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<document id="7AD5B73A7495853B503840B4E5F81DF6" ID-CLB-Dataset="6390" ID-DOI="10.5252/g2014n4a4" ID-GBIF-Dataset="8bc26961-9853-4113-a63d-4cf691aa7100" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538625" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150850454" checkinUser="felipe" docAuthor="Vianey-Liaud, Monique, Comte, Bernard, Marandat, Bernard, Peigné, Stéphane, Rage, Jean-Claude &amp; Sudre, Jean" docDate="2014" docId="03D6987B4454B10BFD67127AFBC7E6E5" docLanguage="en" docName="Geodiversitas.36.4.565-622.pdf" docOrigin="Geodiversitas 36 (4)" docSource="http://dx.doi.org/10.5252/g2014n4a4" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Comtia Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre 2014, n. gen." docType="treatment" docVersion="7" lastPageNumber="604" masterDocId="FFEFE0034471B12CFF8F106FFF81E50F" masterDocTitle="A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)" masterLastPageNumber="622" masterPageNumber="565" pageNumber="602" updateTime="1698929460103" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="1D6E062B68E794365B28930D7693621C">A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)</mods:title>
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<mods:namePart id="6DD06C855CAB321FD7B4A0DE955F10BA">Vianey-Liaud, Monique</mods:namePart>
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<mods:namePart id="D5C55BE49CAC16DD42DFBFF117B1DBD9">Comte, Bernard</mods:namePart>
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<subSubSection id="C3657AE64454B109FD67127AFBF7E720" box="[744,1142,533,559]" pageId="37" pageNumber="602" type="nomenclature">
<paragraph id="8BC0296D4454B109FD67127AFBF7E720" blockId="37.[744,1142,533,559]" box="[744,1142,533,559]" pageId="37" pageNumber="602">
<heading id="D0889E014454B109FD67127AFBF7E720" box="[744,1142,533,559]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="602" reason="2">
Genus
<taxonomicName id="4C7F52EE4454B109FCB9127AFBA0E720" authority="Vianey-Liaud" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[822,1057,533,559]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B90BF57F4454B109FCB9127AFC05E720" box="[822,900,533,559]" italics="true" pageId="37" pageNumber="602">Comtia</emphasis>
Vianey-Liaud
</taxonomicName>
,
<taxonomicNameLabel id="A23848044454B109FBA3127AFBF7E720" box="[1068,1142,533,559]" pageId="37" pageNumber="602" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A1221FCEEE7BF" pageId="37" pageNumber="602" type="diagnosis">
<paragraph id="8BC0296D4454B109FD2A1221FBABE769" blockId="37.[677,1066,590,614]" box="[677,1066,590,614]" pageId="37" pageNumber="602">DIAGNOSIS. — Ŋat of the type-species.</paragraph>
<paragraph id="8BC0296D4454B109FD2A1210FCEEE7BF" blockId="37.[677,1209,637,688]" pageId="37" pageNumber="602">
SPECIES INCLUDED. —
<taxonomicName id="4C7F52EE4454B109FC091211FC74E79A" authority="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre, 2014" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[902,1013,637,661]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="species" species="bernardi" status="sp. nov.">
<emphasis id="B90BF57F4454B109FC091211FC74E79A" box="[902,1013,637,661]" italics="true" pageId="37" pageNumber="602">C. bernardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23848044454B109FC741211FBB2E79A" box="[1019,1075,638,661]" pageId="37" pageNumber="602" rank="species">n. sp.</taxonomicNameLabel>
; ṙ
<taxonomicName id="4C7F52EE4454B109FBC91211FCEAE7BF" authority="(Freudenberg 1941)" baseAuthorityName="Freudenberg" baseAuthorityYear="1941" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B90BF57F4454B109FBC91211FB38E79A" box="[1094,1209,638,661]" italics="true" pageId="37" pageNumber="602">C. giganteus</emphasis>
(
<bibRefCitation id="EFEE549C4454B109FD2412F7FCE2E7BF" author="FREUDENBERG H." box="[683,867,664,688]" pageId="37" pageNumber="602" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg 1941</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD6D137DFC66E642" pageId="37" pageNumber="602" type="description">
