNumber="266">
<emphasis id="B924EAC1FFA0F76D016F12924E47CC9B" bold="true" box="[794,850,727,751]" pageId="49" pageNumber="266">Shell</emphasis>
(
<figureCitation id="136B2A56FFA0F76D012A12924ECACC9A" box="[863,991,727,750]" captionStart="Figure 77" captionStartId="50.[166,229,1086,1107]" captionTargetBox="[166,1466,151,1073]" captionTargetId="figure-74@50.[166,1466,151,1073]" captionTargetPageId="50" captionText="Figure 77. SEM photographs of shell and radula of Amplirhagada camdenensis n.sp., paratype AMS C463722: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first three whorls viewed from above (scale 1 mm); (C) close-up showing central and inner lateral teeth (scale 10 µm); (D) close-up showing inner and middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239255" httpUri="https://zenodo.org/record/5239255/files/figure.png" pageId="49" pageNumber="266">Fig. 77AB</figureCitation>
, Pl. 2.12). Globose, with medium to high spire; solid. Periphery evenly rounded to slightly angulate; upper sector of whorl flattened to slightly shouldered, basal sector rounded. Umbilicus forming a chink to narrowly winding opening, 40100 percent concealed by columellar reflection. Background colour yellowish brown to horn; sub-sutural band diffuse, dark to chestnut brown, becomes more indistinct or blurs towards tip of whorl, may fuse with mid-whorl band to cover shell in light brown colour; mid-whorl band diffuse, dark to chestnut brown, thin to moderately broad, visible on last whorl(s) only; ventral colour whitish cream to yellowish brown; outer and inner lip colour differs from shell, pinkish brown. Protoconch c.
<quantity id="4CA89B36FFA3F76E008D15464DDACB4C" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="50" pageNumber="267" unit="mm" value="3.5">3.5 mm</quantity>
in diameter, comprising 2 whorls, smooth. Teleoconch smooth except of faint axial growth lines. Angle of aperture 30°; outer lip rounded, moderately thick, expanded, not or slightly reflected, basal and palatal node absent. Parietal wall of inner lip absent.
</paragraph>
<caption id="DF2F665BFFA3F76E02D3147B4956CAFC" ID-DOI="http://doi.org/10.5281/zenodo.5239255" ID-Zenodo-Dep="5239255" httpUri="https://zenodo.org/record/5239255/files/figure.png" pageId="50" pageNumber="267" startId="50.[166,229,1086,1107]" targetBox="[166,1466,151,1073]" targetPageId="50">
<paragraph id="8BEF36D3FFA3F76E02D3147B4956CAFC" blockId="50.[166,1467,1086,1160]" pageId="50" pageNumber="267">
Figure 77. SEM photographs of shell and radula of
<taxonomicName id="4C504D50FFA3F76E00E1147B4EB5CA27" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[660,928,1086,1107]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="50" pageNumber="267" phylum="Mollusca" rank="species" species="camdenensis" status="sp. nov.">
<emphasis id="B924EAC1FFA3F76E00E1147B4EB5CA27" box="[660,928,1086,1107]" italics="true" pageId="50" pageNumber="267">Amplirhagada camdenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA3F76E01D0147B4EC0CA27" box="[933,981,1086,1107]" pageId="50" pageNumber="267" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463722: (
<emphasis id="B924EAC1FFA3F76E06A9147B49FECA27" box="[1244,1259,1086,1107]" italics="true" pageId="50" pageNumber="267">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFA3F76E03C8141D4CD9CA19" box="[445,460,1112,1133]" italics="true" pageId="50" pageNumber="267">B</emphasis>
) sculpture on first three whorls viewed from above (scale 1 mm); (
<emphasis id="B924EAC1FFA3F76E0638141D4948CA19" box="[1101,1117,1112,1133]" italics="true" pageId="50" pageNumber="267">C</emphasis>
) close-up showing central and inner lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FFA3F76E03C114364CD0CAFC" box="[436,453,1139,1160]" italics="true" pageId="50" pageNumber="267">D</emphasis>
) close-up showing inner and middle marginal teeth (scale 10 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFA3F76E013214EE4870CA94" blockId="50.[839,1466,1195,1248]" pageId="50" pageNumber="267">
<emphasis id="B924EAC1FFA3F76E013214EE490ACAB7" bold="true" box="[839,1055,1195,1219]" pageId="50" pageNumber="267">Pallial morphology</emphasis>
. Pigmentation on mantle mottled, dark grey. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFA3F76E0132154649DAC95E" blockId="50.[839,1468,1283,1834]" pageId="50" pageNumber="267">
