191 lines
26 KiB
XML
191 lines
26 KiB
XML
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<mods:title id="AB6DE195272F62BEA3AB0807F02C71F0">Integrative taxonomic revision of the African taxa of the Racotis Moore, 1887 generic complex (Lepidoptera, Geometridae, Ennominae, Boarmiini)</mods:title>
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<mods:namePart id="6570443D7E94D1F914A3AB8A06ACF72E">László, Gyula M.</mods:namePart>
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<mods:namePart id="B9BC8C035E531FD1EC19D4F5161A72B0">Hausmann, Axel</mods:namePart>
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<mods:namePart id="A068CB8B96B425434377EC5F7E7CFF80">Karisch, Timm</mods:namePart>
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<mods:title id="E4BE6F9366C605B84503542DEB9F14A7">Zootaxa</mods:title>
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<mods:part id="A971E2BEDE5CA1B08509E536498A7721">
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<mods:date id="9EE312634D987326CEE7F6CFD384F5AE">2023</mods:date>
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<mods:number id="23D67D16F06B8C3A4A04E140AB2E392A">2023-06-26</mods:number>
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<mods:number id="0D1B5B026EC2E09F0CA5B09FDFFE06CB">5308</mods:number>
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<mods:url id="3A84FD65E1C651062E57C5BDCDDFA985">http://dx.doi.org/10.11646/zootaxa.5308.1.1</mods:url>
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<mods:classification id="765DEE2E6D43ECE32B465ACDF8CC3D9F">journal article</mods:classification>
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<treatment id="03EB442B5D74FFF657F55793FE36FE55" ID-DOI="http://doi.org/10.5281/zenodo.8117378" ID-Zenodo-Dep="8117378" LSID="urn:lsid:plazi:treatment:03EB442B5D74FFF657F55793FE36FE55" httpUri="http://treatment.plazi.org/id/03EB442B5D74FFF657F55793FE36FE55" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
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<subSubSection id="C358A6B65D74FFF557F55793FE60FF35" box="[151,455,151,178]" pageId="73" pageNumber="74" type="nomenclature">
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<paragraph id="8BFDF53D5D74FFF557F55793FE60FF35" blockId="73.[151,455,151,178]" box="[151,455,151,178]" pageId="73" pageNumber="74">
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<heading id="D0B542515D74FFF557F55793FE60FF35" bold="true" box="[151,455,151,178]" fontSize="11" level="2" pageId="73" pageNumber="74" reason="4">
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<emphasis id="B936292F5D74FFF557F55793FE60FF35" bold="true" box="[151,455,151,178]" pageId="73" pageNumber="74">
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<taxonomicName id="4C428EBE5D74FFF557F55793FEE7FF36" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,320,151,178]" family="Geometridae" genus="Sokokeracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" rank="genus" status="subgen. nov.">
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<emphasis id="B936292F5D74FFF557F55793FEE7FF36" bold="true" box="[151,320,151,178]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20594545D74FFF5562A5793FE60FF35" box="[328,455,151,177]" pageId="73" pageNumber="74" rank="subGenus">subgen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358A6B65D74FFF557F557D8FD8FFF70" box="[151,552,220,244]" pageId="73" pageNumber="74" type="type_taxon">
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<paragraph id="8BFDF53D5D74FFF557F557D8FD8FFF70" blockId="73.[151,552,220,244]" box="[151,552,220,244]" pageId="73" pageNumber="74">
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<typeStatus id="54F94B9F5D74FFF557F557D8FF6EFF70" box="[151,201,220,244]" pageId="73" pageNumber="74">Type</typeStatus>
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species:
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<taxonomicName id="4C428EBE5D74FFF5564457D8FE4FFF70" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[294,488,220,244]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="staudei" status="sp. nov.">
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<emphasis id="B936292F5D74FFF5564457D8FE4FFF70" box="[294,488,220,244]" italics="true" pageId="73" pageNumber="74">Afroracotis staudei</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D74FFF5568D57D8FD8FFF70" bold="true" box="[495,552,220,244]" pageId="73" pageNumber="74">
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<taxonomicNameLabel id="A20594545D74FFF5568D57D8FD8FFF70" box="[495,552,220,244]" pageId="73" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358A6B65D74FFF557F55625FF4AFBD0" pageId="73" pageNumber="74" type="diagnosis">
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<paragraph id="8BFDF53D5D74FFF557F55625FEEBFD90" blockId="73.[151,1437,289,1108]" pageId="73" pageNumber="74">
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<emphasis id="B936292F5D74FFF557F55625FEADFEBF" bold="true" box="[151,266,289,315]" pageId="73" pageNumber="74">Diagnosis</emphasis>
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. The external habitus of the
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<typeStatus id="54F94B9F5D74FFF5553F5626FD2AFEB8" box="[605,653,290,316]" pageId="73" pageNumber="74">type</typeStatus>
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species of the subgenus
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<taxonomicName id="4C428EBE5D74FFF554D35626FBE9FEB8" box="[945,1102,290,316]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF554D35626FBE9FEB8" box="[945,1102,290,316]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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lacks any notable characters which distinguish it from other members of the