<paragraph id="8BC0296D4454B109FD6D137DFC66E642" blockId="37.[738,1148,786,845]" pageId="37" pageNumber="602">
<taxonomicName id="4C7F52EE4454B109FD6D137DFBB7E623" authority="Vianey-Liaud" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[738,1078,786,813]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="species" species="bernardi" status="sp. nov.">
<emphasis id="B90BF57F4454B109FD6D137DFC18E622" bold="true" box="[738,921,786,813]" italics="true" pageId="37" pageNumber="602">Comtia bernardi</emphasis>
Vianey-Liaud
</taxonomicName>
,
<taxonomicNameLabel id="A23848044454B109FBCE137CFBFDE622" box="[1089,1148,787,813]" pageId="37" pageNumber="602" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="134435E84454B109FC0F135CFC5EE642" box="[896,991,819,845]" captionStart="FIG" captionStartId="40.[113,124,1185,1203]" captionTargetBox="[129,1209,215,1142]" captionTargetId="figure-215@40.[661,936,494,775]" captionTargetPageId="40" captionText="FIG. 15. — A, Comtia bernardi Vianey-Liaud, n. gen., n. sp., right M1-2, from Saint-Privat-des-Vieux (Gard; Lower Chattian, MP 26), SPV190, occlusal view; B-D,? cf. Aplodontidae indet. B, SPV187, anterior fragment of lower molar: B1, occlusal, B2, mesial, B3, lin- gual,B4, distal, B5, labial;C, SPV188,posterior? fragment of lower molar: C1, occlusal,C2, mesial,C3, lingual,C4, distal;D, SPV189, fragment of upper tooth: D1, occlusal, D2, lingual, D3, labial, D4, distal. Scale bars: A, 1.5 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4538659" httpUri="https://zenodo.org/record/4538659/files/figure.png" pageId="37" pageNumber="602">Fig. 15A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A1303FB3BE68C" box="[677,1210,875,899]" pageId="37" pageNumber="602" type="etymology">
<paragraph id="8BC0296D4454B109FD2A1303FB3BE68C" blockId="37.[677,1210,875,899]" box="[677,1210,875,899]" pageId="37" pageNumber="602">ETYMOLOGY. — Dedicated to the late Bernard Comte.</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A13F4FC16E6F2" pageId="37" pageNumber="602" type="materials_examined">
<paragraph id="8BC0296D4454B109FD2A13F4FB0CE6BC" blockId="37.[677,1165,923,947]" box="[677,1165,923,947]" pageId="37" pageNumber="602">
<materialsCitation id="3B1723304454B109FD2A13F4FB0CE6BC" ID-GBIF-Occurrence="3122035301" box="[677,1165,923,947]" collectionCode="SPV, UM" collectorName="Right M &amp; Coll." pageId="37" pageNumber="602" specimenCode="SPV190, UM2" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C497CF4454B109FD2A13F4FC8FE6BE" box="[677,782,923,946]" pageId="37" pageNumber="602" type="holotype">HOLOTYPE</typeStatus>
. —
<collectorName id="268A4CBB4454B109FCB513F4FC11E6BC" box="[826,912,923,947]" pageId="37" pageNumber="602">Right M</collectorName>
ṙ1 (
<specimenCode id="DBD981164454B109FC3E13F4FB86E6BC" box="[945,1031,923,947]" collectionCode="SPV" pageId="37" pageNumber="602">SPV190</specimenCode>
),
<collectorName id="268A4CBB4454B109FB9713F4FBCBE6BC" box="[1048,1098,923,947]" pageId="37" pageNumber="602">Coll.</collectorName>
<specimenCode id="DBD981164454B109FBC013F4FB08E6BC" box="[1103,1161,923,947]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="37" pageNumber="602">UM2</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC0296D4454B109FD2A13A4FC16E6F2" blockId="37.[675,1210,971,1021]" pageId="37" pageNumber="602">
<collectionCode id="ED6EB1A84454B109FD2A13A4FCAEE6EE" box="[677,815,971,994]" pageId="37" pageNumber="602">TYPE LOCALITY</collectionCode>
. — Saint-Privat-des-Vieux (Gard,
<taxonomicName id="4C7F52EE4454B109FBE113A4FB31E6EC" authority="Henri" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[1134,1200,971,995]" genus="France" pageId="37" pageNumber="566" rank="genus" status="gen. nov.">France</taxonomicName>
), lower Chattian (
<collectionCode id="ED6EB1A84454B109FCCD138AFCE9E6F2" box="[834,872,997,1021]" country="0" name="Mohonk Preserve, Inc." pageId="37" pageNumber="602">MP</collectionCode>
26).