<emphasis id="B924EAC1FFA3F76E013215464938CB6F" bold="true" box="[839,1069,1283,1307]" pageId="50" pageNumber="267">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFA3F76E064E15464980CB6E" box="[1083,1173,1283,1306]" captionStart="Figure 76" captionStartId="49.[794,857,1117,1138]" captionTargetBox="[794,1417,1018,2010]" captionTargetId="figure-369@49.[794,1421,1014,2014]" captionTargetPageId="49" captionText="Figure 76. Interior of penial chamber" figureDoi="http://doi.org/10.5281/zenodo.5239251" httpUri="https://zenodo.org/record/5239251/files/figure.png" pageId="50" pageNumber="267">Figs. 76</figureCitation>
,
<figureCitation id="136B2A56FFA3F76E06D5154649ABCB6E" box="[1184,1214,1283,1306]" captionStart="Figure 78" captionStartId="50.[166,229,1932,1953]" captionTargetBox="[175,781,1506,1912]" captionTargetId="figure-75@50.[166,793,1497,1925]" captionTargetPageId="50" captionText="Figure 78. Genitalia of Amplirhagada camdenensis n.sp., paratype AMS C463722 (25 May, scale 10 mm). Compare with Fig. 3 for labelling of structures." figureDoi="http://doi.org/10.5281/zenodo.5239257" httpUri="https://zenodo.org/record/5239257/files/figure.png" pageId="50" pageNumber="267">78</figureCitation>
). Penis straight, shorter than anterior part of oviduct, very thin. Vas deferens forms simple loop before entering penis. Penial retractor muscle as long as penis. Penial verge rather short (
<emphasis id="B924EAC1FFA3F76E076F151E4837CB1D" bold="true" box="[1306,1314,1371,1385]" pageId="50" pageNumber="267">
<superScript id="7C259B9BFFA3F76E076F151E4837CB1D" attach="none" box="[1306,1314,1371,1385]" fontSize="6" pageId="50" pageNumber="267">1</superScript>
</emphasis>
<emphasis id="B924EAC1FFA3F76E075215274850CB1D" bold="true" box="[1319,1349,1371,1394]" pageId="50" pageNumber="267">
<subScript id="17D43496FFA3F76E07521527483ACB04" attach="none" box="[1319,1327,1378,1392]" fontSize="6" pageId="50" pageNumber="267">8</subScript>
<superScript id="7C259B9BFFA3F76E0748151E4850CB1D" attach="right" box="[1341,1349,1371,1385]" fontSize="6" pageId="50" pageNumber="267">1</superScript>
</emphasis>
<emphasis id="B924EAC1FFA3F76E073F15274847CB04" bold="true" box="[1354,1362,1378,1392]" pageId="50" pageNumber="267">
<subScript id="17D43496FFA3F76E073F15274847CB04" attach="left" box="[1354,1362,1378,1392]" fontSize="6" pageId="50" pageNumber="267">4</subScript>
</emphasis>
of length of penial chamber), broad, with pointed to rounded tip. Penial wall pustules absent. Main stimulatory pilaster not well differentiated, narrow, smooth, on apical to median portion of penial chamber. One to three additional, smooth, narrow pilasters on apical portion. Vas deferens entering penial sheath in upper third. Vagina rather short, medially to posteriorly inflated; inner vaginal wall predominantly smooth, with few weakly developed longitudinal pilasters. Spermathecal duct moderately wide, internally smooth. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue, internally smooth, with thin wall. Free oviduct comprising about half of anterior part of oviduct; more or less straight. Spermoviduct much longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<paragraph id="8BEF36D3FFA3F76F013217084CC0C968" blockId="50.[839,1467,1869,2010]" lastBlockId="51.[121,749,1709,1820]" lastPageId="51" lastPageNumber="268" pageId="50" pageNumber="267">
<emphasis id="B924EAC1FFA3F76E013217084929C911" bold="true" box="[839,1084,1869,1893]" pageId="50" pageNumber="267">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFA3F76E063B170B49C3C911" box="[1102,1238,1870,1893]" captionStart="Figure 77" captionStartId="50.[166,229,1086,1107]" captionTargetBox="[166,1466,151,1073]" captionTargetId="figure-74@50.[166,1466,151,1073]" captionTargetPageId="50" captionText="Figure 77. SEM photographs of shell and radula of Amplirhagada camdenensis n.sp., paratype AMS C463722: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first three whorls viewed from above (scale 1 mm); (C) close-up showing central and inner lateral teeth (scale 10 µm); (D) close-up showing inner and middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239255" httpUri="https://zenodo.org/record/5239255/files/figure.png" pageId="50" pageNumber="267">Fig. 77CD</figureCitation>
). Rectangular. Tooth formula C+16+4+?. With 122 rows of teeth, 27.4 rows per mm (n = 1). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular to ovate mesocones, length equal to base of tooth; ectocones and endocones vestigial. Marginal teeth with triangular to broadly flattened mesocones; ectocones shorter and narrower than mesocones; endocones vestigial.