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<taxonomicName id="4C428EBE5D74FFF555D95642FC9DFEE4" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[699,826,326,352]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF555D95642FC9DFEE4" box="[699,826,326,352]" italics="true" pageId="73" pageNumber="74">Afroracotis</emphasis>
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</taxonomicName>
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,
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<emphasis id="B936292F5D74FFF5542B5642FC53FEE4" box="[841,1012,326,352]" italics="true" pageId="73" pageNumber="74">Herbuloracotis</emphasis>
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and
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<emphasis id="B936292F5D74FFF5534F5642FB02FEE4" box="[1069,1189,326,352]" italics="true" pageId="73" pageNumber="74">Zebracotis</emphasis>
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subgenera displaying the typical pale mossy-green wing upperside densely speckled with darker brown scales and a dark marginal band, as well as a large, rounded discal spot on the wing underside. However, based on the single known specimen of the
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<typeStatus id="54F94B9F5D74FFF557F556B6FF60FE48" box="[151,199,434,460]" pageId="73" pageNumber="74">type</typeStatus>
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species, the inner two-thirds of the forewing of
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<taxonomicName id="4C428EBE5D74FFF555BA56B6FCD2FE48" box="[728,885,434,460]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF555BA56B6FCD2FE48" box="[728,885,434,460]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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are noticeably paler with more extensive whitish-grey areas rendering the forewing somewhat lighter than in the species of other subgenera with almost undetectable transverse lines.
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</paragraph>
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<paragraph id="8BFDF53D5D74FFF557A5551AFA36FCF8" blockId="73.[151,1437,289,1108]" pageId="73" pageNumber="74">
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The subgenus is well characterized by the rather unique configuration of the male genitalia expressed in the basally triangular, distally narrow quadrangular uncus bearing two short, pointed latero-apical processi, displaying a morphological transition between the short, apically bifurcate uncus of the subgenus
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<taxonomicName id="4C428EBE5D74FFF553215562FB65FD04" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1091,1218,614,640]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF553215562FB65FD04" box="[1091,1218,614,640]" italics="true" pageId="73" pageNumber="74">Afroracotis</emphasis>
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</taxonomicName>
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and the moderately long, simple uncus of
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<taxonomicName id="4C428EBE5D74FFF556F7558EFDE7FD20" box="[405,576,650,676]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="105" phylum="Arthropoda" rank="subGenus" subGenus="Herbuloracotis">
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<emphasis id="B936292F5D74FFF556F7558EFDE7FD20" box="[405,576,650,676]" italics="true" pageId="73" pageNumber="74">Herbuloracotis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C428EBE5D74FFF55518558EFD55FD20" box="[634,754,650,676]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
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<emphasis id="B936292F5D74FFF55518558EFD55FD20" box="[634,754,650,676]" italics="true" pageId="73" pageNumber="74">Zebracotis</emphasis>
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</taxonomicName>
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. The tegumen of
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<taxonomicName id="4C428EBE5D74FFF554DF558EFBFDFD20" box="[957,1114,650,676]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF554DF558EFBFDFD20" box="[957,1114,650,676]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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is considerably broader and shorter than that of the other subgenera, rather quadrangular with an almost straight distal margin, while the tegumen is rounded trapezoidal in the other
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<taxonomicName id="4C428EBE5D74FFF5557555D6FD31FD68" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[535,662,722,748]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF5557555D6FD31FD68" box="[535,662,722,748]" italics="true" pageId="73" pageNumber="74">Afroracotis</emphasis>
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</taxonomicName>
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subgenera. The valva configuration of
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<taxonomicName id="4C428EBE5D74FFF5532555D6FB43FD68" box="[1095,1252,722,748]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF5532555D6FB43FD68" box="[1095,1252,722,748]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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is unique within the genus due to the presence of an evenly curved heavily sclerotised medial crest running along the dorsal margin of the sacculus connected with the rather broad valva costa, forming a pointed, sclerotised valva apex directed medio-caudad. The vesica configuration of
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<taxonomicName id="4C428EBE5D74FFF555F2543AFC8AFCDC" box="[656,813,830,856]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF555F2543AFC8AFCDC" box="[656,813,830,856]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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is also rather distinctive, characterised by the robust, apically rounded, stick-like distal cornutus, while the cornutus is pointed, rather spike-like in the other subgenera.