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A1479FC43E05F" pageId="37" pageNumber="602" type="diagnosis">
<paragraph id="8BC0296D4454B109FD2A1479FCA4E00E" blockId="37.[675,1212,1045,1572]" pageId="37" pageNumber="602">
DIAGNOSIS. — Squirrel of large size (M1-2: L × w
<emphasis id="B90BF57F4454B109FB241479FB38E122" box="[1195,1209,1046,1069]" italics="true" pageId="37" pageNumber="602">=</emphasis>
3.4 ×
<quantity id="4C8784884454B109FD51145FFCBAE147" box="[734,827,1072,1096]" metricMagnitude="-3" metricUnit="m" metricValue="4.02" pageId="37" pageNumber="602" unit="mm" value="4.02">4.02 mm</quantity>
); lingual flank of the tooth higher than the labial one. Cusps high and acute (parastyle, paracone, mesostyle, metacone, small hypocone and protocone); one small paraconule, two metaconules stronger than the paraconule; straight protoloph and metaloph higher than the anterior and posterior cinguli; enamel slightly wrinkled all along the cusps and lophs as well as in the basins. Differs from:
</paragraph>
<paragraph id="8BC0296D4454B109FD2A156BFC43E05F" blockId="37.[675,1212,1045,1572]" pageId="37" pageNumber="602">
<taxonomicName id="4C7F52EE4454B109FD38156CFC22E014" authority="Heissig, 1979" authorityName="Heissig" authorityYear="1979" box="[695,931,1283,1307]" class="Mammalia" family="Sciuridae" genus="Oligopetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4454B109FD38156CFC94E014" box="[695,789,1283,1307]" italics="true" pageId="37" pageNumber="602">Oligopetes</emphasis>
Heissig, 1979
</taxonomicName>
, in the absence of postparacrista connected to the mesostyle and the well marked paraconule and metaconules;
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A153CFC1AE32B" pageId="37" pageNumber="602" type="description">
<paragraph id="8BC0296D4454B109FD2A153CFB95E0B5" blockId="37.[675,1212,1045,1572]" pageId="37" pageNumber="602">
<taxonomicName id="4C7F52EE4454B109FD35153CFC35E064" authority="Dehm, 1950" authorityName="Dehm" authorityYear="1950" box="[698,948,1363,1387]" class="Mammalia" family="Aplodontidae" genus="Paracitellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4454B109FD35153CFCA6E064" box="[698,807,1363,1387]" italics="true" pageId="37" pageNumber="602">Paracitellus</emphasis>
Dehm, 1950
</taxonomicName>
, in the connection of the paraloph and metaloph to the protocone, and in the absence of labial connection between parastyle-paraconemesostyle-metacone and posteroloph;
</paragraph>
<paragraph id="8BC0296D4454B109FD2A15D2FC28E0E0" blockId="37.[675,1212,1045,1572]" pageId="37" pageNumber="602">
<taxonomicName id="4C7F52EE4454B109FD3915D2FC44E0DB" authority="Kretzoi, 1962" authorityName="Kretzoi" authorityYear="1962" box="[694,965,1468,1493]" class="Mammalia" family="Sciuridae" genus="Miopetaurista" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="37" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4454B109FD3915D2FCB8E0DB" box="[694,825,1469,1492]" italics="true" pageId="37" pageNumber="602">Miopetaurista</emphasis>
Kretzoi, 1962
</taxonomicName>
, or
<taxonomicName id="4C7F52EE4454B109FC6815D2FB34E0DA" authority="Mein, 1970" authorityName="Mein" authorityYear="1970" box="[999,1205,1469,1493]" class="Magnoliopsida" family="Oleaceae" genus="Forsythia" kingdom="Plantae" order="Lamiales" pageId="37" pageNumber="602" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BF57F4454B109FC6815D2FBBDE0DA" box="[999,1084,1469,1493]" italics="true" pageId="37" pageNumber="602">Forsythia</emphasis>
Mein, 1970
</taxonomicName>
, in its non-crescentic cusps;
</paragraph>
<paragraph id="8BC0296D4454B109FD2A159DFC1AE32B" blockId="37.[675,1212,1045,1572]" pageId="37" pageNumber="602">
<taxonomicName id="4C7F52EE4454B109FD37159EFCFCE305" authority="Mein, 1970" authorityName="Mein" authorityYear="1970" box="[696,893,1521,1546]" class="Magnoliopsida" family="Melastomataceae" genus="Blackia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="37" pageNumber="602" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BF57F4454B109FD37159EFD7FE306" box="[696,766,1521,1545]" italics="true" pageId="37" pageNumber="602">Blackia</emphasis>
Mein, 1970
</taxonomicName>
, in the presence of conules, and the less wrinkled enamel.
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64454B109FD2A1653FB3BE3AC" pageId="37" pageNumber="602" type="materials_examined">
<paragraph id="8BC0296D4454B109FD2A1653FB3BE3AC" blockId="37.[677,1211,1595,1699]" pageId="37" pageNumber="602">
MATERIAL
<collectionCode id="ED6EB1A84454B109FC9F162EFCBBE35D" box="[784,826,1601,1618]" pageId="37" pageNumber="602">AND</collectionCode>
MEASUREMENTS (in mm). — A single isolated right upper molar has been collected at Saint- Privat-des-Vieux.
<collectionCode id="ED6EB1A84454B109FCC7161EFCF5E386" box="[840,884,1649,1673]" pageId="37" pageNumber="602">SPV</collectionCode>
190 Mṙ1: L × w
<emphasis id="B90BF57F4454B109FB8A161EFB92E387" box="[1029,1043,1649,1672]" italics="true" pageId="37" pageNumber="602">=</emphasis>
3.4 × 4.02, protocone height
<emphasis id="B90BF57F4454B109FC9B16E3FCA3E3AC" box="[788,802,1676,1699]" italics="true" pageId="37" pageNumber="602">=</emphasis>
2.48, paracone and metacone height
<emphasis id="B90BF57F4454B109FBF816E3FB04E3AC" box="[1143,1157,1676,1699]" italics="true" pageId="37" pageNumber="602">=</emphasis>
1.95.
</paragraph>
</subSubSection>
<caption id="DF0079E54457B10AFFFE138AFEE4E15E" ID-DOI="http://doi.org/10.5281/zenodo.4538657" ID-Zenodo-Dep="4538657" httpUri="https://zenodo.org/record/4538657/files/figure.png" pageId="38" pageNumber="603" startId="38.[113,124,997,1015]" targetBox="[113,1207,222,957]" targetPageId="38">
<paragraph id="8BC0296D4457B10AFFFE138AFEE4E15E" blockId="38.[113,1211,996,1105]" pageId="38" pageNumber="603">
FIG. 14.
<emphasis id="B90BF57F4457B10AFF3E138AFF45E6F8" box="[177,196,997,1015]" italics="true" pageId="38" pageNumber="603"></emphasis>
<taxonomicName id="4C7F52EE4457B10AFF48138AFE52E6F8" authority="Schlosser, 1884" authorityName="Schlosser" authorityYear="1884" box="[199,467,996,1015]" class="Mammalia" family="Eomyidae" genus="Eomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="603" phylum="Chordata" rank="species" species="zitteli">
<emphasis id="B90BF57F4457B10AFF48138AFF7EE6F8" box="[199,255,997,1015]" italics="true" pageId="38" pageNumber="603">Eomys</emphasis>
aff.
<emphasis id="B90BF57F4457B10AFEAF138AFECBE6F8" box="[288,330,997,1015]" italics="true" pageId="38" pageNumber="603">zitteli</emphasis>
Schlosser,1884
</taxonomicName>
,from Saint-Privat-des-Vieux (Gard;Lower Chattian,MP 26):
<emphasis id="B90BF57F4457B10AFC4F138AFC4CE6F8" bold="true" box="[960,973,997,1015]" pageId="38" pageNumber="603">A</emphasis>
, left m1-2, SPV184,(drawing reversed):
<emphasis id="B90BF57F4457B10AFF461394FF5EE102" bold="true" box="[201,223,1019,1037]" pageId="38" pageNumber="603">A1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4457B10AFEEC1394FEFAE102" bold="true" box="[355,379,1019,1037]" pageId="38" pageNumber="603">A2</emphasis>
, labial view;
<emphasis id="B90BF57F4457B10AFE691394FE72E102" bold="true" box="[486,499,1019,1037]" pageId="38" pageNumber="603">B</emphasis>
, digested and worn m1-m3, SPV183, occlusal view;
<emphasis id="B90BF57F4457B10AFC221394FC3AE102" bold="true" box="[941,955,1019,1037]" pageId="38" pageNumber="603">C</emphasis>
, left P4, SPV180:
<emphasis id="B90BF57F4457B10AFBDD1394FBEBE102" bold="true" box="[1106,1130,1019,1037]" pageId="38" pageNumber="603">C1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4457B10AFF2E147DFF38E12B" bold="true" box="[161,185,1042,1060]" pageId="38" pageNumber="603">C2</emphasis>
, lingual view;
<emphasis id="B90BF57F4457B10AFEBD147DFEC1E12B" bold="true" box="[306,320,1042,1060]" pageId="38" pageNumber="603">D</emphasis>
, left M1-2, SPV181:
<emphasis id="B90BF57F4457B10AFE7E147DFD88E12B" bold="true" box="[497,521,1042,1060]" pageId="38" pageNumber="603">D1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4457B10AFD1F147DFD29E12B" bold="true" box="[656,680,1042,1060]" pageId="38" pageNumber="603">D2</emphasis>
, lingual view;
<emphasis id="B90BF57F4457B10AFCAE147DFCACE12B" bold="true" box="[801,813,1042,1060]" pageId="38" pageNumber="603">E</emphasis>
,
<emphasis id="B90BF57F4457B10AFCB6147DFCC5E12B" bold="true" box="[825,836,1042,1060]" pageId="38" pageNumber="603">F</emphasis>
,
<taxonomicName id="4C7F52EE4457B10AFCC0147DFB36E12B" authority="Comte &amp; Vianey-Liaud, 1987" authorityName="Comte &amp; Vianey-Liaud" authorityYear="1987" box="[847,1207,1042,1060]" class="Mammalia" family="Eomyidae" genus="Eomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="603" phylum="Chordata" rank="species" species="minor">
<emphasis id="B90BF57F4457B10AFCC0147DFC3DE12B" box="[847,956,1042,1060]" italics="true" pageId="38" pageNumber="603">Eomys minor</emphasis>
Comte &amp; Vianey-Liaud, 1987
</taxonomicName>
, from Mas-de-Pauffié (Lot; Lower Chattian, MP 26):
<emphasis id="B90BF57F4457B10AFD951446FDA7E134" bold="true" box="[538,550,1065,1083]" pageId="38" pageNumber="603">E</emphasis>
, left M2, MPF215:
<emphasis id="B90BF57F4457B10AFD4B1446FD5AE134" bold="true" box="[708,731,1065,1083]" pageId="38" pageNumber="603">E1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4457B10AFCD21446FCF5E134" bold="true" box="[861,884,1065,1083]" pageId="38" pageNumber="603">E2</emphasis>
, lingual view;
<emphasis id="B90BF57F4457B10AFC661446FC75E134" bold="true" box="[1001,1012,1065,1083]" pageId="38" pageNumber="603">F</emphasis>
, right M3, MPF216, oc- clusal view. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C3657AE64457B10AFFFE14E0FCEDE32B" pageId="38" pageNumber="603" type="description">
<paragraph id="8BC0296D4457B10AFFFE14E0FE83E1A8" blockId="38.[111,648,1167,1700]" box="[113,258,1167,1193]" pageId="38" pageNumber="603">DESCRIPTION</paragraph>
<paragraph id="8BC0296D4457B10AFFFE14C1FBE3E089" blockId="38.[111,648,1167,1700]" lastBlockId="38.[675,1212,1166,1572]" pageId="38" pageNumber="603">Ŋis unworn molar has well-individualized cusps. Ŋe anterior cingulum runs from the base of the protocone to the anterior flank of the paracone which it is connected by a low ridge.It has a bulbous parastyle. Ŋe hypocone is small, but well protrud - ing at the lingual end of the posterior cingulum. Ŋe hypocone is connected to the distal end of the protocone through a low endoloph. Ŋe metacone is a little smaller than the paracone. Ŋe mesostyle is also bulbous, developed in the middle of the valley between the two labial cusps. It extends lingually by a short mesoloph. Ŋe protoloph is connected transversely to the anterior arm of the protocone. It displays one well distinct protoconule, and a weak other one, the more pronounced being the most labial. Ŋe metaloph is connected to the rear of the protocone in front of the endoloph anterior end. It is slightly oblique relative to the protoloph. It has two metaconules, wider than the protoconules. Ŋe labial metaconule is the most marked. Both lophs with their conules are lower than the main cones and styles, which are prominent. A short and low spur descends from the top of the protocone labially in the median valley of the tooth.</paragraph>
<paragraph id="8BC0296D4457B10AFD3415E3FCEDE32B" blockId="38.[675,1212,1166,1572]" pageId="38" pageNumber="603">Ŋe entire crown is relatively high, the lingual flank of the protocone + hypocone being the highest. Enamel is slightly crenulated on the surface of the valleys and transverse lophes, but without clear-cut crestules.</paragraph>
</subSubSection>
<subSubSection id="C3657AE64457B10BFD2A1625FBC7E6E5" lastPageId="39" lastPageNumber="604" pageId="38" pageNumber="603" type="discussion">
<paragraph id="8BC0296D4457B10AFD2A1625FCA7E36D" blockId="38.[677,1210,1610,1700]" box="[677,806,1610,1636]" pageId="38" pageNumber="603">DISCUSSION</paragraph>
<paragraph id="8BC0296D4457B10AFD2A1605FB3BE3AB" blockId="38.[677,1210,1610,1700]" pageId="38" pageNumber="603">Even if the material is restricted to one tooth, we decided to describe a new taxon because it is easy</paragraph>
<paragraph id="8BC0296D4456B10BFFFE10B7FE55E4BF" blockId="39.[111,648,216,1700]" pageId="39" pageNumber="604">
to distinguish this form from the other large Oligocene and lower Miocene squirrels. One lower m3 of close size (L
<emphasis id="B90BF57F4456B10BFE821177FE9CE43E" box="[269,285,280,305]" italics="true" pageId="39" pageNumber="604">=</emphasis>
<quantity id="4C8784884456B10BFEAD1178FE09E43D" box="[290,392,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="4.07" pageId="39" pageNumber="604" unit="mm" value="4.07">4.07 mm</quantity>
, l
<emphasis id="B90BF57F4456B10BFE111177FE2FE43E" box="[414,430,280,305]" italics="true" pageId="39" pageNumber="604">=</emphasis>
<quantity id="4C8784884456B10BFE3C1178FD97E43D" box="[435,534,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="3.41" pageId="39" pageNumber="604" unit="mm" value="3.41">3.41 mm</quantity>
) is known from Pech Desse (Quercy, MP 28a), but its crown is clearly lower (H of metaconid
<emphasis id="B90BF57F4456B10BFE201138FE3EE47F" box="[431,447,343,368]" italics="true" pageId="39" pageNumber="604">=</emphasis>
<quantity id="4C8784884456B10BFE4C1138FDA6E47E" box="[451,551,343,369]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="39" pageNumber="604" unit="mm" value="1.79">1.79 mm</quantity>
, strongly overhanging the other cuspids and the low talonid basin) and the cuspids less acute.
</paragraph>
<paragraph id="8BC0296D4456B10BFF0711DAFD07E684" blockId="39.[111,648,216,1700]" pageId="39" pageNumber="604">
Another large sciurid is summarily reported as
<emphasis id="B90BF57F4456B10BFFFC11B9FEB0E4E0" box="[115,305,470,496]" italics="true" pageId="39" pageNumber="604">
<taxonomicName id="4C7F52EE4456B10BFFF311B9FF49E4FF" box="[124,200,470,496]" class="Mammalia" family="Sciuridae" genus="Sciurus" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus">Sciurus</taxonomicName>
<taxonomicName id="4C7F52EE4456B10BFF5A11B9FEB0E4E0" baseAuthorityName="Freudenberg" baseAuthorityYear="1941" box="[213,305,470,495]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="species" species="giganteus">giganteus</taxonomicName>
</emphasis>
in
<bibRefCitation id="EFEE549C4456B10BFEDF11BAFDA3E4E0" author="FREUDENBERG H." box="[336,546,469,495]" pageId="39" pageNumber="604" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg (1941)</bibRefCitation>
and Werner (1994) from the lower Miocene of Eckingen (Southern
<collectingCountry id="F36869FD4456B10BFF6C127AFECDE720" box="[227,332,533,559]" name="Germany" pageId="39" pageNumber="604">Germany</collectingCountry>
). Only
<quantity id="4C8784884456B10BFE17127AFE45E720" box="[408,452,533,559]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="604" unit="m" value="2.0">2 m</quantity>
3 (4.20 × 4.06 and 4.09 × 4.00) and 1 M3 (3.53 × 3.74) are known.Ŋis species shares wrinkled enamel with
<taxonomicName id="4C7F52EE4456B10BFE51123BFF7FE781" authority="Vianey-Liaud" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="species" species="bernardi" status="n. gen., n. sp.">
<emphasis id="B90BF57F4456B10BFE51123BFD05E761" box="[478,644,596,622]" italics="true" pageId="39" pageNumber="604">Comtia bernardi</emphasis>
Vianey-Liaud
</taxonomicName>
,
<taxonomicNameLabel id="A23848044456B10BFE89121BFE0DE781" box="[262,396,628,654]" pageId="39" pageNumber="604" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Ŋe wrinkles seem more numerous in the former. It is difficult to compare the M3 of Freudenberg with our M1-2, mainly the height of the protocone and other cusps, the shape and number of buccal cusps and their mesio-distal connections, the occurrence of conules.Ŋis species is tentatively included in the genus
<taxonomicName id="4C7F52EE4456B10BFE68135DFDB5E643" authority="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre, 2014" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[487,564,818,844]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B90BF57F4456B10BFE68135DFDB5E643" box="[487,564,818,844]" italics="true" pageId="39" pageNumber="604">Comtia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23848044456B10BFDB2135DFD06E643" box="[573,647,818,844]" pageId="39" pageNumber="604" rank="genus">n. gen.</taxonomicNameLabel>
We keep provisionaly the two species:
<taxonomicName id="4C7F52EE4456B10BFD8D133EFDD8E664" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" box="[514,601,849,875]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="species" species="bernardi">
<emphasis id="B90BF57F4456B10BFD8D133EFDD8E664" box="[514,601,849,875]" italics="true" pageId="39" pageNumber="604">bernardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4456B10BFFFE131DFF4BE684" baseAuthorityName="Freudenberg" baseAuthorityYear="1941" box="[113,202,882,907]" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B90BF57F4456B10BFFFE131DFF4BE684" box="[113,202,882,907]" italics="true" pageId="39" pageNumber="604">giganteus</emphasis>
</taxonomicName>
, waiting for more material and information.
</paragraph>
<paragraph id="8BC0296D4456B10BFF0713FEFDAEE146" blockId="39.[111,648,216,1700]" pageId="39" pageNumber="604">
A slightly smaller upper teeth is described as ṙ
<taxonomicName id="4C7F52EE4456B10BFFF413DEFF45E6C4" authorityName="J.E.Gray" authorityYear="1867" box="[123,196,945,971]" class="Mammalia" family="Sciuridae" genus="Ratufa" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4456B10BFFF413DEFF45E6C4" box="[123,196,945,971]" italics="true" pageId="39" pageNumber="604">Ratufa</emphasis>
</taxonomicName>
sp. from Ulm-Westtangente by Werner (1994: 171). It differs from the M1-2 from Saint- Privat-des-Vieux in its clearly lower crown, its less prominent cusps, especially the hypocone, the absence of metaconules, its smooth enamel.
</paragraph>
<paragraph id="8BC0296D4456B10BFF071420FEEDE0A9" blockId="39.[111,648,216,1700]" pageId="39" pageNumber="604">
Some teeth of a large
<taxonomicName id="4C7F52EE4456B10BFEEF1420FE41E166" box="[352,448,1103,1129]" class="Mammalia" family="Sciuridae" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="family">Sciuridae</taxonomicName>
from an Oligocene Turkish locality (Kocayarmalar) have been attributed to the flying squirrels
<taxonomicName id="4C7F52EE4456B10BFEEF14E1FD07E1A6" authority="Brandt, 1855" authorityName="Brandt" authorityYear="1855" box="[352,646,1166,1193]" class="Mammalia" family="Sciuridae" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="subFamily" subFamily="Petauristinae">Petauristinae Brandt, 1855</taxonomicName>
and with doubt to
<taxonomicName id="4C7F52EE4456B10BFECF14C0FE50E1C7" authorityName="Kretzoi" authorityYear="1962" box="[320,465,1199,1224]" class="Mammalia" family="Sciuridae" genus="Miopetaurista" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4456B10BFECF14C0FE50E1C7" box="[320,465,1199,1224]" italics="true" pageId="39" pageNumber="604">Miopetaurista</emphasis>
</taxonomicName>
, by de
<bibRefCitation id="EFEE549C4456B10BFDAD14C1FE80E1E7" author="BRUIJN H. DE &amp; UNAY E." pageId="39" pageNumber="604" pagination="139 - 146" refId="ref31860" refString="BRUIJN H. DE &amp; UNAY E. 1989. - Petauristinae (Mammalia, Rodentia) from the Oligocene of Spain, Belgium and Turkisch Nrace. Natural History Museum Los Angeles County, Science Series 33: 139 - 146." type="book chapter" year="1989">Bruijn &amp; Ünay (1989)</bibRefCitation>
. Ŋorington
<emphasis id="B90BF57F4456B10BFE1C14A1FE41E1E8" box="[403,448,1229,1255]" italics="true" pageId="39" pageNumber="604">et al</emphasis>
. (2005) discussed the difficulties to identify fossil flying squirrels on the basis of dental morphology alone, considering many cases of convergent evolution. Ŋerefore, we assign the species of Saint-Privat-des-Vieux, represented by a single tooth, to the family
<taxonomicName id="4C7F52EE4456B10BFDAC1503FD06E089" box="[547,647,1388,1414]" class="Mammalia" family="Sciuridae" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="family">Sciuridae</taxonomicName>
without further details.
</paragraph>
<paragraph id="8BC0296D4456B10BFF0715C4FC2AE720" blockId="39.[111,648,216,1700]" lastBlockId="39.[675,1212,216,781]" pageId="39" pageNumber="604">
Slightly smaller than the one of Saint-Privat-des- Vieux, the teeth from Kocayarmalar exhibit some similarities to it, such as the central mesostyle, separated both from paracone and metacone, the lingual mesoloph indicated by an outgrowth in the central basin (here drawn lingually by a spur from the protocone and labially by a marked ridge from the mesostyle) and by the low hypocone. Ŋe strong wear on teeth from Kocayarmalar makes them difficult to compare. However, some significant differences can be pointed, highlighting the strong metaconule, while the labial swellings of the metaloph are lower, in contrast to what occurs at Saint-Privat-des-Vieux. Despite the wear, accessory ridges from the mesostyle and metacone are well marked and the enamel surface is smooth (de
<bibRefCitation id="EFEE549C4456B10BFD2B11BAFC12E4E0" author="BRUIJN H. DE &amp; UNAY E." box="[676,915,469,499]" pageId="39" pageNumber="604" pagination="139 - 146" refId="ref31860" refString="BRUIJN H. DE &amp; UNAY E. 1989. - Petauristinae (Mammalia, Rodentia) from the Oligocene of Spain, Belgium and Turkisch Nrace. Natural History Museum Los Angeles County, Science Series 33: 139 - 146." type="book chapter" year="1989">Bruijn &amp; Ünay 1989</bibRefCitation>
: figs 10, 11). Ŋere are no particular indications on the height of the crown for the Turkish material.
</paragraph>
<paragraph id="8BC0296D4456B10BFD34125BFBBFE603" blockId="39.[675,1212,216,781]" pageId="39" pageNumber="604">
Otherwise, being of the same size as the large Miocene
<taxonomicName id="4C7F52EE4456B10BFC9F123AFC24E761" authorityName="Kretzoi" authorityYear="1962" box="[784,933,597,622]" class="Mammalia" family="Sciuridae" genus="Miopetaurista" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4456B10BFC9F123AFC24E761" box="[784,933,597,622]" italics="true" pageId="39" pageNumber="604">Miopetaurista</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4456B10BFC6E123BFBD7E761" authorityName="Daxner-Hock &amp; Mein" authorityYear="1975" box="[993,1110,596,622]" class="Mammalia" family="Sciuridae" genus="Albanensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4456B10BFC6E123BFBD7E761" box="[993,1110,596,622]" italics="true" pageId="39" pageNumber="604">Albanensia</emphasis>
</taxonomicName>
Daxner- Höck &amp; Mein, 1975, the tooth from Saint-Privatdes-Vieux differs from them by a bunodonty more pronounced, the hypocone shorter, the protoloph and metaloph less convergent, and the crenulations of enamel seemingly more attenuated.
</paragraph>
<paragraph id="8BC0296D4456B10BFD2A135DFC43E645" blockId="39.[675,1211,818,1003]" box="[677,962,818,844]" pageId="39" pageNumber="604">NOMENCLATURAL REMARK</paragraph>
<paragraph id="8BC0296D4456B10BFD2A133EFBC7E6E5" blockId="39.[675,1211,818,1003]" pageId="39" pageNumber="604">
Ŋis new species must be referred to “
<taxonomicName id="4C7F52EE4456B10BFB98133DFCFBE684" authority="Vianey-Liaud, 2014" authorityName="Vianey-Liaud &amp; Comte &amp; Marandat &amp; Peigné &amp; Rage &amp; Sudre" authorityYear="2014" class="Mammalia" family="Sciuridae" genus="Comtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="39" pageNumber="604" phylum="Chordata" rank="species" species="bernardi">
<emphasis id="B90BF57F4456B10BFB98133DFB38E664" box="[1047,1209,849,876]" italics="true" pageId="39" pageNumber="604">Comtia bernardi</emphasis>
Vianey-Liaud, 2014
</taxonomicName>
”, following the article 50.1 and the “recommendation 50A concerning multiple authors” of the International Code of Zoological Nomenclature (ICZN 1999: 52, 182).
</paragraph>
</subSubSection>
</treatment>
</document>