</paragraph>
<caption id="DF2F665BFFA3F76E02D317C94C6AC9A3" ID-DOI="http://doi.org/10.5281/zenodo.5239257" ID-Zenodo-Dep="5239257" httpUri="https://zenodo.org/record/5239257/files/figure.png" pageId="50" pageNumber="267" startId="50.[166,229,1932,1953]" targetBox="[175,781,1506,1912]" targetPageId="50">
<paragraph id="8BEF36D3FFA3F76E02D317C94C6AC9A3" blockId="50.[166,794,1932,2007]" pageId="50" pageNumber="267">
Figure 78. Genitalia of
<taxonomicName id="4C504D50FFA3F76E03F417C84F9EC9D6" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[385,651,1933,1954]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="50" pageNumber="267" phylum="Mollusca" rank="species" species="camdenensis" status="sp. nov.">
<emphasis id="B924EAC1FFA3F76E03F417C84F9EC9D6" box="[385,651,1933,1954]" italics="true" pageId="50" pageNumber="267">Amplirhagada camdenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA3F76E00FA17C94FAAC9D5" box="[655,703,1932,1953]" pageId="50" pageNumber="267" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463722 (25 May, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFA2F76F020C16554866C801" ID-DOI="http://doi.org/10.5281/zenodo.5239261" ID-Zenodo-Dep="5239261" httpUri="https://zenodo.org/record/5239261/files/figure.png" pageId="51" pageNumber="268" startId="51.[121,184,1552,1573]" targetBox="[121,1421,151,1543]" targetPageId="51">
<paragraph id="8BEF36D3FFA2F76F020C16554866C801" blockId="51.[121,1421,1552,1653]" pageId="51" pageNumber="268">
Figure 79. SEM photographs of
<taxonomicName id="4C504D50FFA2F76F03DA16554FD7C851" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[431,706,1552,1573]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="268" phylum="Mollusca" rank="species" species="kimberleyana" status="sp. nov.">
<emphasis id="B924EAC1FFA2F76F03DA16554FD7C851" box="[431,706,1552,1573]" italics="true" pageId="51" pageNumber="268">Amplirhagada kimberleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA2F76F00B216554FE0C851" box="[711,757,1552,1573]" pageId="51" pageNumber="268" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FFA2F76F017616554E3AC851" box="[771,815,1552,1573]" italics="true" pageId="51" pageNumber="268">AD</emphasis>
) Shell, paratype FMNH 219055: (
<emphasis id="B924EAC1FFA2F76F060F1655499CC851" box="[1146,1161,1552,1573]" italics="true" pageId="51" pageNumber="268">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFA2F76F0323166E4C70C834" box="[342,357,1579,1600]" italics="true" pageId="51" pageNumber="268">B</emphasis>
) sculpture on first three whorls viewed from above (scale 1 mm); (
<emphasis id="B924EAC1FFA2F76F01AF166E4EFFC834" box="[986,1002,1579,1600]" italics="true" pageId="51" pageNumber="268">C</emphasis>
) close-up showing sculpture on penultimate whorl (scale 200 µm); (
<emphasis id="B924EAC1FFA2F76F032316004C72C82E" box="[342,359,1605,1626]" italics="true" pageId="51" pageNumber="268">D</emphasis>
) close-up showing surface of last whorl (scale 200 µm). (
<emphasis id="B924EAC1FFA2F76F01F116004E86C82E" box="[900,915,1605,1626]" italics="true" pageId="51" pageNumber="268">E</emphasis>
) Jaw, paratype AMS C463760 (scale 100 µm). (
<emphasis id="B924EAC1FFA2F76F072C16004890C82E" box="[1369,1413,1605,1626]" italics="true" pageId="51" pageNumber="268">FG</emphasis>
) Radula, paratype AMS C463760: (
<emphasis id="B924EAC1FFA2F76F03B216254CC3C801" box="[455,470,1632,1653]" italics="true" pageId="51" pageNumber="268">F</emphasis>
) central and lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FFA2F76F011416254E67C801" box="[865,882,1632,1653]" italics="true" pageId="51" pageNumber="268">G</emphasis>
) outer lateral and inner marginal teeth (scale 20 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFA2F76F020C176F4C00C9A0" blockId="51.[121,748,1834,2004]" pageId="51" pageNumber="268">
<emphasis id="B924EAC1FFA2F76F020C176F4C62C936" bold="true" box="[121,375,1834,1858]" pageId="51" pageNumber="268">Comparative remarks</emphasis>
. Material of this species has been labelled as “
<taxonomicName id="4C504D50FFA2F76F0373170D4CDEC92B" box="[262,459,1864,1887]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="268" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FFA2F76F0373170D4CB7C92B" box="[262,418,1864,1887]" italics="true" pageId="51" pageNumber="268">Amplirhagada</emphasis>
sp.
</taxonomicName>
65” by Solem. It has the smallest of all dome-shaped shells with a significantly higher H/D ratio as most other species (
<tableCitation id="C6D20368FFA2F76F03B917C64F0EC9EE" box="[460,539,1923,1946]" captionStart="Table 1" captionStartId="61.[120,176,474,495]" captionText="Table 1. Shell parameters of Amplirhagada species, given in the form maximumminimum (mean±standard deviation) measures of shells for n specimens." pageId="51" pageNumber="268">Table 1</tableCitation>
). Absence of penial wall pustules and pattern of smooth, longitudinal pilasters are diagnostic.
</paragraph>
</subSubSection>
</treatment>
</document>