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</paragraph>
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<paragraph id="8BFDF53D5D74FFF557A55482FF4AFBD0" blockId="73.[151,1437,289,1108]" pageId="73" pageNumber="74">
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Despite sharing the typical external features of the genera
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<taxonomicName id="4C428EBE5D74FFF554165483FC6FFC24" authorityName="Moore" authorityYear="1887" box="[884,968,903,928]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF554165483FC6FFC24" box="[884,968,903,928]" italics="true" pageId="73" pageNumber="74">Racotis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C428EBE5D74FFF553675482FB23FC24" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1029,1156,902,928]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF553675482FB23FC24" box="[1029,1156,902,928]" italics="true" pageId="73" pageNumber="74">Afroracotis</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="54F94B9F5D74FFF553A05482FB55FC24" box="[1218,1266,902,928]" pageId="73" pageNumber="74">type</typeStatus>
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species of the subgenus
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<taxonomicName id="4C428EBE5D74FFF5566554AEFE03FC40" box="[263,420,938,964]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="subGenus" subGenus="Sokokeracotis">
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<emphasis id="B936292F5D74FFF5566554AEFE03FC40" box="[263,420,938,964]" italics="true" pageId="73" pageNumber="74">Sokokeracotis</emphasis>
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</taxonomicName>
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has a highly distinctive male genitalia configuration suggesting the potential for a generic level classification within the
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<taxonomicName id="4C428EBE5D74FFF5569254CBFDE3FC6C" authorityName="Moore" authorityYear="1887" box="[496,580,975,1000]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D74FFF5569254CBFDE3FC6C" box="[496,580,975,1000]" italics="true" pageId="73" pageNumber="74">Racotis</emphasis>
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</taxonomicName>
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generic complex. Nevertheless, because of the unavailability of molecular data and information on female genitalia, the taxon is considered here as a member of the genus
|
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<taxonomicName id="4C428EBE5D74FFF5538F54F6FACBFB88" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1261,1388,1010,1036]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B936292F5D74FFF5538F54F6FACBFB88" box="[1261,1388,1010,1036]" italics="true" pageId="73" pageNumber="74">Afroracotis</emphasis>
|
||
</taxonomicName>
|
||
and distinguished tentatively at subgeneric rank until further material becomes available for in-depth phylogenetic studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358A6B65D74FFF657F553A1FE36FE55" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="description">
|
||
<paragraph id="8BFDF53D5D74FFF557F553A1FE8FFB3B" blockId="73.[151,296,1189,1215]" box="[151,296,1189,1215]" pageId="73" pageNumber="74">
|
||
<emphasis id="B936292F5D74FFF557F553A1FE8FFB3B" bold="true" box="[151,296,1189,1215]" pageId="73" pageNumber="74">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D74FFF557F553EAFE0DF8E8" blockId="73.[151,1437,1261,2008]" pageId="73" pageNumber="74">
|
||
