treatments-xml/data/03/DC/90/03DC9030FFA3FFAD6EFAFD96FBE1E4E4.xml
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<document id="53F3328AF7A838D681A3A631DE1013C7" ID-DOI="10.11646/zootaxa.5397.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10468563" ID-ZooBank="C55178A5-DABB-4615-8443-59B388F99A36" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704706602791" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2024" docId="03DC9030FFA3FFAD6EFAFD96FBE1E4E4" docLanguage="en" docName="zootaxa.5397.2.2.pdf" docOrigin="Zootaxa 5397 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysdera azerbajdzhanica Charitonov 1956" docType="treatment" docVersion="2" lastPageNumber="199" masterDocId="FFE5E848FFA7FFA96E6DFFB4FFFBE051" masterDocTitle="New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records" masterLastPageNumber="217" masterPageNumber="195" pageNumber="199" updateTime="1704707340498" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0E3D13E182CD50C81453F4A1D2E7B7C4">New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records</mods:title>
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<mods:namePart id="0F2369DC6506B03B8F88822C134BCEE7">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="D5511130F2EAA1D2AC5BA2AB37DFC74F">Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa. &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Institute for Biological Problems of the North, Portovaya Str. 18, Magadan 685000, Russia.</mods:affiliation>
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<taxonomicName id="4C755AA5FFA3FFAD6EFAFD96FD5DE26D" ID-CoL="388YK" authority="Charitonov, 1956" authorityName="Charitonov" authorityYear="1956" box="[151,678,546,573]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="azerbajdzhanica">
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<emphasis id="B901FD34FFA3FFAD6EFAFD96FE3CE26C" bold="true" box="[151,455,546,573]" italics="true" pageId="4" pageNumber="199">Dysdera azerbajdzhanica</emphasis>
<bibRefCitation id="EFE45CD7FFA3FFAD6FA3FD96FD5DE26D" author="Charitonov, D. E." box="[462,678,546,572]" pageId="4" pageNumber="199" pagination="17 - 39" refId="ref9811" refString="Charitonov, D. E. (1956) Obzor paukov semeistva Dysderidae fauny SSSR. Uchenye Zapiski Molotovskogo Gosudarstvennogo Universiteta Imeni M. A. Gor'kogo, 10, 17 - 39. [in Russian]" type="journal article" year="1956">Charitonov, 1956</bibRefCitation>
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<figureCitation id="134E3DA3FFA3FFAD6EFAFDF3FEEEE230" box="[151,277,583,609]" captionStart="FIGURE 3" captionStartId="3.[152,255,1935,1959]" captionTargetBox="[161,1435,1382,1896]" captionTargetId="figure-93@3.[151,1436,1354,1911]" captionTargetPageId="3" captionText="FIGURE 3. Male of Dysdera azerbajdzhanica, left bulb. A retrolateral B prolateral C dorsal D ventral. Scale bar = 0.2 mm. Abbreviations: AL—additional lateral sheet, C—crest, DH—distal haematodocha, L—lateral sheet, LML—lateral margin of the lateral sheet, P—posterior apophysis, T—tegulum." figureDoi="http://doi.org/10.5281/zenodo.10468571" httpUri="https://zenodo.org/record/10468571/files/figure.png" pageId="4" pageNumber="199">Figs 3AD</figureCitation>
,
<figureCitation id="134E3DA3FFA3FFAD6F4CFDF3FE8EE230" box="[289,373,583,609]" captionStart="FIGURE 11" captionStartId="10.[152,255,1860,1884]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@10.[151,1435,181,1837]" captionTargetPageId="10" captionText="FIGURE 11. Males of Dysdera azerbajdzhanica (AD), D. mazini (E, F), and D. hungarica atra (GI), SEM images of left palps. A, E, I bulbs, retrolateral C, G close-ups of psembolus, retrolateral H same, retrodorsal D, F same, dorsal B cymbium, ventral. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10468587" httpUri="https://zenodo.org/record/10468587/files/figure.png" pageId="4" pageNumber="199">11AD</figureCitation>
,
<figureCitation id="134E3DA3FFA3FFAD6FEDFDF3FE54E230" box="[384,431,583,609]" captionStart="FIGURE 13" captionStartId="12.[152,255,1920,1944]" captionTargetBox="[151,1436,921,1897]" captionTargetId="figure-421@12.[151,1436,921,1897]" captionTargetPageId="12" captionText="FIGURE 13. Females of Dysdera atabekia sp. n. (A), D. azerbajdzhanica (B), D. dunini (C), D. daghestanica (D), D. dunini (E), and D. hungarica atra (F), dorsal view of endogynes. Scale bars = 0.2 mm.Abbreviations: DA—dorsal arch, Re—receptacle, TB—transverse bar." figureDoi="http://doi.org/10.5281/zenodo.10468591" httpUri="https://zenodo.org/record/10468591/files/figure.png" pageId="4" pageNumber="199">13B</figureCitation>
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<emphasis id="B901FD34FFA3FFAD6EFAFD3FFE61E2F2" box="[151,410,651,675]" italics="true" pageId="4" pageNumber="199">Dysdera azerbajdzhanica</emphasis>
<bibRefCitation id="EFE45CD7FFA3FFAD6FCCFD3FFD81E2F2" author="Charitonov, D. E." box="[417,634,651,675]" pageId="4" pageNumber="199" pagination="17 - 39" refId="ref9811" refString="Charitonov, D. E. (1956) Obzor paukov semeistva Dysderidae fauny SSSR. Uchenye Zapiski Molotovskogo Gosudarstvennogo Universiteta Imeni M. A. Gor'kogo, 10, 17 - 39. [in Russian]" type="journal article" year="1956">Charitonov, 1956: 34</bibRefCitation>
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, fig. 14 (
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).
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<treatmentCitationGroup id="AB650608FFA3FFAD6EFAFD1EFDBCE2B2" pageId="4" pageNumber="199">
<taxonomicName id="4C755AA5FFA3FFAD6EFAFD1EFE67E293" ID-CoL="388YK" authorityName="Charitonov" authorityYear="1956" box="[151,412,682,706]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="azerbajdzhanica">
<emphasis id="B901FD34FFA3FFAD6EFAFD1EFE67E293" box="[151,412,682,706]" italics="true" pageId="4" pageNumber="199">Dysdera azerbajdzhanica</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AD40737FFA3FFAD6FC1FD1EFC8EE293" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[428,885,682,706]" page="222" pageId="4" pageNumber="199" year="1988">
<bibRefCitation id="EFE45CD7FFA3FFAD6FC1FD1EFC8EE293" author="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R." box="[428,885,682,706]" pageId="4" pageNumber="199" pagination="141 - 269" refId="ref9849" refString="Deeleman-Reinhold, C. L. &amp; Deeleman, P. R. (1988) Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie, 131, 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988: 222</bibRefCitation>
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, fig. 263, 266 (
<specimenCount id="9D73EAAFFFA3FFAD6A76FD1FFBC0E295" box="[1051,1083,683,708]" count="3" pageId="4" pageNumber="199" type="female">³♀</specimenCount>
);
<treatmentCitation id="0AD40737FFA3FFAD6A38FD1EFAFCE293" author="Dunin, P. M." box="[1109,1287,682,706]" page="66" pageId="4" pageNumber="199" year="1992">
<bibRefCitation id="EFE45CD7FFA3FFAD6A38FD1EFAFCE293" author="Dunin, P. M." box="[1109,1287,682,706]" pageId="4" pageNumber="199" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin 1992b: 66</bibRefCitation>
</treatmentCitation>
, fig. 11 (
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);
<treatmentCitation id="0AD40737FFA3FFAD6EAAFD7CFE76E2B1" author="Mcheidze, T. S." box="[199,397,712,737]" page="77" pageId="4" pageNumber="199" year="1997">
<bibRefCitation id="EFE45CD7FFA3FFAD6EAAFD7CFE76E2B1" author="Mcheidze, T. S." box="[199,397,712,737]" pageId="4" pageNumber="199" refId="ref10876" refString="Mcheidze, T. S. (1997) [Spiders of Georgia: Systematics, Ecology, Zoogeographic Review]. Tbilisi University, Tbilisi, 390 pp. [in Georgian]" type="book" year="1997">Mcheidze 1997: 77</bibRefCitation>
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, figs 69, 70 (
<specimenCount id="9D73EAAFFFA3FFAD6C78FD7EFDCDE2B2" box="[533,566,714,739]" count="3" pageId="4" pageNumber="199" type="female">³♀</specimenCount>
).
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<emphasis id="B901FD34FFA3FFAD6EFAFCBAFEFDE379" bold="true" box="[151,262,782,808]" pageId="4" pageNumber="199">Material.</emphasis>
<materialsCitation id="3B1D2B7BFFA3FFAD6F79FCBBFCBAE378" box="[276,833,783,811]" collectionCode="ZMMU" country="Azerbaijan" location="Baku Region" pageId="4" pageNumber="199" specimenCount="1" specimenCount-female="1" stateProvince="Baku">
<collectingCountry id="F36261B6FFA3FFAD6F79FCBBFE3EE378" box="[276,453,783,809]" name="Azerbaijan" pageId="4" pageNumber="199">AZERBAIJAN</collectingCountry>
:
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</emphasis>
: 3³
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(
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)
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,
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,
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, [
<geoCoordinate id="EE4147E1FFA3FFAD6B50FCBBFA67E378" box="[1341,1436,783,809]" degrees="40" direction="north" minutes="30" orientation="latitude" pageId="4" pageNumber="199" precision="925" value="40.5">40°30'N</geoCoordinate>
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], 04.2010 (
<collectorName id="268044F0FFA3FFAD6F1CFC87FDE8E31C" box="[369,531,819,845]" pageId="4" pageNumber="199">N. Snegovaya</collectorName>
); 2³ (
<collectionCode id="ED64B9E3FFA3FFAD6C3BFC87FD48E31C" box="[598,691,819,845]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="199" type="Museum">ZMMU</collectionCode>
)
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,
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same locality, 03.2010 (
<collectorName id="268044F0FFA3FFAD6DA2FC87FB8AE31C" box="[975,1137,819,845]" pageId="4" pageNumber="199">N. Snegovaya</collectorName>
); 1³ (
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)
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,
<materialsCitation id="3B1D2B7BFFA3FFAD6B49FC87FC5BE320" collectorName="N. Snegovaya" country="Azerbaijan" latitude="40.5" location="Institute of Zoology" longLatPrecision="1162" longitude="49.883335" pageId="4" pageNumber="199" specimenCount="1" stateProvince="Baku">
<collectingRegion id="49B1EFC4FFA3FFAD6B49FC87FA99E31C" box="[1316,1378,819,845]" country="Azerbaijan" name="Baku" pageId="4" pageNumber="199">Baku</collectingRegion>
, near
<location id="8EAA77FDFFA3FFAD6EFAFCE3FE83E320" LSID="urn:lsid:plazi:treatment:03DC9030FFA3FFAD6EFAFD96FBE1E4E4:8EAA77FDFFA3FFAD6EFAFCE3FE83E320" box="[151,376,855,881]" country="Azerbaijan" latitude="40.5" longLatPrecision="1162" longitude="49.883335" name="Institute of Zoology" pageId="4" pageNumber="199" stateProvince="Baku">Institute of Zoology</location>
, [
<geoCoordinate id="EE4147E1FFA3FFAD6FE7FCE3FE12E320" box="[394,489,855,881]" degrees="40" direction="north" minutes="30" orientation="latitude" pageId="4" pageNumber="199" precision="925" value="40.5">40°30'N</geoCoordinate>
<geoCoordinate id="EE4147E1FFA3FFAD6F9DFCE2FDB2E320" box="[496,585,854,881]" degrees="49" direction="east" minutes="53" orientation="longitude" pageId="4" pageNumber="199" precision="925" value="49.883335">49°53'E</geoCoordinate>
], 0304.2010 (
<collectorName id="268044F0FFA3FFAD6C9BFCE3FC63E320" box="[758,920,855,881]" pageId="4" pageNumber="199">N. Snegovaya</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFA3FFAD6EAAFCCEFB36E474" pageId="4" pageNumber="199" type="discussion">
<paragraph id="8BCA2126FFA3FFAD6EAAFCCEFB36E474" blockId="4.[151,1437,782,1205]" pageId="4" pageNumber="199">
<emphasis id="B901FD34FFA3FFAD6EAAFCCEFEB4E3C5" bold="true" box="[199,335,890,916]" pageId="4" pageNumber="199">Comments.</emphasis>
This species has been considered in four taxonomic publications (
<bibRefCitation id="EFE45CD7FFA3FFAD6A47FCCFFB52E3C4" author="WSC" box="[1066,1193,891,917]" pageId="4" pageNumber="199" refId="ref11291" refString="WSC (2023) World Spider Catalog. Version 24. Natural History Museum Bern, online at http: // wsc. nmbe. ch (accessed 23 May 2023) https: // doi. org / 10.24436 / 2" type="url" year="2023">WSC 2023</bibRefCitation>
) and is relatively well studied. A large amount of material collected in the type locality,
<collectingRegion id="49B1EFC4FFA3FFAD6DE7FC2BFC33E3E8" box="[906,968,927,953]" country="Azerbaijan" name="Baku" pageId="4" pageNumber="199">Baku</collectingRegion>
, has shown some variability in spination (
<bibRefCitation id="EFE45CD7FFA3FFAD6EF2FC77FECCE38C" author="Dunin, P. M." box="[159,311,963,989]" pageId="4" pageNumber="199" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin 1992b</bibRefCitation>
), although most of the specimens have spination similar to that of the
<typeStatus id="54CE9F84FFA3FFAD6A29FC77FB51E38C" box="[1092,1194,963,989]" pageId="4" pageNumber="199" type="holotype">holotype</typeStatus>
. In the lowland of the Kura-Aras (
<collectingCountry id="F36261B6FFA3FFAD6F70FC53FE65E450" box="[285,414,999,1025]" name="Azerbaijan" pageId="4" pageNumber="199">Azerbaijan</collectingCountry>
), it inhabits the dry foothills of the Caucasus Major up to
<quantity id="4C8D8CC3FFA3FFAD6A43FC53FB7DE450" box="[1070,1158,999,1025]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="199" unit="m" value="1000.0">1000 m</quantity>
, and can be found under stones, in soil cracks, rodent burrows, and often in cotton or cereal (grain) fields (
<bibRefCitation id="EFE45CD7FFA3FFAD6A4AFBBFFB3AE474" author="Dunin, P. M." box="[1063,1217,1035,1061]" pageId="4" pageNumber="199" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin 1992b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFA3FFAD6EAAFB9AFBE1E4E4" pageId="4" pageNumber="199" type="distribution">
<paragraph id="8BCA2126FFA3FFAD6EAAFB9AFBE1E4E4" blockId="4.[151,1437,782,1205]" pageId="4" pageNumber="199">
<emphasis id="B901FD34FFA3FFAD6EAAFB9AFE9AE419" bold="true" box="[199,353,1070,1096]" pageId="4" pageNumber="199">Distribution.</emphasis>
Caucasus (from ca. 43ºN to
<geoCoordinate id="EE4147E1FFA3FFAD6CCAFB9BFCFEE418" box="[679,773,1071,1097]" degrees="38" direction="north" minutes="40" orientation="latitude" pageId="4" pageNumber="199" precision="925" value="38.666668">38º40'N</geoCoordinate>
). There are over 30 records of this species in the Caucasus region, including
<collectingCountry id="F36261B6FFA3FFAD6F31FBE7FE21E43C" box="[348,474,1107,1133]" name="Azerbaijan" pageId="4" pageNumber="199">Azerbaijan</collectingCountry>
,
<collectingCountry id="F36261B6FFA3FFAD6F88FBE7FDB9E43C" box="[485,578,1107,1133]" name="Georgia" pageId="4" pageNumber="199">Georgia</collectingCountry>
, and the North Caucasus (Dagestan, North Ossetia-Alania) (
<bibRefCitation id="EFE45CD7FFA3FFAD6A8AFBE7FAA4E43C" author="Otto, S." box="[1255,1375,1107,1133]" pageId="4" pageNumber="199" refId="ref10910" refString="Otto, S. (2022) Caucasian Spiders. A faunistic database on the spiders of the Caucasus. Version February 2022. Available from: https: // caucasus-spiders. info / (accessed 23 May 2023)" type="url" year="2022">Otto 2022</bibRefCitation>
). The majority of the records are from the eastern part of the region. However, the record from
<collectingRegion id="49B1EFC4FFA3FFAD6AE9FBC3FB2BE4C0" box="[1156,1232,1143,1169]" country="Georgia" name="Ajaria" pageId="4" pageNumber="199">Adjara</collectingRegion>
in
<collectingCountry id="F36261B6FFA3FFAD6A98FBC3FAB4E4C0" box="[1269,1359,1143,1169]" name="Georgia" pageId="4" pageNumber="199">Georgia</collectingCountry>
stands out from the other known localities and may potentially refer to another species.
</paragraph>
</subSubSection>
</treatment>
</document>