<emphasis id="B936292F5D74FFF557F553EAFD28FA83" bold="true" box="[151,655,1261,1288]" pageId="73" pageNumber="74">External features of male body and wings</emphasis>
|
||
(
|
||
<figureCitation id="1379E9B85D74FFF555FD53EAFCA2FA8C" box="[671,773,1262,1288]" captionStart="FIGURES 101–120" captionStartId="65.[151,264,1643,1667]" captionTargetBox="[185,1402,188,1614]" captionTargetId="figure-17@65.[176,1412,181,1620]" captionTargetPageId="65" captionText="FIGURES 101–120. Adults, dorsal view (all in coll. ANHRT, unless otherwise stated). 101. A. smithi sp. n., HT, ♁, Zambia; 102. Ibidem, PT, ♁, Gabon; 103. Ibidem, PT, ♀, Zambia; 104. Ibidem, PT, ♀, Cameroon (ZSM); 105. A. milesi sp. n., HT, ♁, Gabon; 106. Ibidem, PT, ♁, Cameroon; 107. Ibidem, PT, ♀, Cameroon; 108. Ibidem, PT, ♁, Zambia; 109. A. ochsei sp. n., HT, ♁, Gabon; 110. Ibidem, PT, ♁, Gabon; 111. Ibidem, PT, ♀, Gabon; 112. Ibidem, PT, ♁, Gabon; 113. A. helicalis sp. n., HT, ♁, Gabon; 114. Ibidem, PT, ♁, Gabon; 115. Ibidem, PT, ♀, Gabon; 116. A. takanoi sp. n., HT, ♁, Sierra Leone; 117. Ibidem, PT, ♁, Guinea; 118. Ibidem, PT, ♁, Liberia; 119. Ibidem, PT, ♀, Liberia; 120. A. staudei sp. n., HT, ♁, Kenya (RCHS)." figureDoi="http://doi.org/10.5281/zenodo.8080943" httpUri="https://zenodo.org/record/8080943/files/figure.png" pageId="73" pageNumber="74">Figs 120</figureCitation>
|
||
,
|
||
<figureCitation id="1379E9B85D74FFF5547153EAFCE6FA8D" box="[787,833,1262,1289]" captionStart="FIGURES 191–210" captionStartId="81.[151,264,1641,1665]" captionTargetBox="[151,1433,188,1610]" captionTargetId="figure-17@81.[151,1436,181,1617]" captionTargetPageId="81" captionText="FIGURES 191–210. Adults, ventral view (all in coll. ANHRT, unless otherwise stated). 191. Afroracotis milesi sp. n., HT, ♁, Gabon; 192. Ibidem, PT, ♁, Cameroon; 193. Ibidem, PT, ♁, Zambia; 194. Ibidem, PT, ♀, Gabon; 195. A. ochsei sp. n., HT, ♁, Gabon; 196. Ibidem, PT, ♁, Gabon; 197. Ibidem, PT, ♀, Gabon; 198. A. helicalis sp. n., HT, ♁, Gabon; 199. Ibidem, PT, ♁, Gabon; 200. Ibidem, PT, ♀, Gabon; 201. A. takanoi sp. n., HT, ♁, Sierra Leone; 202. Ibidem, PT, ♁, Guinea; 203. Ibidem, PT, ♀, Ivory Coast; 204. Ibidem, PT, ♀, Liberia; 205. A. staudei sp. n., HT, ♁, Kenya (RCHS); 206. Racotis boarmiaria, LT, ♁, India (NHMUK); 207. Chorocotis ugandaria, HT of Boarmia ugandaria, ♁, Uganda (NHMUK); 208. Ibidem, ♀, Cameroon (NHMUK); 209. Colocleora breijeri, ♁, RSA (RCDS); 210. C. incauta, ♁, RSA (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.8080954" httpUri="https://zenodo.org/record/8080954/files/figure.png" pageId="73" pageNumber="74">205</figureCitation>
|
||
). Forewing length
|
||
<quantity id="4CBA58D85D74FFF5537553EAFBCCFA8C" box="[1047,1131,1262,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="73" pageNumber="74" unit="mm" value="21.0">21 mm</quantity>
|
||
. Male antenna bipectinate-fasciculate, rami moderately long, distal two-fifths filiform. Head moderately large, proboscis well-developed, labial palp short, length as the diameter of eye, porrect, first and second segment dilated, third segment rather short and thin. Compound eye moderately large. Frons, vertex, collar (patagium), tegula, thorax and abdomen colouration as of wing upperside. Legs rather long, colouration as of wing underside, full set of spurs could not be counted due to missing midlegs of the single accessed specimen. Forewing broad, triangular, costa straight, apical section slightly curved, termen very slightly arched, ventral margin almost straight, slightly arched in basal fifth. Ground colour pale mossy-green, speckled with groups of brownish and whithish scales. Basal and medial area lighter than postmedialterminal areas with more extensive whitish patches. Transverse lines diffuse, interrupted, barely detectable except for the broad, slightly sinuous, shadow-like antemedial and postmedial lines and interrupted terminal line consisting of small black dots between veins. Fringe (cilia) short, concolourous with forewing, chequered with dark brown. Discal spot obliterated, patch-like, round. Hindwing apex rounded, termen evenly arcuate, slightly scalloped, anal margin straight. Ground colour as of forewing, densely speckled with groups of brownish- and whitish-grey scales. Transverse lines undetectable except for the interrupted terminal line, consisting of short arches between veins. Discal spot absent. Fringe (cilia) short, colour as of hindwing, chequered with dark brown. Wing underside pale whitish-grey in basal two-thirds, sparsley suffused with greyish scales, distal third with broad, dark brownish-grey postmedial band; apex and medial section of forewing subterminal area with off-white patch. Transverse lines and discal spot undetectable.
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D74FFF657A55072FB81FE29" blockId="73.[151,1437,1261,2008]" lastBlockId="74.[151,1436,151,465]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
|
||
<emphasis id="B936292F5D74FFF557A55072FED4F80B" bold="true" box="[199,371,1909,1936]" pageId="73" pageNumber="74">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1379E9B85D74FFF556E15072FE4FF814" box="[387,488,1910,1936]" captionStart="FIGURES 280–284" captionStartId="93.[151,264,1824,1848]" captionTargetBox="[155,1431,189,1794]" captionTargetId="figure-17@93.[151,1436,181,1800]" captionTargetPageId="93" captionText="FIGURES 280–284. Male genitalia, a: clasping apparatus, b: aedeagus. 280. Afroracotis helicalis sp. n., PT, Gabon, slide No. LG 5399 (ANHRT); 281. A. takanoi sp. n., HT, Sierra Leone, slide No. LG 4810 (ANHRT); 282. Ibidem, PT, Guinea, slide No. LG 5445 (ANHRT); 283. Ibidem, PT, Liberia, slide No. LG 4982 (ANHRT); 284. A. staudei sp. n., HT, Kenya, slide No. LG 5540 (RCHS)." figureDoi="http://doi.org/10.5281/zenodo.8080987" httpUri="https://zenodo.org/record/8080987/files/figure.png" pageId="73" pageNumber="74">Fig. 284</figureCitation>
|
||
). Uncus short, proximal section broad-triangular, distal section produced into short, quadrangular projection, apex with short, latero-caudal teeth. Tegumen very short and conspicuously broad, more or less quadrangular, gnathos well-developed, arms narrow, medial plate triangular. Valva short, broad at base, distally tapered, costa widely sclerotised, slightly concave. Ventral margin of valva evenly convex, sparsely setose, submedial area of valva with heavily sclerotised, narrow, evenly arched longitudinal crest adjacent to sacculus, running in parallel with ventral margin, connected with costa, forming a triangular, pointed apex curved medio-caudad. Sacculus long and narrow, weakly sclerotised, without process. Juxta weakly sclerotised, consisting of two very short and narrow, parallel longitudinal plates. Vinculum very short and broad V-shaped, medially with short projection. Aedeagus medium thick, short, straight, coecum penis short, rounded, carina absent. Vesica short and narrow, bearing a small cornuti field consisting of ca. 10 short cornuti ventro-basally and a robust, almost straight, apically rounded, stick-like cornutus distally at base of thick vesica ejaculatorius.
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D77FFF657A556B3FE36FE55" blockId="74.[151,1436,151,465]" box="[199,401,439,465]" pageId="74" pageNumber="75">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |