911 lines
141 KiB
XML
911 lines
141 KiB
XML
<document id="DAE2FCDB01A6C83F47BF53825D57B3D3" ID-CLB-Dataset="29954" ID-DOI="10.11646/zootaxa.4429.2.4" ID-GBIF-Dataset="309f28f9-2ce7-43c1-921f-5eefb09bffc7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1283981" ID-ZooBank="52FE0737-5A32-4587-89F4-496669975F42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528274093740" checkinUser="plazi" docAuthor="Kosterin, Oleg E. & Kompier, Tom" docDate="2018" docId="03D687A99801FFA9FF7EFC7AFE56FF10" docLanguage="en" docName="zootaxa.4429.2.4.pdf" docOrigin="Zootaxa 4429 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphicnemis valentini Kosterin & Kompier 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="291" masterDocId="FFEFFFD19800FFA2FFE9FFD5FFB1FFC5" masterDocTitle="Amphicnemis valentini sp. nov. from the Cardamom ecoregion in Cambodia and Vietnam (Odonata: Coenagrionidae)" masterLastPageNumber="294" masterPageNumber="281" pageNumber="282" updateTime="1698547725694" updateUser="ExternalLinkService">
|
||
<mods:mods id="638AC04797F9630B82941301B90BAA69" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EF63CB8779B033F5088CFEF42154DD70">
|
||
<mods:title id="C2C4577AE6C78DB476C3594C382059CB">Amphicnemis valentini sp. nov. from the Cardamom ecoregion in Cambodia and Vietnam (Odonata: Coenagrionidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5085A688388971ED3FEFE108E2F7DC3F" type="personal">
|
||
<mods:role id="C4FC7567B9F9DB0E22AE64FBF7B2DEC7">
|
||
<mods:roleTerm id="674687105B7BB503BD12929C1DA1EE59">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C0841333326A1FFD3DB29C75E5722562">Kosterin, Oleg E.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0400A8E1529B0CA931C081D74B801F94" type="personal">
|
||
<mods:role id="E64FE940950BFDAA16FD77F3F156FBD8">
|
||
<mods:roleTerm id="ECF058674C23BE25A78A32815C90096C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="430764DABB6329AACCE6908972638D8E">Kompier, Tom</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4B1FB02C6559B5DFC6A080023874A735">text</mods:typeOfResource>
|
||
<mods:relatedItem id="791E95BFE8284AA55884C1549CD1B6BF" type="host">
|
||
<mods:titleInfo id="7AD3E6B92E1619A7C35B91FAA823D78E">
|
||
<mods:title id="58CFD48F2CE39285E2712BC40681E9F8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="04650DD34BD345090782748444696419">
|
||
<mods:date id="BB10231CCDB57F2615C7301876CCA04F">2018</mods:date>
|
||
<mods:detail id="C29164274956B89CEC398522463CA87D" type="pubDate">
|
||
<mods:number id="448087A750236A8E664CAAEE7E9C8D63">2018-06-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="7B399A4BDFEAC55F19AE253D55E5319F" type="volume">
|
||
<mods:number id="F541E3F421FED3A57ED6090695E3101F">4429</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6F9FAC2F4731FD0E6170635EB5FCEE2B" type="issue">
|
||
<mods:number id="AAC90951055E88734CE1021EBB60993F">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A90D2FEF5886B08928848BF6673AC95C" unit="page">
|
||
<mods:start id="F8AA0A72AEAD1AE25E570BF7D3BB009B">281</mods:start>
|
||
<mods:end id="06F63B0792F90531B629D2DB221DC909">294</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="FD490C3B41CD49018694D67E44F10238">journal article</mods:classification>
|
||
<mods:identifier id="538A3EEB4CD56A53FC4C8660435F07F4" type="CLB-Dataset">29954</mods:identifier>
|
||
<mods:identifier id="178E166B9B4841D2FF9062178B1B5C16" type="DOI">10.11646/zootaxa.4429.2.4</mods:identifier>
|
||
<mods:identifier id="E3BA7E049FC47794AEBB7D110847255F" type="GBIF-Dataset">309f28f9-2ce7-43c1-921f-5eefb09bffc7</mods:identifier>
|
||
<mods:identifier id="A281A52C929FEF947C4F810DC38045CD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="87B8FA15DC817D5EBDFDAF1AB7C602E6" type="Zenodo-Dep">1283981</mods:identifier>
|
||
<mods:identifier id="1BE2FC83A2A5B096E172D5266B063ADF" type="ZooBank">52FE0737-5A32-4587-89F4-496669975F42</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D687A99801FFA9FF7EFC7AFE56FF10" ID-DOI="http://doi.org/10.5281/zenodo.5974170" ID-GBIF-Taxon="145907993" ID-Zenodo-Dep="5974170" LSID="urn:lsid:plazi:treatment:03D687A99801FFA9FF7EFC7AFE56FF10" httpUri="http://treatment.plazi.org/id/03D687A99801FFA9FF7EFC7AFE56FF10" lastPageId="11" lastPageNumber="291" pageId="1" pageNumber="282">
|
||
<subSubSection id="C36565349801FFA3FF7EFC7AFEFEFC2E" pageId="1" pageNumber="282" type="nomenclature">
|
||
<paragraph id="8BC036BF9801FFA3FF7EFC7AFDABFC0F" blockId="1.[151,538,943,1003]" box="[151,538,943,970]" pageId="1" pageNumber="282">
|
||
<heading id="D08881D39801FFA3FF7EFC7AFDABFC0F" bold="true" box="[151,538,943,970]" fontSize="11" level="1" pageId="1" pageNumber="282" reason="1">
|
||
<emphasis id="B90BEAAD9801FFA3FF7EFC7AFDABFC0F" bold="true" box="[151,538,943,970]" pageId="1" pageNumber="282">
|
||
<emphasis id="B90BEAAD9801FFA3FF7EFC7AFE04FC0C" bold="true" box="[151,437,943,969]" italics="true" pageId="1" pageNumber="282">
|
||
<taxonomicName id="4C7F4D3C9801FFA3FF7EFC7AFE1EFC0C" ID-CoL="9WZYH" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[151,431,943,969]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">Amphicnemis valentini</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A23857D69801FFA3FE55FC65FDABFC0F" box="[444,538,944,970]" pageId="1" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9801FFA3FF7EFC07FEFEFC2E" blockId="1.[151,538,943,1003]" box="[151,335,978,1003]" pageId="1" pageNumber="282">
|
||
(
|
||
<figureCitation id="13442A3A9801FFA3FF76FC07FEB5FC2E" box="[159,260,978,1003]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1170,1192]" captionStartId-1="3.[151,250,1350,1372]" captionStartId-2="4.[151,250,1325,1347]" captionStartId-3="6.[151,250,1941,1963]" captionStartId-4="7.[151,250,1973,1995]" captionStartId-5="8.[151,250,1878,1900]" captionTargetBox-0="[151,1436,187,1155]" captionTargetBox-1="[151,1436,193,1329]" captionTargetBox-2="[151,1436,517,1304]" captionTargetBox-3="[151,1436,193,1880]" captionTargetBox-4="[275,1299,193,1916]" captionTargetBox-5="[151,1436,193,1827]" captionTargetId-0="figure@2.[151,1436,187,1155]" captionTargetId-1="figure@3.[151,1436,193,1329]" captionTargetId-2="figure@4.[151,1436,517,1304]" captionTargetId-3="figure@6.[151,1436,193,1944]" captionTargetId-4="figure@7.[275,1299,193,1916]" captionTargetId-5="figure@8.[151,1436,193,1856]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Males of Amphicnemis valentini, sp. nov. in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K." captionText-1="FIGURE 2. Immature (a,c) and mature (b,d) females of Amphicnemis valentini, sp. nov. in nature, Vietnam, Kiên Giang Province, Phú Quốc Island, 19 (a-b) and 20 (c-d) viii 2016, photos by T.K." captionText-2="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." captionText-3="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." captionText-4="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." captionText-5="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri-0="https://zenodo.org/record/1283983/files/figure.png" httpUri-1="https://zenodo.org/record/1283985/files/figure.png" httpUri-2="https://zenodo.org/record/1283987/files/figure.png" httpUri-3="https://zenodo.org/record/1283989/files/figure.png" httpUri-4="https://zenodo.org/record/1283991/files/figure.png" httpUri-5="https://zenodo.org/record/1283993/files/figure.png" pageId="1" pageNumber="282">Figs 1–6</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9801FFA3FEE6FC06FEF6FC2E" box="[271,327,979,1003]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="1" pageNumber="282">7e–g</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565349801FFA3FF7EFBCFFBC7FBF5" box="[151,1142,1050,1072]" pageId="1" pageNumber="282" type="description">
|
||
<paragraph id="8BC036BF9801FFA3FF7EFBCFFBC7FBF5" blockId="1.[151,1142,1050,1072]" box="[151,1142,1050,1072]" pageId="1" pageNumber="282">
|
||
<treatmentCitationGroup id="AB6F11919801FFA3FF7EFBCFFBC7FBF5" box="[151,1142,1050,1072]" pageId="1" pageNumber="282">
|
||
<taxonomicName id="4C7F4D3C9801FFA3FF7EFBCFFE4CFBF5" ID-CoL="CZSV" authority="Kruger, 1898" authorityName="Kruger" authorityYear="1898" box="[151,509,1050,1072]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9801FFA3FF7EFBCFFEC3FBF5" box="[151,370,1050,1072]" italics="true" pageId="1" pageNumber="282">Amphicnemis gracilis</emphasis>
|
||
Krüger, 1898
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0ADE10AE9801FFA3FDF1FBCFFD60FBF5" author="" box="[536,721,1050,1072]" page="55" pageId="1" pageNumber="282">
|
||
<bibRefCitation id="EFEE4B4E9801FFA3FDF1FBCFFD1DFBF5" box="[536,684,1050,1072]" pageId="1" pageNumber="282" refId="ref6396" refString="Do, M. C., Bui, H. M. & Nguen, V. K. (2011) Dragonflies of Phu Quoc Island, South Vietnam. Agrion, 15 (2), 54 - 57." type="journal article">
|
||
Do
|
||
<emphasis id="B90BEAAD9801FFA3FDD7FBCEFDC0FBF5" box="[574,625,1050,1072]" italics="true" pageId="1" pageNumber="282">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
: 55
|
||
</treatmentCitation>
|
||
, Photo 4—Report from Phú Quốc
|
||
<collectingCountry id="F368762F9801FFA3FBD8FBCFFBC3FBF5" box="[1073,1138,1050,1072]" name="Iceland" pageId="1" pageNumber="282">Island</collectingCountry>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565349801FFA3FF7EFB88FEFFFA1B" pageId="1" pageNumber="282" type="materials_examined">
|
||
<paragraph id="8BC036BF9801FFA3FF7EFB88FC08FAEF" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="282">
|
||
<materialsCitation id="3B173CE29801FFA3FF7EFB88FCDEFB7B" ID-GBIF-Occurrence="1848506035" collectingDate="2017-03-05" collectionCode="RMNH" collectorName="Oleg E. Kosterin & Netherlands" country="Cambodia" elevation="16" latitude="10.52258" location="Ream Peninsula" longLatPrecision="1" longitude="103.69556" pageId="1" pageNumber="282" specimenCount="1" specimenCount-male="1" stateProvince="Preah Sihanouk Province" typeStatus="holotype">
|
||
<emphasis id="B90BEAAD9801FFA3FF7EFB88FE72FBB3" bold="true" box="[151,451,1117,1142]" pageId="1" pageNumber="282">
|
||
<typeStatus id="54C4881D9801FFA3FF7EFB88FF63FBB3" box="[151,210,1117,1142]" pageId="1" pageNumber="282">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54C4881D9801FFA3FEA4FB88FE0FFBB3" box="[333,446,1117,1142]" pageId="1" pageNumber="282" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D79FD369801FFA3FE25FB8BFE50FBB0" box="[460,481,1118,1141]" pageId="1" pageNumber="282" type="male">♂</specimenCount>
|
||
(
|
||
<figureCitation id="13442A3A9801FFA3FE18FB88FDF2FBB3" box="[497,579,1117,1142]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="1" pageNumber="282">Fig. 3a</figureCitation>
|
||
),
|
||
<collectingCountry id="F368762F9801FFA3FDB0FB88FD7EFBB3" box="[601,719,1117,1142]" name="Cambodia" pageId="1" pageNumber="282">Cambodia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49BBF85D9801FFA3FD32FB88FC4DFBB3" box="[731,1020,1117,1142]" inferredBySuffix="true" pageId="1" pageNumber="282">Preah Sihanouk Province</collectingRegion>
|
||
,
|
||
<location id="8EA060649801FFA3FBEEFB8BFB70FBB3" LSID="urn:lsid:plazi:treatment:03D687A99801FFA9FF7EFC7AFE56FF10:8EA060649801FFA3FBEEFB8BFB70FBB3" box="[1031,1217,1117,1142]" country="Cambodia" latitude="10.52258" longLatPrecision="1" longitude="103.69556" name="Ream Peninsula" pageId="1" pageNumber="282" stateProvince="Preah Sihanouk Province">Ream Peninsula</location>
|
||
, lower reaches of a rivulet in lowland evergreen forest,
|
||
<geoCoordinate id="EE4B50789801FFA3FDD6FB54FD76FB5C" box="[575,711,1153,1178]" direction="north" orientation="latitude" pageId="1" pageNumber="282" precision="1" value="10.52258">10.52258 N</geoCoordinate>
|
||
<geoCoordinate id="EE4B50789801FFA3FD3AFB54FCD8FB5C" box="[723,873,1153,1178]" direction="east" orientation="longitude" pageId="1" pageNumber="282" precision="1" value="103.69556">103.69556 E</geoCoordinate>
|
||
,
|
||
<elevation id="0052D18C9801FFA3FC93FB54FC49FB5F" box="[890,1016,1153,1178]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="282" unit="m" value="16.0">
|
||
<quantity id="4C879B5A9801FFA3FC93FB54FC06FB5C" box="[890,951,1153,1178]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="282" unit="m" value="16.0">16 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFC1107F9801FFA3FBE0FB54FB30FB5F" box="[1033,1153,1153,1178]" pageId="1" pageNumber="282" value="2017-03-05">
|
||
<collectingDate id="EF85E9979801FFA3FBE0FB54FB30FB5F" box="[1033,1153,1153,1178]" pageId="1" pageNumber="282" value="2017-03-05">5 iii 2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268A53699801FFA3FB78FB57FB76FB5F" box="[1169,1223,1154,1178]" name="Oleg E. Kosterin" pageId="1" pageNumber="282">O.K.</collectorName>
|
||
leg.; deposited in Naturalis Biodiversity Centre, Leiden, the
|
||
<collectorName id="268A53699801FFA3FD9FFB70FD4CFB7B" box="[630,765,1189,1214]" pageId="1" pageNumber="282">Netherlands</collectorName>
|
||
(
|
||
<collectionCode id="ED6EAE7A9801FFA3FCE5FB73FCD7FB7B" box="[780,870,1190,1214]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="282">RMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B90BEAAD9801FFA3FC90FB70FC4AFB7B" bold="true" box="[889,1019,1189,1214]" pageId="1" pageNumber="282">Paratypes:</emphasis>
|
||
1 ♂ (preserved in alcohol), the same place, 10.5238 N 103.6955 E,
|
||
<quantity id="4C879B5A9801FFA3FE03FB1FFD93FB24" box="[490,546,1226,1250]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="282" unit="m" value="20.0">20 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC1107F9801FFA3FD83FB1FFD69FB27" box="[618,728,1225,1250]" pageId="1" pageNumber="282" value="2017-03-04">4 iii 2017</date>
|
||
, N.V. Priydak leg.; 1 ♂, 1 ♀ (
|
||
<figureCitation id="13442A3A9801FFA3FBC9FB1CFBC3FB27" box="[1056,1138,1225,1250]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="1" pageNumber="282">Fig. 3b</figureCitation>
|
||
), the same place, 10.5226- 5238 N 103.6955 E,
|
||
<quantity id="4C879B5A9801FFA3FE97FB38FE6AFAC0" box="[382,475,1261,1286]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="1" pageNumber="282" unit="m" value="18.0" valueMax="20.0" valueMin="16.0">16-20 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFC1107F9801FFA3FDCAFB38FD21FAC3" box="[547,656,1261,1286]" pageId="1" pageNumber="282" value="2017-03-05">5 iii 2017</date>
|
||
, O.K. leg.;
|
||
<materialsCitation id="3B173CE29801FFA3FCE6FB3BFC08FAEF" ID-GBIF-Occurrence="1848506037" collectingDate="2016-08-20" collectorName="Tom Kompier" country="Vietnam" latitude="10.251" location="Island" longLatPrecision="77" longitude="104.029" municipality="Phu Qu" pageId="1" pageNumber="282" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Kien Giang Province" typeStatus="holotype">
|
||
<specimenCount id="9D79FD369801FFA3FCE6FB3BFCFEFAC0" box="[783,847,1262,1286]" pageId="1" pageNumber="282" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D79FD369801FFA3FCB2FB38FC28FAC0" box="[859,921,1261,1286]" pageId="1" pageNumber="282" type="female">6 ♀♀</specimenCount>
|
||
(5 mature, 1 immature),
|
||
<collectingCountry id="F368762F9801FFA3FB46FB38FAA0FAC3" box="[1199,1297,1261,1286]" name="Vietnam" pageId="1" pageNumber="282">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion id="49BBF85D9801FFA3FAF2FB38FF4FFAEF" country="Vietnam" name="Kien Giang" pageId="1" pageNumber="282">
|
||
Kiên
|
||
<collectingRegion id="49BBF85D9801FFA3FAB1FB38FF4FFAEF" inferredBySuffix="true" pageId="1" pageNumber="282">Giang Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BA4ACC59801FFA3FEE1FAC4FEEFFAEF" box="[264,350,1297,1322]" pageId="1" pageNumber="282">Phú Qu</collectingMunicipality>
|
||
ốc
|
||
<location id="8EA060649801FFA3FE97FAC4FE74FAEF" LSID="urn:lsid:plazi:treatment:03D687A99801FFA9FF7EFC7AFE56FF10:8EA060649801FFA3FE97FAC4FE74FAEF" box="[382,453,1297,1322]" country="Vietnam" latitude="10.251" longLatPrecision="77" longitude="104.029" municipality="Phu Qu" name="Island" pageId="1" pageNumber="282" stateProvince="Kien Giang Province">Island</location>
|
||
,
|
||
<geoCoordinate id="EE4B50789801FFA3FE38FAC4FD88FAEC" box="[465,569,1297,1322]" direction="north" orientation="latitude" pageId="1" pageNumber="282" precision="55" value="10.251">10.251 N</geoCoordinate>
|
||
<geoCoordinate id="EE4B50789801FFA3FDA9FAC7FD04FAEC" box="[576,693,1298,1322]" direction="east" orientation="longitude" pageId="1" pageNumber="282" precision="55" value="104.029">104.029 E</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF85E9979801FFA3FD28FAC7FCFDFAEF" box="[705,844,1297,1322]" pageId="1" pageNumber="282" value="2016-08-20">
|
||
<date id="FFC1107F9801FFA3FD28FAC7FCFDFAEF" box="[705,844,1297,1322]" pageId="1" pageNumber="282" value="2016-08-20">20 viii 2016</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="268A53699801FFA3FCBEFAC7FC39FAEC" box="[855,904,1298,1321]" name="Tom Kompier" pageId="1" pageNumber="282">T.K.</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9801FFA3FF2EFAE0FEFFFA1B" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="282">
|
||
<emphasis id="B90BEAAD9801FFA3FF2EFAE0FC9BFA8B" bold="true" box="[199,810,1333,1358]" pageId="1" pageNumber="282">Additional data: visual and photographic records</emphasis>
|
||
. 1 ♂ (a photo,
|
||
<figureCitation id="13442A3A9801FFA3FC0EFAE0FB8BFA8B" box="[999,1082,1333,1358]" captionStart="FIGURE 1" captionStartId="2.[151,250,1170,1192]" captionTargetBox="[151,1436,187,1155]" captionTargetId="figure@2.[151,1436,187,1155]" captionTargetPageId="2" captionText="FIGURE 1. Males of Amphicnemis valentini, sp. nov. in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K." httpUri="https://zenodo.org/record/1283983/files/figure.png" pageId="1" pageNumber="282">Fig. 1a</figureCitation>
|
||
), the type locality, 10.5238 N 103.6955 E,
|
||
<date id="FFC1107F9801FFA3FEC0FA8FFE2BFAB7" box="[297,410,1369,1394]" pageId="1" pageNumber="282" value="2017-03-04">4 iii 2017</date>
|
||
, O.K. 10 ♂♂, 13♀♀ (photos,
|
||
<figureCitation id="13442A3A9801FFA3FD1DFA8CFCE2FAB7" box="[756,851,1369,1394]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1170,1192]" captionStartId-1="3.[151,250,1350,1372]" captionTargetBox-0="[151,1436,187,1155]" captionTargetBox-1="[151,1436,193,1329]" captionTargetId-0="figure@2.[151,1436,187,1155]" captionTargetId-1="figure@3.[151,1436,193,1329]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Males of Amphicnemis valentini, sp. nov. in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K." captionText-1="FIGURE 2. Immature (a,c) and mature (b,d) females of Amphicnemis valentini, sp. nov. in nature, Vietnam, Kiên Giang Province, Phú Quốc Island, 19 (a-b) and 20 (c-d) viii 2016, photos by T.K." httpUri-0="https://zenodo.org/record/1283983/files/figure.png" httpUri-1="https://zenodo.org/record/1283985/files/figure.png" pageId="1" pageNumber="282">Figs 1-2</figureCitation>
|
||
), Vietnam,
|
||
<materialsCitation id="3B173CE29801FFA3FC30FA8CFC7CFA53" ID-GBIF-Occurrence="1848506036" collectingDate="2016-08-19" collectingDateMax="2016-08-20" collectingDateMin="2016-08-19" collectorName="Tom Kompier" country="Iceland" latitude="10.251" location="Phu Qu" longLatPrecision="77" longitude="104.029" pageId="1" pageNumber="282" specimenCount="1" specimenCount-female="1" stateProvince="Kien Giang Province">
|
||
<collectingRegion id="49BBF85D9801FFA3FC30FA8CFB7DFAB7" box="[985,1228,1369,1394]" country="Vietnam" name="Kien Giang" pageId="1" pageNumber="282">
|
||
Kiên
|
||
<collectingRegion id="49BBF85D9801FFA3FBF1FA8CFB7DFAB7" box="[1048,1228,1369,1394]" inferredBySuffix="true" pageId="1" pageNumber="282">Giang Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<location id="8EA060649801FFA3FB31FA8CFA9EFAB7" LSID="urn:lsid:plazi:treatment:03D687A99801FFA9FF7EFC7AFE56FF10:8EA060649801FFA3FB31FA8CFA9EFAB7" box="[1240,1327,1369,1394]" country="Iceland" latitude="10.251" longLatPrecision="77" longitude="104.029" name="Phu Qu" pageId="1" pageNumber="282" stateProvince="Kien Giang Province">Phú Qu</location>
|
||
ốc
|
||
<collectingCountry id="F368762F9801FFA3FAB8FA8CFA29FAB7" box="[1361,1432,1369,1394]" name="Iceland" pageId="1" pageNumber="282">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE4B50789801FFA3FF7EFAA8FEB4FA50" box="[151,261,1405,1430]" direction="north" orientation="latitude" pageId="1" pageNumber="282" precision="55" value="10.251">10.251 N</geoCoordinate>
|
||
<geoCoordinate id="EE4B50789801FFA3FEFBFAABFE3CFA50" box="[274,397,1406,1430]" direction="east" orientation="longitude" pageId="1" pageNumber="282" precision="55" value="104.029">104.029 E</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF85E9979801FFA3FE76FAABFDEBFA53" box="[415,602,1405,1430]" pageId="1" pageNumber="282" value="2016-08-19" valueMax="2016-08-20" valueMin="2016-08-19">
|
||
<date id="FFC1107F9801FFA3FE76FAABFDEBFA53" box="[415,602,1405,1430]" pageId="1" pageNumber="282" value="2016-08-19" valueMax="2016-08-20" valueMin="2016-08-19">19-20 viii 2016</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="268A53699801FFA3FD82FAABFD2CFA50" box="[619,669,1406,1429]" name="Tom Kompier" pageId="1" pageNumber="282">T.K.</collectorName>
|
||
1 immature
|
||
<specimenCount id="9D79FD369801FFA3FCD5FAABFCE0FA50" box="[828,849,1406,1429]" pageId="1" pageNumber="282" type="female">♀</specimenCount>
|
||
(a photo)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B173CE29801FFA3FC37FAA8FEFFFA1B" ID-GBIF-Occurrence="1848506038" collectingDate="2017-12-09" collectorName="Gerard Chartier." country="Cambodia" latitude="11.5186" location="Tatai Commune" longLatPrecision="7" longitude="103.1233" municipality="Tatai Riverestuary" pageId="1" pageNumber="282" specimenCount="1" stateProvince="Koh Kong Province">
|
||
<collectingCountry id="F368762F9801FFA3FC37FAA8FBE5FA53" box="[990,1108,1405,1430]" name="Cambodia" pageId="1" pageNumber="282">Cambodia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49BBF85D9801FFA3FB8CFAA8FAE5FA53" box="[1125,1364,1405,1430]" country="Cambodia" name="Koh Kong" pageId="1" pageNumber="282">
|
||
Koh
|
||
<collectingRegion id="49BBF85D9801FFA3FB4BFAABFAE5FA53" box="[1186,1364,1405,1430]" inferredBySuffix="true" pageId="1" pageNumber="282">Kong Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<location id="8EA060649801FFA3FA8DFAA8FEBEFA7F" LSID="urn:lsid:plazi:treatment:03D687A99801FFA9FF7EFC7AFE56FF10:8EA060649801FFA3FA8DFAA8FEBEFA7F" country="Cambodia" latitude="11.5186" longLatPrecision="7" longitude="103.1233" municipality="Tatai Riverestuary" name="Tatai Commune" pageId="1" pageNumber="282" stateProvince="Koh Kong Province">Tatai Commune</location>
|
||
, a lower course of an unnamed tributary of the
|
||
<collectingMunicipality id="6BA4ACC59801FFA3FCD8FA74FBB3FA7F" box="[817,1026,1441,1466]" pageId="1" pageNumber="282">Tatai Riverestuary</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE4B50789801FFA3FBE7FA74FB34FA7C" box="[1038,1157,1441,1466]" direction="north" orientation="latitude" pageId="1" pageNumber="282" precision="5" value="11.5186">11.5186 N</geoCoordinate>
|
||
<geoCoordinate id="EE4B50789801FFA3FB67FA77FAA2FA7C" box="[1166,1299,1442,1466]" direction="east" orientation="longitude" pageId="1" pageNumber="282" precision="5" value="103.1233">103.1233 E</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF85E9979801FFA3FAC9FA77FA29FA7F" box="[1312,1432,1441,1466]" pageId="1" pageNumber="282" value="2017-12-09">
|
||
<date id="FFC1107F9801FFA3FAC9FA77FA29FA7F" box="[1312,1432,1441,1466]" pageId="1" pageNumber="282" value="2017-12-09">9 xii 2017</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="268A53699801FFA3FF7EFA10FEFFFA1B" box="[151,334,1477,1502]" pageId="1" pageNumber="282">Gerard Chartier.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565349801FFA3FF2EFA3CFB23F9E3" pageId="1" pageNumber="282" type="etymology">
|
||
<paragraph id="8BC036BF9801FFA3FF2EFA3CFB23F9E3" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="282">
|
||
<emphasis id="B90BEAAD9801FFA3FF2EFA3CFEFFF9C7" bold="true" box="[199,334,1513,1538]" pageId="1" pageNumber="282">Etymology.</emphasis>
|
||
The specific name, a noun in genitive case, is derived from the name of O.K.’s son Valentin, who, at the age of six, accompanied his parents at the
|
||
<typeStatus id="54C4881D9801FFA3FD5CF9DBFD54F9E3" box="[693,741,1550,1574]" pageId="1" pageNumber="282">type</typeStatus>
|
||
locality when they found this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565349801FFA7FF2EF9E4FAEBFAF5" lastPageId="5" lastPageNumber="286" pageId="1" pageNumber="282" type="description">
|
||
<paragraph id="8BC036BF9801FFA3FF2EF9E4FD39F98F" blockId="1.[151,1437,1117,2006]" box="[199,648,1585,1610]" pageId="1" pageNumber="282">
|
||
<emphasis id="B90BEAAD9801FFA3FF2EF9E4FE6CF98F" bold="true" box="[199,477,1585,1610]" pageId="1" pageNumber="282">
|
||
Description of
|
||
<typeStatus id="54C4881D9801FFA3FE9FF9E4FE6CF98F" box="[374,477,1585,1610]" pageId="1" pageNumber="282" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13442A3A9801FFA3FE05F9E4FDF0F98F" box="[492,577,1585,1610]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="1" pageNumber="282">Figs 3a</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9801FFA3FDA4F9E4FDCAF98F" box="[589,635,1585,1610]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1941,1963]" captionStartId-1="7.[151,250,1973,1995]" captionTargetBox-0="[151,1436,193,1880]" captionTargetBox-1="[275,1299,193,1916]" captionTargetId-0="figure@6.[151,1436,193,1944]" captionTargetId-1="figure@7.[275,1299,193,1916]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." captionText-1="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri-0="https://zenodo.org/record/1283989/files/figure.png" httpUri-1="https://zenodo.org/record/1283991/files/figure.png" pageId="1" pageNumber="282">4–5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9801FFA3FF2EF980FA2CF813" blockId="1.[151,1437,1117,2006]" pageId="1" pageNumber="282">
|
||
<emphasis id="B90BEAAD9801FFA3FF2EF980FEB5F9AB" box="[199,260,1621,1646]" italics="true" pageId="1" pageNumber="282">Head</emphasis>
|
||
(
|
||
<figureCitation id="13442A3A9801FFA3FEFCF980FED6F9AB" box="[277,359,1621,1646]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="1" pageNumber="282">Fig. 4e</figureCitation>
|
||
) mostly metallic black, dorsal side with a green shine, face with some yellowish markings as follows: Labium yellowish. Labrum glittering black shading to yellowish on lower one-third. Mandible bases yellowish along sides, brownish in middle (border of colour indistinct). Vertical surface of clypeus saturated yellow below, dark-brown above. The border of these colours, looking like a pair of fused broadly triangular spots, mostly coincides with the wavy suture on the vertical surface of the clypeus, but laterally the yellow colour extends above it. Horizontal surface of clypeus glittering black. Genae dull greenish below, brownish-black above, dull greenish colour extending dorsally along margins of compound eyes and alongside clypeus towards antefrons (
|
||
<figureCitation id="13442A3A9801FFA3FAD0F8F8FA3EF883" box="[1337,1423,1837,1862]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="1" pageNumber="282">Fig. 4e</figureCitation>
|
||
). Ventral half of antefrons dull greenish-yellow (only narrowly so towards middle), dorsal half black with a pair of large dull greenish spots immediately below antenna bases. Antennae blackish-brown, but apex of segment 1 and anterior part of segment 2 yellowish. Occiput with long golden bristles. Compound eyes in life (
|
||
<figureCitation id="13442A3A9801FFA3FB0EF84CFA98F877" box="[1255,1321,1945,1970]" captionStart="FIGURE 1" captionStartId="2.[151,250,1170,1192]" captionTargetBox="[151,1436,187,1155]" captionTargetId="figure@2.[151,1436,187,1155]" captionTargetPageId="2" captionText="FIGURE 1. Males of Amphicnemis valentini, sp. nov. in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K." httpUri="https://zenodo.org/record/1283983/files/figure.png" pageId="1" pageNumber="282">Fig. 1</figureCitation>
|
||
) greenish brown above, gradually turning yellowish green below; darker upper part with a narrow greenish rim along margin.
|
||
</paragraph>
|
||
<caption id="DF0066379802FFA0FF7EFB47FB39FB02" httpUri="https://zenodo.org/record/1283983/files/figure.png" pageId="2" pageNumber="283" startId="2.[151,250,1170,1192]" targetBox="[151,1436,187,1155]" targetPageId="2">
|
||
<paragraph id="8BC036BF9802FFA0FF7EFB47FB39FB02" blockId="2.[151,1436,1170,1223]" pageId="2" pageNumber="283">
|
||
<emphasis id="B90BEAAD9802FFA0FF7EFB47FEA5FB62" bold="true" box="[151,276,1170,1192]" pageId="2" pageNumber="283">FIGURE 1.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C7F4D3C9802FFA0FE95FB47FDD4FB6D" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[380,613,1170,1192]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9802FFA0FE95FB47FDD4FB6D" box="[380,613,1170,1192]" italics="true" pageId="2" pageNumber="283">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69802FFA0FD9AFB47FD75FB62" box="[627,708,1170,1191]" pageId="2" pageNumber="283" rank="species">
|
||
<emphasis id="B90BEAAD9802FFA0FD9AFB47FD75FB62" bold="true" box="[627,708,1170,1191]" pageId="2" pageNumber="283">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036BF9802FFA0FF2EFB3BFA23F8AE" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="283">
|
||
<emphasis id="B90BEAAD9802FFA0FF2EFB3BFEAEFAC2" box="[199,287,1262,1287]" italics="true" pageId="2" pageNumber="283">Thorax.</emphasis>
|
||
Prothorax (
|
||
<figureCitation id="13442A3A9802FFA0FE45FB3BFDACFAC2" box="[428,541,1262,1287]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4c–d</figureCitation>
|
||
) metallic black with variable metallic gloss above, yellowish below (border between the two colours indistinct). Anterior lobe with a rounded margin, brown at sides, black with a green shine at middle, and with large roundish dorsal markings of yellowish colour laterally on either side (
|
||
<figureCitation id="13442A3A9802FFA0FB0BFAE3FA86FA8A" box="[1250,1335,1334,1359]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4c</figureCitation>
|
||
). Medial lobe black with a bluish-green shine on its proximal two-thirds and bronze-green shine on its distal third; posterior lobe black with a strong violet shine (
|
||
<figureCitation id="13442A3A9802FFA0FDA3FAABFD11FA52" box="[586,672,1406,1431]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4d</figureCitation>
|
||
). Posterior lobe produced into a robust and long hook-like process (
|
||
<figureCitation id="13442A3A9802FFA0FF76FA77FF5EFA7E" box="[159,239,1442,1467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4c</figureCitation>
|
||
) generally inclined to the left side (
|
||
<figureCitation id="13442A3A9802FFA0FD95FA77FD7FFA7E" box="[636,718,1442,1467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4d</figureCitation>
|
||
) and evenly curving anteriorly so that its tip is nearly horizontal (
|
||
<figureCitation id="13442A3A9802FFA0FF76FA13FF40FA1A" box="[159,241,1478,1503]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4c</figureCitation>
|
||
), its margins rough. Distance from anterior lobe base to process tip slightly less than median lobe length. Lateral posterior extremities of posterior lobe bluntly angulated (
|
||
<figureCitation id="13442A3A9802FFA0FC90FA3FFC7CF9C6" box="[889,973,1514,1539]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4c</figureCitation>
|
||
). In dorsal view (
|
||
<figureCitation id="13442A3A9802FFA0FB7FFA3FFB5AF9C6" box="[1174,1259,1514,1539]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4d</figureCitation>
|
||
), posterior lobe margins are broadly and evenly concave at sides and more narrowly and pointedly concave at center, so that its outline resembles a bow composed of two horns. Mesostigmal plate easily observed from above (
|
||
<figureCitation id="13442A3A9802FFA0FB06F9E7FAF4F98E" box="[1263,1349,1586,1611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4d</figureCitation>
|
||
), of the typical male coenagrionid form, as composed by a pair of transverse hollowed parts, long triangular with slightly convex margins in outline, brownish with a green shine, and two slightly semicircular anterior ridges at middorsal suture. Synthorax (
|
||
<figureCitation id="13442A3A9802FFA0FE90F94BFE61F972" box="[377,464,1694,1719]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">Fig. 4a</figureCitation>
|
||
) glittering bronze-black (with mixed purple and green shine) dorsally, yellowish ventrally, the transition rather indistinct, with an uneven border and transitioning through brown over the middle of mesinfraepisternum (where black projects unevenly ventrally) through spiracle to ventroposterior corner of metepisternum. Dorsoposterior part of metapleural suture with a blackish-brown darkening connecting through vague brownish spots at posterior part of metepimeron to black dorsum of synthorax; posterior ridge of metepimeron brownish-black. Pleural sclerites brown, some with a greenish shine. Venter of synthorax yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9802FFA0FF2EF8A2FC13F812" blockId="2.[151,1437,1262,2007]" pageId="2" pageNumber="283">
|
||
Legs (
|
||
<figureCitation id="13442A3A9802FFA0FEE7F8A3FED7F84A" box="[270,358,1910,1935]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="2" pageNumber="283">Figs 3a</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9802FFA0FE9DF8A2FE21F84A" box="[372,400,1911,1935]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="2" pageNumber="283">4a</figureCitation>
|
||
) yellowish with blackish brown parts as follows: stripes along extensor surface of femora (not extending to both ends of metafemora), apices of femora, bases of tibia (markedly at protibia), all segment joints of tarsi; spines blackish-brown, claws brown with black apices.
|
||
</paragraph>
|
||
<caption id="DF0066379803FFA1FF7EFA93FC3BFABE" httpUri="https://zenodo.org/record/1283985/files/figure.png" pageId="3" pageNumber="284" startId="3.[151,250,1350,1372]" targetBox="[151,1436,193,1329]" targetPageId="3">
|
||
<paragraph id="8BC036BF9803FFA1FF7EFA93FC3BFABE" blockId="3.[151,1436,1350,1403]" pageId="3" pageNumber="284">
|
||
<emphasis id="B90BEAAD9803FFA1FF7EFA93FEA7FA99" bold="true" box="[151,278,1350,1372]" pageId="3" pageNumber="284">FIGURE 2.</emphasis>
|
||
Immature (a,c) and mature (b,d) females of
|
||
<taxonomicName id="4C7F4D3C9803FFA1FD1CFA93FC50FA99" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[757,993,1350,1372]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9803FFA1FD1CFA93FC50FA99" box="[757,993,1350,1372]" italics="true" pageId="3" pageNumber="284">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69803FFA1FC18FA92FBF4FA99" box="[1009,1093,1351,1372]" pageId="3" pageNumber="284" rank="species">
|
||
<emphasis id="B90BEAAD9803FFA1FC18FA92FBF4FA99" bold="true" box="[1009,1093,1351,1372]" pageId="3" pageNumber="284">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in nature, Vietnam, Kiên Giang Province, Phú Quốc Island, 19 (a-b) and 20 (c-d) viii 2016, photos by T.K.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036BF9803FFA1FF2EFA71FED3F98E" blockId="3.[151,1437,1442,2007]" pageId="3" pageNumber="284">
|
||
<emphasis id="B90BEAAD9803FFA1FF2EFA71FEBBFA7E" box="[199,266,1444,1467]" italics="true" pageId="3" pageNumber="284">Wings</emphasis>
|
||
hyaline; venation blackish-brown; 13 (left) - 14 (right) Px in FW, 11 (left) - 12 (right) in HW. Arculus noticeably distal to Ax2. 1A arising at level of arculus. IR3 and R4 arising distal of subnodus. Pterostigmata (
|
||
<figureCitation id="13442A3A9803FFA1FA86FA13FF05F9C6" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="3" pageNumber="284">Fig. 4b</figureCitation>
|
||
) similar in both wings, brownish-black, with a lighter margin, costal side shorter than anal side, which is slightly shorter than underlying cell; both transversal sides convex, proximal one slanting, distal one perpendicular; ventral margin thickened.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9803FFA1FF2EF983FE59F882" blockId="3.[151,1437,1442,2007]" pageId="3" pageNumber="284">
|
||
<emphasis id="B90BEAAD9803FFA1FF2EF983FE80F9AA" box="[199,305,1622,1647]" italics="true" pageId="3" pageNumber="284">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="13442A3A9803FFA1FEAFF983FE2CF9AA" box="[326,413,1622,1647]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="3" pageNumber="284">Fig. 3a</figureCitation>
|
||
) with tergites brownish-black dorsally (S1–2 with a green shine,
|
||
<figureCitation id="13442A3A9803FFA1FB55F983FAA5F9AA" box="[1212,1300,1622,1647]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="3" pageNumber="284">Fig. 4a</figureCitation>
|
||
), yellowish ventrally; border of colors distinct on S1–2 and S10, but indistinct on S3–9. S1–7 tergites with indistinctly darkened apices, so that dark colour extends to ventral sides and becomes more saturated-black on dorsum, and indistinctly lightened bases, so that yellowish-grey colour extends dorsally; there is also some lightening proximally of apical darkened parts. S10 tergite cream white with a large triangular dorsal brown spot with indistinct borders; appendages cream white. All sternites yellowish. S10 posterior margin with a small, almost rectangular central projection.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9803FFA6FF2EF886FC56FE15" blockId="3.[151,1437,1442,2007]" lastBlockId="4.[151,1436,151,464]" lastPageId="4" lastPageNumber="285" pageId="3" pageNumber="284">
|
||
Cercus bifurcate, its dorsal branch moderately curved down in lateral view (
|
||
<figureCitation id="13442A3A9803FFA1FBFDF887FBD8F8AE" box="[1044,1129,1874,1899]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="3" pageNumber="284">Figs 5a</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9803FFA1FB9DF886FB20F8AE" box="[1140,1169,1875,1899]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="3" pageNumber="284">7e</figureCitation>
|
||
), with a slightly convex dorsal margin and convex for proximal two-thirds of ventral margin while distal third very slightly convex so that dorsal branch expands in its distal half; apex bluntly rounded. In dorsal view (
|
||
<figureCitation id="13442A3A9803FFA1FBC4F84FFB39F876" box="[1069,1160,1946,1971]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="3" pageNumber="284">Figs 5b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9803FFA1FB7EF84FFB03F876" box="[1175,1202,1946,1971]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="3" pageNumber="284">7f</figureCitation>
|
||
), dorsal branches in general look similar to their lateral view, gradually curving towards each other, their outer margins convex, proximal two-thirds of inner margin concave, especially in middle third; distal third of inner margin very slightly convex, apices bluntly rounded. A small process of dorsal branch of cercus at ca one-fourth of its length, directed up and inside, visible in both dorsal and lateral view (
|
||
<figureCitation id="13442A3A9804FFA6FCEFFF0AFCCBFF3D" box="[774,890,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="4" pageNumber="285">Figs 5a–b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9804FFA6FC61FF0AFC0AFF3D" box="[904,955,223,248]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="4" pageNumber="285">7e–f</figureCitation>
|
||
) and looking bigger and blunter in dorsal view. Ventral branches of cerci 0.6 times as long as dorsal branches, in ventral (
|
||
<figureCitation id="13442A3A9804FFA6FBE4FED1FBD0FED8" box="[1037,1121,260,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="4" pageNumber="285">Figs 5a</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9804FFA6FB84FED1FB39FED9" box="[1133,1160,260,284]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="4" pageNumber="285">7e</figureCitation>
|
||
) and dorsal (
|
||
<figureCitation id="13442A3A9804FFA6FAF0FED1FAC0FED8" box="[1305,1393,260,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="4" pageNumber="285">Figs 5b</figureCitation>
|
||
.
|
||
<figureCitation id="13442A3A9804FFA6FA92FED1FA25FED8" box="[1403,1428,260,285]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="4" pageNumber="285">7f</figureCitation>
|
||
) views narrow, slightly tapering but with expanded apices truncated at ca 45° to body axis, with very slightly concave, almost straight posterioadaxial margins, dorsally with black sclerotised ridges. In lateral view (
|
||
<figureCitation id="13442A3A9804FFA6FAA9FE99FA26FEA0" box="[1344,1431,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="4" pageNumber="285">Figs 5a</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9804FFA6FF7EFEA5FF05FE4D" box="[151,180,368,392]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="4" pageNumber="285">7e</figureCitation>
|
||
), ventral branch narrows considerably to a ‘neck’ just before apex, which expands in a long triangular manner, with a rounded ventral angle, acute dorsoposterior angle and straight ventroposterior margin. Paraprocts very short, bluntly triangular with rounded apices as seen in ventral view (
|
||
<figureCitation id="13442A3A9804FFA6FCB1FE62FC05FE15" box="[856,948,439,464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="4" pageNumber="285">Figs. 5c</figureCitation>
|
||
.
|
||
<figureCitation id="13442A3A9804FFA6FC29FE62FC6BFE15" box="[960,986,439,464]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="4" pageNumber="285">7f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0066379804FFA6FF7EFAF8FDDAFAA7" httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="4" pageNumber="285" startId="4.[151,250,1325,1347]" targetBox="[151,1436,517,1304]" targetPageId="4">
|
||
<paragraph id="8BC036BF9804FFA6FF7EFAF8FDDAFAA7" blockId="4.[151,1436,1325,1378]" pageId="4" pageNumber="285">
|
||
<emphasis id="B90BEAAD9804FFA6FF7EFAF8FEA5FA87" bold="true" box="[151,276,1325,1347]" pageId="4" pageNumber="285">FIGURE 3.</emphasis>
|
||
The freshly collected holotype, male (a) and female paratype (b) of
|
||
<taxonomicName id="4C7F4D3C9804FFA6FC34FAF8FB77FA86" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[989,1222,1325,1347]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9804FFA6FC34FAF8FB77FA86" box="[989,1222,1325,1347]" italics="true" pageId="4" pageNumber="285">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69804FFA6FB3DFAF8FA97FA87" box="[1236,1318,1325,1346]" pageId="4" pageNumber="285" rank="species">
|
||
<emphasis id="B90BEAAD9804FFA6FB3DFAF8FA97FA87" bold="true" box="[1236,1318,1325,1346]" pageId="4" pageNumber="285">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
at the type locality. 4.3 mm between lines. Photo by O.K.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036BF9804FFA6FF2EFA5CFBD4FA03" blockId="4.[151,1437,1417,2018]" pageId="4" pageNumber="285">
|
||
Genital valves (
|
||
<figureCitation id="13442A3A9804FFA6FE93FA5CFE78FA67" box="[378,457,1417,1442]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="4" pageNumber="285">Fig. 4f</figureCitation>
|
||
) leafy, touching each other with their convex inner margins, with triangularly pointed posterior projections at level of segment joint, and an expanded lateroposterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9804FFA6FF2EFA07FBDBFA2C" blockId="4.[151,1437,1417,2018]" box="[199,1130,1489,1514]" pageId="4" pageNumber="285">Measurements (in mm). Abdomen without appendages 33; HW 19; total length 40.</paragraph>
|
||
<paragraph id="8BC036BF9804FFA6FF2EFA20FB04F9BF" blockId="4.[151,1437,1417,2018]" pageId="4" pageNumber="285">
|
||
<emphasis id="B90BEAAD9804FFA6FF2EFA20FE10F9CB" bold="true" box="[199,417,1525,1550]" pageId="4" pageNumber="285">Variation in males</emphasis>
|
||
scarce (
|
||
<figureCitation id="13442A3A9804FFA6FE17FA20FDF1F9CB" box="[510,576,1525,1550]" captionStart="FIGURE 1" captionStartId="2.[151,250,1170,1192]" captionTargetBox="[151,1436,187,1155]" captionTargetId="figure@2.[151,1436,187,1155]" captionTargetPageId="2" captionText="FIGURE 1. Males of Amphicnemis valentini, sp. nov. in nature: a—the type locality, Cambodia, Ream Peninsula, 4 iii 2017, photo by O.K.; b-d—Vietnam, Kiên Giang Province, Phú Quốc Island, 20 viii 2016, photos by T.K." httpUri="https://zenodo.org/record/1283983/files/figure.png" pageId="4" pageNumber="285">Fig. 1</figureCitation>
|
||
). The HW length
|
||
<quantity id="4C879B5A9804FFA6FCE6FA23FC3BF9C8" box="[783,906,1526,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="4" pageNumber="285" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">18–21 mm</quantity>
|
||
, the number of FW Px 11–14. Total length
|
||
<quantity id="4C879B5A9804FFA6FA9BFA23FF5FF9F4" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="4" pageNumber="285" unit="mm" value="40.0" valueMax="41.0" valueMin="39.0">39– 41 mm</quantity>
|
||
, abdomen without appendages
|
||
<quantity id="4C879B5A9804FFA6FD8DF9CCFD57F9F4" box="[612,742,1561,1585]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="4" pageNumber="285" unit="mm" value="32.0" valueMax="33.0" valueMin="31.0">31–33 mm</quantity>
|
||
. In one Cambodian male
|
||
<typeStatus id="54C4881D9804FFA6FBC8F9CFFB30F9F7" box="[1057,1153,1562,1586]" pageId="4" pageNumber="285" type="paratype">paratype</typeStatus>
|
||
the prothoracic horn is strongly inclined to the left side as in the
|
||
<typeStatus id="54C4881D9804FFA6FD7EF9E8FD48F993" box="[663,761,1597,1622]" pageId="4" pageNumber="285" type="holotype">holotype</typeStatus>
|
||
(perhaps just individual deviations of this long and thin structure) and is broken in the other, but is directed perfectly ahead in all Phú Quốc
|
||
<typeStatus id="54C4881D9804FFA6FBABF9B7FB00F9BF" box="[1090,1201,1634,1658]" pageId="4" pageNumber="285" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9804FFA6FF2EF950FCCFF95B" blockId="4.[151,1437,1417,2018]" box="[199,894,1669,1694]" pageId="4" pageNumber="285">
|
||
<emphasis id="B90BEAAD9804FFA6FF2EF950FD92F95B" bold="true" box="[199,547,1669,1694]" pageId="4" pageNumber="285">Description of mature female</emphasis>
|
||
(from
|
||
<collectingCountry id="F368762F9804FFA6FD98F950FD5BF95B" box="[625,746,1669,1694]" name="Cambodia" pageId="4" pageNumber="285">Cambodia</collectingCountry>
|
||
). (
|
||
<figureCitation id="13442A3A9804FFA6FCEAF950FCEBF95B" box="[771,858,1669,1694]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="4" pageNumber="285">Figs 3b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9804FFA6FC8FF950FCC4F95B" box="[870,885,1669,1694]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="4" pageNumber="285">6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9804FFA6FF2EF97CFEE8F923" blockId="4.[151,1437,1417,2018]" pageId="4" pageNumber="285">Resembles male but ca 15% larger (but abdomen only ca 12% longer), and with different colouration of thorax and abdomen tip.</paragraph>
|
||
<paragraph id="8BC036BF9804FFA7FF2EF924FBF6FED8" blockId="4.[151,1437,1417,2018]" lastBlockId="5.[151,1437,151,1941]" lastPageId="5" lastPageNumber="286" pageId="4" pageNumber="285">
|
||
<emphasis id="B90BEAAD9804FFA6FF2EF924FEB5F8CF" box="[199,260,1777,1802]" italics="true" pageId="4" pageNumber="285">Head</emphasis>
|
||
(
|
||
<figureCitation id="13442A3A9804FFA6FEFAF924FED2F8CF" box="[275,355,1777,1802]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="4" pageNumber="285">Fig. 6e</figureCitation>
|
||
) mostly glossy black, its dorsal side with a strong purple lustre at sides of ocellar bulge and two patches of strong bluish-green lustre near compound eyes; face black with some pale markings. Labium yellowish. Lower half of labrum yellowish, upper one black, the latter colour extending to middle of labrum centrally, but to four-fifth laterally; border of colour indistinct. Mandible bases yellowish with large, somewhat indistinct, rounded brown spots centrally. Clypeus coloured as in male. Genae greenish below, brownish-black above, with greenish colour extending dorsally along compound eye margins and towards anteclypeus, bordering narrowly triangular downward projections of black colour. Antefrons mostly greenish, its dorsal part black, border of colours distinct and wavy; below antenna bases there is a pair of rounded trapezoid greenish spots; remainder of postfrons black. Antennae blackish-brown but anterior parts of segments 1 and 2 and apex of segment 2 yellowish. Occiput with long light-brown bristles and a short yellowish-brown streak along occipital ridge. Eyes in life saturated bluishgreen, above changing to darker olive-green, yellowish-green in the dried specimen.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFEF2FA9EFE30" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFEF2FEAEFE85" box="[199,287,295,320]" italics="true" pageId="5" pageNumber="286">Thorax.</emphasis>
|
||
Prothorax (
|
||
<figureCitation id="13442A3A9805FFA7FE4EFEF2FDA4FE85" box="[423,533,295,320]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6c–d</figureCitation>
|
||
) entirely light bluish-green when alive, greenish-brown in the dried specimen; posterior lobe with a long central horn (
|
||
<figureCitation id="13442A3A9805FFA7FDBCFE99FD16FEA0" box="[597,679,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6e</figureCitation>
|
||
), curving anteriorly in basal half and straight at distal half, directed up and forward at about 45° angle and inclined to the left (
|
||
<figureCitation id="13442A3A9805FFA7FCC0FEBAFCCDFE4D" box="[809,892,367,392]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6d</figureCitation>
|
||
). Horn looks like a half-rolled flat plate, with its anterior (dorsal) surface convex and posterior (ventral) surface concave, apex rounded. Lateral projections of posterior lobe large, pointed and separated from central horn base by deep rounded incisions. In dorsal view, prothorax posterior lobe smoothly rounded, but incision of central horn, with lateral lobes hooked inside.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFE2AFBD1FD40" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
Mesostigmal plate seen from above (
|
||
<figureCitation id="13442A3A9805FFA7FD9FFE2AFD78FDDD" box="[630,713,511,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6c</figureCitation>
|
||
) composed of two yellowish, transverse, hollowed parts, long triangular with slightly convex margins and slight rounded anterior ridges at middorsal suture. Synthorax (
|
||
<figureCitation id="13442A3A9805FFA7FAA9FDF1FA26FDF8" box="[1344,1431,548,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="5" pageNumber="286">Figs 3b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9805FFA7FF7EFD92FF03FDA5" box="[151,178,583,608]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">6a</figureCitation>
|
||
) saturated olivaceous-green dorsally changing to saturated bluish-green at sides when alive, ochraceous to dirty bluish-greenish in the dried specimen; synthorax venter whitish without dark marking.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFD45FD16FD35" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
Legs (
|
||
<figureCitation id="13442A3A9805FFA7FEE6FD5AFED8FD6D" box="[271,361,655,680]" captionStart="FIGURE 3" captionStartId="4.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@4.[151,1436,517,1304]" captionTargetPageId="4" captionText="FIGURE 3. The freshly collected holotype, male (a) and female paratype (b) of Amphicnemis valentini, sp. nov. at the type locality. 4.3 mm between lines. Photo by O.K." httpUri="https://zenodo.org/record/1283987/files/figure.png" pageId="5" pageNumber="286">Figs 3b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9805FFA7FE9EFD5AFE23FD6D" box="[375,402,655,680]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">6a</figureCitation>
|
||
) yellowish-white with blackish-brown parts as follows: continuous stripes along extensor surface of femora (broader at profemora), femora apices, protibia bases, segment joints of tarsi; spines blackishbrown, claws light brown with blackish apices.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFD28FE07FC45" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFD28FEBBFCD1" box="[199,266,765,788]" italics="true" pageId="5" pageNumber="286">Wings</emphasis>
|
||
as in male but larger: antenodals Px:
|
||
<quantity id="4C879B5A9805FFA7FD59FD29FD58FCD0" box="[688,745,764,789]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="286" unit="in" value="13.0">13 in</quantity>
|
||
FW,
|
||
<quantity id="4C879B5A9805FFA7FCCCFD29FCEFFCD0" box="[805,862,764,789]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="286" unit="in" value="12.0">12 in</quantity>
|
||
HW. Pterostigmata (
|
||
<figureCitation id="13442A3A9805FFA7FBAEFD29FB28FCD0" box="[1095,1177,764,789]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6b</figureCitation>
|
||
) similar in both wings, brownish-black, with a light margin, costal side twice as short as inner side, which is slightly shorter than underlying cell; both transversal sides convex, proximal one slanting, distal one perpendicular; distal half of its ventral margin thickened.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFC59FC5CFC28" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFC59FE80FC60" box="[199,305,908,933]" italics="true" pageId="5" pageNumber="286">Abdomen</emphasis>
|
||
as in male with differences at last segments (
|
||
<figureCitation id="13442A3A9805FFA7FC6AFC59FC6CFC60" box="[899,989,908,933]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">Fig. 6f</figureCitation>
|
||
): S8 tergite indistinctly lightened at anterioventral corner, S9 with a pair of large dorsolateral whitish spots, S10 whitish, changing pale brownish below, appendages whitish, ovipositor yellowish-white, brownish at margin.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFC22FEB5FB9D" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFC22FE1DFBD5" bold="true" box="[199,428,1015,1040]" pageId="5" pageNumber="286">Immature females.</emphasis>
|
||
Immature females have crimson red thorax (including prothorax) and legs, somewhat paler ventrally, but are otherwise colored as mature ones (
|
||
<figureCitation id="13442A3A9805FFA7FCABFBC9FC08FBF1" box="[834,953,1052,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1350,1372]" captionTargetBox="[151,1436,193,1329]" captionTargetId="figure@3.[151,1436,193,1329]" captionTargetPageId="3" captionText="FIGURE 2. Immature (a,c) and mature (b,d) females of Amphicnemis valentini, sp. nov. in nature, Vietnam, Kiên Giang Province, Phú Quốc Island, 19 (a-b) and 20 (c-d) viii 2016, photos by T.K." httpUri="https://zenodo.org/record/1283985/files/figure.png" pageId="5" pageNumber="286">Figs 2a, c</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9805FFA7FC20FBC9FC68FBF1" box="[969,985,1052,1076]" captionStart="FIGURE 8" captionStartId="10.[151,250,1339,1361]" captionTargetBox="[159,1423,193,1315]" captionTargetId="figure@10.[159,1423,193,1315]" captionTargetPageId="10" captionText="FIGURE 8. Details of paratypes of Amphicnemis valentini, sp. nov. from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—ditto, dorsal view. Not to scale." httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="5" pageNumber="286">8</figureCitation>
|
||
). Transitory stages to mature state not observed.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFBB1FABAFAC9" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFBB1FE0EFBB8" bold="true" box="[199,447,1124,1149]" pageId="5" pageNumber="286">Variation in females</emphasis>
|
||
scarce (
|
||
<figureCitation id="13442A3A9805FFA7FDCDFBB1FD2AFBB8" box="[548,667,1124,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1350,1372]" captionTargetBox="[151,1436,193,1329]" captionTargetId="figure@3.[151,1436,193,1329]" captionTargetPageId="3" captionText="FIGURE 2. Immature (a,c) and mature (b,d) females of Amphicnemis valentini, sp. nov. in nature, Vietnam, Kiên Giang Province, Phú Quốc Island, 19 (a-b) and 20 (c-d) viii 2016, photos by T.K." httpUri="https://zenodo.org/record/1283985/files/figure.png" pageId="5" pageNumber="286">Fig. 2b, d</figureCitation>
|
||
). All female
|
||
<typeStatus id="54C4881D9805FFA7FCD3FBB0FC17FBB8" box="[826,934,1125,1149]" pageId="5" pageNumber="286" type="paratype">paratypes</typeStatus>
|
||
from Phú Quốc have the prothoracic horn directed anteriorly, not inclined to any side. In the Phú Quốc female series, it is strongly curved, as in males, in three
|
||
<typeStatus id="54C4881D9805FFA7FF3EFB78FEF3FB00" box="[215,322,1197,1221]" pageId="5" pageNumber="286" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="13442A3A9805FFA7FEBDFB79FE28FB01" box="[340,409,1196,1221]" captionStart="FIGURE 8" captionStartId="10.[151,250,1339,1361]" captionTargetBox="[159,1423,193,1315]" captionTargetId="figure@10.[159,1423,193,1315]" captionTargetPageId="10" captionText="FIGURE 8. Details of paratypes of Amphicnemis valentini, sp. nov. from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—ditto, dorsal view. Not to scale." httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="5" pageNumber="286">Fig. 8</figureCitation>
|
||
), slightly curved in one, virtually straight and raised upward more than in the Cambodian
|
||
<typeStatus id="54C4881D9805FFA7FF7EFB04FF46FB2C" box="[151,247,1233,1257]" pageId="5" pageNumber="286" type="paratype">paratype</typeStatus>
|
||
in one (and is distorted in one). The lateral projections are almost absent and blunt in the immature female (
|
||
<figureCitation id="13442A3A9805FFA7FF76FB21FF52FAC9" box="[159,227,1268,1293]" captionStart="FIGURE 8" captionStartId="10.[151,250,1339,1361]" captionTargetBox="[159,1423,193,1315]" captionTargetId="figure@10.[159,1423,193,1315]" captionTargetPageId="10" captionText="FIGURE 8. Details of paratypes of Amphicnemis valentini, sp. nov. from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—ditto, dorsal view. Not to scale." httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="5" pageNumber="286">Fig. 8</figureCitation>
|
||
), but well developed in the five mature ones, somewhat varying in curvature. Px in HW12–14.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFAC2FAEBFAF5" blockId="5.[151,1437,151,1941]" box="[199,1370,1303,1328]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFAC2FD8FFAF5" bold="true" box="[199,574,1303,1328]" pageId="5" pageNumber="286">Measurements of females (mm)</emphasis>
|
||
. Abdomen without appendages 33–37; HW 21–23; total length 39–43.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565349805FFA8FF2EFAE9FB35F910" lastPageId="10" lastPageNumber="291" pageId="5" pageNumber="286" type="diagnosis">
|
||
<paragraph id="8BC036BF9805FFA7FF2EFAE9FBE7FA58" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFAE9FE38FA90" bold="true" box="[199,393,1340,1365]" pageId="5" pageNumber="286">Short diagnosis.</emphasis>
|
||
An
|
||
<taxonomicName id="4C7F4D3C9805FFA7FE54FAE9FDE5FA90" authorityName="Selys" authorityYear="1863" box="[445,596,1340,1365]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD9805FFA7FE54FAE9FDE5FA90" box="[445,596,1340,1365]" italics="true" pageId="5" pageNumber="286">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
species with a long process on prothorax posterior lobe, strongly curved anteriorly in males, straight to strongly curved and directed dorsally and anteriorly in females, a slightly arched upper male cercus branch with a small basodorsal projection directed up- and inward.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EFA72FE53F978" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
<emphasis id="B90BEAAD9805FFA7FF2EFA72FE61FA05" bold="true" box="[199,464,1447,1472]" pageId="5" pageNumber="286">Differential diagnosis.</emphasis>
|
||
The new species belongs to the
|
||
<taxonomicName id="4C7F4D3C9805FFA7FCA5FA72FC52FA05" authorityName="Selys" authorityYear="1863" box="[844,995,1447,1472]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD9805FFA7FCA5FA72FC52FA05" box="[844,995,1447,1472]" italics="true" pageId="5" pageNumber="286">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C7F4D3C9805FFA7FC1FFA72FBE1FA05" authorityName="Selys" authorityYear="1863" box="[1014,1104,1447,1472]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="wallacii">
|
||
<emphasis id="B90BEAAD9805FFA7FC1FFA72FBE1FA05" box="[1014,1104,1447,1472]" italics="true" pageId="5" pageNumber="286">wallacii</emphasis>
|
||
</taxonomicName>
|
||
) species group suggested by
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FF7EFA18FE92FA21" box="[151,291,1484,1508]" pageId="5" pageNumber="286" refId="ref6472" refString="Dow, R. A. (2014) Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae). Odonatologica, 43 (1 / 2), 67 - 77." type="journal article">Dow (2014)</bibRefCitation>
|
||
, which includes 12 species characterised by unicolorous pterostigmata, similar in all wings (
|
||
<figureCitation id="13442A3A9805FFA7FAD6FA19FA26FA20" box="[1343,1431,1484,1509]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="5" pageNumber="286">Figs 4b</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9805FFA7FF7EFA3AFF07F9CD" box="[151,182,1519,1544]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="5" pageNumber="286">6b</figureCitation>
|
||
). In this group, five species, including the current new species, have males with a long prothoracic horn. Of these species,
|
||
<taxonomicName id="4C7F4D3C9805FFA7FED2F9C0FDEBF9E9" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" box="[315,602,1556,1581]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="annae">
|
||
<emphasis id="B90BEAAD9805FFA7FED2F9C0FE10F9E9" box="[315,417,1557,1580]" italics="true" pageId="5" pageNumber="286">A. annae</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FE42F9C1FDEBF9E9" box="[427,602,1556,1581]" pageId="5" pageNumber="286" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
has this horn much longer than the prothorax median lobe; in
|
||
<taxonomicName id="4C7F4D3C9805FFA7FAD8F9C0FEA0F995" authority="Dow, 2014" authorityName="Dow" authorityYear="2014" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="triplex">
|
||
<emphasis id="B90BEAAD9805FFA7FAD8F9C0FA2DF9E8" box="[1329,1436,1556,1581]" italics="true" pageId="5" pageNumber="286">A. triplex</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FF7EF9ECFEA0F995" box="[151,273,1592,1616]" pageId="5" pageNumber="286" refId="ref6472" refString="Dow, R. A. (2014) Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae). Odonatologica, 43 (1 / 2), 67 - 77." type="journal article">Dow, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
it is slightly longer; and in
|
||
<taxonomicName id="4C7F4D3C9805FFA7FDAFF9EDFCCEF995" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" box="[582,895,1591,1616]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="billitonis">
|
||
<emphasis id="B90BEAAD9805FFA7FDAFF9EDFD78F995" box="[582,713,1591,1616]" italics="true" pageId="5" pageNumber="286">A. billitonis</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FD26F9E2FCCEF995" box="[719,895,1591,1616]" pageId="5" pageNumber="286" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D3C9805FFA7FC63F9EDFB37F995" authority="Selys, 1863" authorityName="Selys" authorityYear="1863" box="[906,1158,1591,1616]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="wallacii">
|
||
<emphasis id="B90BEAAD9805FFA7FC63F9EDFC4CF995" box="[906,1021,1591,1616]" italics="true" pageId="5" pageNumber="286">A wallacii</emphasis>
|
||
Selys, 1863
|
||
</taxonomicName>
|
||
and the new species it is shorter than the prothorax median lobe (
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FD89F989FCBDF9B1" box="[608,780,1628,1653]" pageId="5" pageNumber="286" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck 1940</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FCF0F989FCE5F9B0" box="[793,852,1628,1653]" pageId="5" pageNumber="286" refId="ref7128" refString="Lieftinck, M. A. (1953) Additions to the odonate fauna of the Indo-Australian archipelago. Treubia, 22, 233 - 269." type="journal article">1953</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FC88F988FBACF9B1" box="[865,1053,1628,1653]" pageId="5" pageNumber="286" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9805FFA7FC49F988FC6BF9B0" box="[928,986,1628,1653]" italics="true" pageId="5" pageNumber="286">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FBC3F988FB17F9B1" box="[1066,1190,1628,1652]" pageId="5" pageNumber="286" refId="ref6472" refString="Dow, R. A. (2014) Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae). Odonatologica, 43 (1 / 2), 67 - 77." type="journal article">Dow 2014</bibRefCitation>
|
||
). In males of the new species this horn (
|
||
<figureCitation id="13442A3A9805FFA7FE81F9AAFE0BF95D" box="[360,442,1663,1688]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="5" pageNumber="286">Fig. 4c</figureCitation>
|
||
) is similar to that of
|
||
<taxonomicName id="4C7F4D3C9805FFA7FD5BF955FC9CF95D" authorityName="Selys" authorityYear="1863" box="[690,813,1663,1688]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="wallacii">
|
||
<emphasis id="B90BEAAD9805FFA7FD5BF955FC9CF95D" box="[690,813,1663,1688]" italics="true" pageId="5" pageNumber="286">A. wallacii</emphasis>
|
||
</taxonomicName>
|
||
and especially of
|
||
<taxonomicName id="4C7F4D3C9805FFA7FBEAF955FAFFF95D" authority="(Lieftinck 1940)" authorityName="Lieftinck" authorityYear="1940" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[1027,1358,1663,1688]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="billitonis">
|
||
<emphasis id="B90BEAAD9805FFA7FBEAF955FB39F95D" box="[1027,1160,1663,1688]" italics="true" pageId="5" pageNumber="286">A. billitonis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FB71F9AAFAF7F95D" box="[1176,1350,1663,1688]" pageId="5" pageNumber="286" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but is much more curved anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9805FFA7FF2EF912FBB7F851" blockId="5.[151,1437,151,1941]" pageId="5" pageNumber="286">
|
||
While the new species is most similar to
|
||
<taxonomicName id="4C7F4D3C9805FFA7FD75F91DFC91F925" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[668,800,1735,1760]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="billitonis">
|
||
<emphasis id="B90BEAAD9805FFA7FD75F91DFC91F925" box="[668,800,1735,1760]" italics="true" pageId="5" pageNumber="286">A. billitonis</emphasis>
|
||
</taxonomicName>
|
||
in the structure of its prothorax and differs from that of the hornless males of
|
||
<taxonomicName id="4C7F4D3C9805FFA7FE76F938FDA9F8C0" authorityName="Kruger" authorityYear="1898" box="[415,536,1772,1797]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9805FFA7FE76F938FDA9F8C0" box="[415,536,1772,1797]" italics="true" pageId="5" pageNumber="286">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FDB4F938FC94F8C1" box="[605,805,1772,1797]" pageId="5" pageNumber="286" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9805FFA7FD49F938FD6CF8C0" box="[672,733,1772,1797]" italics="true" pageId="5" pageNumber="286">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
:
|
||
<figureCitation id="13442A3A9805FFA7FCDDF939FCCAF8C1" box="[820,891,1772,1797]" captionStart="FIGURE 8" captionStartId="10.[151,250,1339,1361]" captionTargetBox="[159,1423,193,1315]" captionTargetId="figure@10.[159,1423,193,1315]" captionTargetPageId="10" captionText="FIGURE 8. Details of paratypes of Amphicnemis valentini, sp. nov. from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—ditto, dorsal view. Not to scale." httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="5" pageNumber="286">Fig. 8</figureCitation>
|
||
), its male appendages (
|
||
<figureCitation id="13442A3A9805FFA7FB7FF939FB55F8C0" box="[1174,1252,1772,1797]" captionStart="FIGURE 5" captionStartId="7.[151,250,1973,1995]" captionTargetBox="[275,1299,193,1916]" captionTargetId="figure@7.[275,1299,193,1916]" captionTargetPageId="7" captionText="FIGURE 5. Amphicnemis valentini, sp. nov., anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="5" pageNumber="286">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="13442A3A9805FFA7FB1CF939FA9DF8C1" box="[1269,1324,1772,1796]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="5" pageNumber="286">7e–g</figureCitation>
|
||
) strongly resemble those in
|
||
<taxonomicName id="4C7F4D3C9805FFA7FE82F8C5FE52F8ED" authorityName="Kruger" authorityYear="1898" box="[363,483,1807,1832]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9805FFA7FE82F8C5FE52F8ED" box="[363,483,1807,1832]" italics="true" pageId="5" pageNumber="286">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13442A3A9805FFA7FE1FF8DAFDD5F8ED" box="[502,612,1807,1832]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="5" pageNumber="286">Fig. 7a–d</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FD97F8DAFC9DF8ED" box="[638,812,1807,1832]" pageId="5" pageNumber="286" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck 1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9805FFA7FCD2F8C4FC4FF8ED" box="[827,1022,1807,1832]" pageId="5" pageNumber="286" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9805FFA7FC95F8C4FC06F8ED" box="[892,951,1807,1832]" italics="true" pageId="5" pageNumber="286">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). The upper branch of the cercus is similar, but the expanded distal part is relatively longer, and in lateral view (
|
||
<figureCitation id="13442A3A9805FFA7FC02F8E1FB8AF889" box="[1003,1083,1844,1869]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="5" pageNumber="286">Fig. 7e</figureCitation>
|
||
) better defined, with a hint of a slight ‘heel’ at its proximal part, and with a much less convex lower margin. The branch itself is more curved in lateral view, with a more concave ventral margin of its proximal part (
|
||
<figureCitation id="13442A3A9805FFA7FC4EF8A9FC48F851" box="[935,1017,1916,1941]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="5" pageNumber="286">Fig. 7e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0066379806FFA4FF7EF840FF5AF82C" httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="6" pageNumber="287" startId="6.[151,250,1941,1963]" targetBox="[151,1436,193,1880]" targetPageId="6">
|
||
<paragraph id="8BC036BF9806FFA4FF7EF840FF5AF82C" blockId="6.[151,1435,1941,2025]" pageId="6" pageNumber="287">
|
||
<emphasis id="B90BEAAD9806FFA4FF7EF840FEA4F86E" bold="true" box="[151,277,1941,1963]" pageId="6" pageNumber="287">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C7F4D3C9806FFA4FEF7F840FDB9F86E" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[286,520,1941,1963]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9806FFA4FEF7F840FDB9F86E" box="[286,520,1941,1963]" italics="true" pageId="6" pageNumber="287">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69806FFA4FDF1F843FDDBF86E" box="[536,618,1942,1963]" pageId="6" pageNumber="287" rank="species">
|
||
<emphasis id="B90BEAAD9806FFA4FDF1F843FDDBF86E" bold="true" box="[536,618,1942,1963]" pageId="6" pageNumber="287">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—
|
||
<emphasis id="B90BEAAD9806FFA4FDB1F861FD37F80F" box="[600,646,1972,1994]" italics="true" pageId="6" pageNumber="287">ditto</emphasis>
|
||
, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0066379807FFA5FF7EF860FE69F82F" httpUri="https://zenodo.org/record/1283991/files/figure.png" pageId="7" pageNumber="288" startId="7.[151,250,1973,1995]" targetBox="[275,1299,193,1916]" targetPageId="7">
|
||
<paragraph id="8BC036BF9807FFA5FF7EF860FE69F82F" blockId="7.[151,1435,1973,2026]" pageId="7" pageNumber="288">
|
||
<emphasis id="B90BEAAD9807FFA5FF7EF860FEA4F80F" bold="true" box="[151,277,1973,1995]" pageId="7" pageNumber="288">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C7F4D3C9807FFA5FEF7F860FDB9F80E" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[286,520,1973,1995]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9807FFA5FEF7F860FDB9F80E" box="[286,520,1973,1995]" italics="true" pageId="7" pageNumber="288">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69807FFA5FDFEF860FDDBF80F" box="[535,618,1973,1994]" pageId="7" pageNumber="288" rank="species">
|
||
<emphasis id="B90BEAAD9807FFA5FDFEF860FDDBF80F" bold="true" box="[535,618,1973,1994]" pageId="7" pageNumber="288">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, anal appendages of the holotype, male: a—lateral view; b—dorsal view; cventral view. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0066379808FFAAFF7EF883FDE7F86C" httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="8" pageNumber="289" startId="8.[151,250,1878,1900]" targetBox="[151,1436,193,1827]" targetPageId="8">
|
||
<paragraph id="8BC036BF9808FFAAFF7EF883FDE7F86C" blockId="8.[151,1436,1878,1961]" pageId="8" pageNumber="289">
|
||
<emphasis id="B90BEAAD9808FFAAFF7EF883FEA2F8AE" bold="true" box="[151,275,1878,1900]" pageId="8" pageNumber="289">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C7F4D3C9808FFAAFEF2F883FDB2F8A9" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[283,515,1878,1900]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9808FFAAFEF2F883FDB2F8A9" box="[283,515,1878,1900]" italics="true" pageId="8" pageNumber="289">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69808FFAAFDF9F883FDD3F8AE" box="[528,610,1878,1899]" pageId="8" pageNumber="289" rank="species">
|
||
<emphasis id="B90BEAAD9808FFAAFDF9F883FDD3F8AE" bold="true" box="[528,610,1878,1899]" pageId="8" pageNumber="289">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—
|
||
<emphasis id="B90BEAAD9808FFAAFC59F8A1FC6FF84F" box="[944,990,1908,1930]" italics="true" pageId="8" pageNumber="289">ditto</emphasis>
|
||
, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0066379809FFABFF7EFC53FD66FC1C" httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="9" pageNumber="290" startId="9.[151,250,902,924]" targetBox="[168,1418,203,843]" targetPageId="9">
|
||
<paragraph id="8BC036BF9809FFABFF7EFC53FD66FC1C" blockId="9.[151,1436,902,985]" pageId="9" pageNumber="290">
|
||
<emphasis id="B90BEAAD9809FFABFF7EFC53FEA2FC5E" bold="true" box="[151,275,902,924]" pageId="9" pageNumber="290">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C7F4D3C9809FFABFEF0FC53FE13FC59" authorityName="Selys" authorityYear="1863" box="[281,418,902,924]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD9809FFABFEF0FC53FE13FC59" box="[281,418,902,924]" italics="true" pageId="9" pageNumber="290">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
spp., male anal appendages of
|
||
<taxonomicName id="4C7F4D3C9809FFABFD09FC52FCF8FC59" authorityName="Kruger" authorityYear="1898" box="[736,841,902,924]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9809FFABFD09FC52FCF8FC59" box="[736,841,902,924]" italics="true" pageId="9" pageNumber="290">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow
|
||
<emphasis id="B90BEAAD9809FFABFD1EFC70FC9BFC7F" box="[759,810,932,954]" italics="true" pageId="9" pageNumber="290">et al.</emphasis>
|
||
(2010: Figs 20, 24) and of
|
||
<taxonomicName id="4C7F4D3C9809FFABFBD2FC70FB1EFC7F" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[1083,1199,932,954]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD9809FFABFBD2FC70FB1EFC7F" box="[1083,1199,932,954]" italics="true" pageId="9" pageNumber="290">A. valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D69809FFABFB52FC70FABAFC7F" box="[1211,1291,933,954]" pageId="9" pageNumber="290" rank="species">
|
||
<emphasis id="B90BEAAD9809FFABFB52FC70FABAFC7F" bold="true" box="[1211,1291,933,954]" pageId="9" pageNumber="290">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036BF9809FFABFF2EFBD5FDD8FB6C" blockId="9.[151,1437,1024,2021]" pageId="9" pageNumber="290">
|
||
The lower cercus branch has a more defined ‘neck’ before its broadened apical part, especially in dorsal and ventral views (
|
||
<figureCitation id="13442A3A9809FFABFED5FBF1FE16FBF8" box="[316,423,1060,1085]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="9" pageNumber="290">Fig. 7f–g</figureCitation>
|
||
). However, this may be an artifact of drying, especially after treatment with acetone, as the structure is quite soft and delicate. The second author made a close-up photo of the appendages of one of the Phú Quốc male
|
||
<typeStatus id="54C4881D9809FFABFEF2FBB8FE36FB40" box="[283,391,1133,1157]" pageId="9" pageNumber="290" type="paratype">paratypes</typeStatus>
|
||
when it was still fresh, and it shows (
|
||
<figureCitation id="13442A3A9809FFABFCD5FBB9FC1BFB40" box="[828,938,1132,1157]" captionStart="FIGURE 8" captionStartId="10.[151,250,1339,1361]" captionTargetBox="[159,1423,193,1315]" captionTargetId="figure@10.[159,1423,193,1315]" captionTargetPageId="10" captionText="FIGURE 8. Details of paratypes of Amphicnemis valentini, sp. nov. from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—ditto, dorsal view. Not to scale." httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="9" pageNumber="290">Fig. 8b–c</figureCitation>
|
||
) the lower branch with only a slight ‘neck’ identical to that of
|
||
<taxonomicName id="4C7F4D3C9809FFABFE83FB44FE6EFB6C" authorityName="Kruger" authorityYear="1898" box="[362,479,1168,1193]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9809FFABFE83FB44FE6EFB6C" box="[362,479,1168,1193]" italics="true" pageId="9" pageNumber="290">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13442A3A9809FFABFE07FB45FDECFB6C" box="[494,605,1168,1193]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="9" pageNumber="290">Fig. 7a–d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9809FFABFF2EFB60FA34FB08" blockId="9.[151,1437,1024,2021]" box="[199,1413,1204,1229]" pageId="9" pageNumber="290">
|
||
In addition, the paraprocts of the new species look smaller in ventral view than those of
|
||
<taxonomicName id="4C7F4D3C9809FFABFB48FB60FAA4FB08" authorityName="Kruger" authorityYear="1898" box="[1185,1301,1204,1229]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9809FFABFB48FB60FAA4FB08" box="[1185,1301,1204,1229]" italics="true" pageId="9" pageNumber="290">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13442A3A9809FFABFACCFB61FAC6FB08" box="[1317,1399,1204,1229]" captionStart="FIGURE 7" captionStartId="9.[151,250,902,924]" captionTargetBox="[168,1418,203,843]" captionTargetId="figure@9.[151,1436,193,880]" captionTargetPageId="9" captionText="FIGURE 7. Amphicnemis spp., male anal appendages of A. gracilis (a–d) from East Sumatra after Lieftinck (1940: Pl. 16—2) and from Peninsular Malaysia, Pahang, Sg Bebar after Dow et al. (2010: Figs 20, 24) and of A. valentini, sp. nov., the holotype; a,c,e—lateral view; b,d,f—dorsal view; g—ventral view." httpUri="https://zenodo.org/record/1283995/files/figure.png" pageId="9" pageNumber="290">Fig. 7e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9809FFABFF2EFB0DFBA6FAD0" blockId="9.[151,1437,1024,2021]" pageId="9" pageNumber="290">
|
||
Male genital valves (
|
||
<figureCitation id="13442A3A9809FFABFE59FB0DFE4CFB34" box="[432,509,1240,1265]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[151,1436,193,1880]" captionTargetId="figure@6.[151,1436,193,1944]" captionTargetPageId="6" captionText="FIGURE 4. Amphicnemis valentini, sp. nov., details of the holotype, male: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—genital valvae, ventral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283989/files/figure.png" pageId="9" pageNumber="290">Fig. 4f</figureCitation>
|
||
) in the new species are of the same shape as in
|
||
<taxonomicName id="4C7F4D3C9809FFABFBE4FB0CFB30FB34" authorityName="Kruger" authorityYear="1898" box="[1037,1153,1240,1265]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B90BEAAD9809FFABFBE4FB0CFB30FB34" box="[1037,1153,1240,1265]" italics="true" pageId="9" pageNumber="290">A. gracilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D3C9809FFABFB5EFB0CFF6AFAD0" authority="(Dow et al. 2010)" authorityName="Dow et al." authorityYear="2010" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="bebar">
|
||
<emphasis id="B90BEAAD9809FFABFB5EFB0CFAA6FB34" box="[1207,1303,1240,1265]" italics="true" pageId="9" pageNumber="290">A. bebar</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B4E9809FFABFACFFB0CFF62FAD0" pageId="9" pageNumber="290" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9809FFABFA8DFB0FFA2DFB34" box="[1380,1436,1240,1265]" italics="true" pageId="9" pageNumber="290">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but their inner margins are nearly touching, rather than clearly separated.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9809FFABFF2EFAF4FC5CF99D" blockId="9.[151,1437,1024,2021]" pageId="9" pageNumber="290">
|
||
A more or less long prothoracic horn is found in females of 10 of 12 species of the
|
||
<taxonomicName id="4C7F4D3C9809FFABFB66FAF5FA97FAFC" authorityName="Selys" authorityYear="1863" box="[1167,1318,1312,1337]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD9809FFABFB66FAF5FA97FAFC" box="[1167,1318,1312,1337]" italics="true" pageId="9" pageNumber="290">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C7F4D3C9809FFABFAD1FAF5FA23FAFC" authorityName="Selys" authorityYear="1863" box="[1336,1426,1312,1337]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="wallacii">
|
||
<emphasis id="B90BEAAD9809FFABFAD1FAF5FA23FAFC" box="[1336,1426,1312,1337]" italics="true" pageId="9" pageNumber="290">wallacii</emphasis>
|
||
</taxonomicName>
|
||
) group including the abovementioned ones and
|
||
<taxonomicName id="4C7F4D3C9809FFABFD4BFA90FB82FA98" authority="Dow, Choong & Ng, 2010" authorityName="Dow, Choong & Ng" authorityYear="2010" box="[674,1075,1348,1373]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="bebar">
|
||
<emphasis id="B90BEAAD9809FFABFD4BFA90FCB3FA98" box="[674,770,1348,1373]" italics="true" pageId="9" pageNumber="290">A. bebar</emphasis>
|
||
Dow,
|
||
<bibRefCitation id="EFEE4B4E9809FFABFCA5FA91FB82FA98" box="[844,1075,1348,1373]" pageId="9" pageNumber="290" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">Choong & Ng, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D3C9809FFABFBD7FA90FADCFA98" authority="Lieftinck, 1953" authorityName="Lieftinck" authorityYear="1953" box="[1086,1389,1348,1373]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="erminea">
|
||
<emphasis id="B90BEAAD9809FFABFBD7FA90FB08FA98" box="[1086,1209,1349,1373]" italics="true" pageId="9" pageNumber="290">A. erminea</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9809FFABFB29FA91FADCFA98" box="[1216,1389,1348,1373]" pageId="9" pageNumber="290" refId="ref7128" refString="Lieftinck, M. A. (1953) Additions to the odonate fauna of the Indo-Australian archipelago. Treubia, 22, 233 - 269." type="journal article">Lieftinck, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D3C9809FFABFF7EFABCFE36FA44" authority="Dow, 2014" authorityName="Dow" authorityYear="2014" box="[151,391,1384,1409]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="triplex">
|
||
<emphasis id="B90BEAAD9809FFABFF7EFABCFEB1FA44" box="[151,256,1384,1409]" italics="true" pageId="9" pageNumber="290">A. triplex</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9809FFABFEE1FABCFE36FA44" box="[264,391,1385,1409]" pageId="9" pageNumber="290" refId="ref6472" refString="Dow, R. A. (2014) Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae). Odonatologica, 43 (1 / 2), 67 - 77." type="journal article">Dow, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, and is absent only in
|
||
<taxonomicName id="4C7F4D3C9809FFABFD6DFABCFC18FA44" authority="Lieftinck, 1937" authorityName="Lieftinck" authorityYear="1937" box="[644,937,1384,1409]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="kuiperi">
|
||
<emphasis id="B90BEAAD9809FFABFD6DFABCFD42FA44" box="[644,755,1384,1409]" italics="true" pageId="9" pageNumber="290">A. kuiperi</emphasis>
|
||
Lieftinck, 1937
|
||
</taxonomicName>
|
||
(no horn) and
|
||
<taxonomicName id="4C7F4D3C9809FFABFBBAFABCFACFFA44" authority="Laidlaw, 1926" authorityName="Laidlaw" authorityYear="1926" box="[1107,1406,1384,1409]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="smedleyi">
|
||
<emphasis id="B90BEAAD9809FFABFBBAFABCFB67FA44" box="[1107,1238,1384,1409]" italics="true" pageId="9" pageNumber="290">A. smedleyi</emphasis>
|
||
Laidlaw, 1926
|
||
</taxonomicName>
|
||
(a very short horn) (
|
||
<bibRefCitation id="EFEE4B4E9809FFABFEB1FA59FDB3FA60" box="[344,514,1420,1445]" pageId="9" pageNumber="290" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck 1940</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEE4B4E9809FFABFDE5FA59FDF6FA60" box="[524,583,1420,1445]" pageId="9" pageNumber="290" refId="ref7128" refString="Lieftinck, M. A. (1953) Additions to the odonate fauna of the Indo-Australian archipelago. Treubia, 22, 233 - 269." type="journal article">1953</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9809FFABFDBBFA58FCB9FA60" box="[594,776,1420,1445]" pageId="9" pageNumber="290" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9809FFABFD66FA5BFD76FA60" box="[655,711,1420,1445]" italics="true" pageId="9" pageNumber="290">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9809FFABFCFAFA58FC3DFA60" box="[787,908,1421,1445]" pageId="9" pageNumber="290" refId="ref6472" refString="Dow, R. A. (2014) Amphicnemis triplex sp. nov. from Central Kalimantan, Indonesia (Odonata: Coenagrionidae). Odonatologica, 43 (1 / 2), 67 - 77." type="journal article">Dow 2014</bibRefCitation>
|
||
), but no female descriptions were published for
|
||
<taxonomicName id="4C7F4D3C9809FFABFF7EFA64FE72FA0C" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" box="[151,451,1456,1481]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B90BEAAD9809FFABFF7EFA64FEB8FA0C" box="[151,265,1457,1481]" italics="true" pageId="9" pageNumber="290">A. mariae</emphasis>
|
||
<bibRefCitation id="EFEE4B4E9809FFABFEFAFA65FE72FA0C" box="[275,451,1456,1481]" pageId="9" pageNumber="290" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D3C9809FFABFDE9FA64FC9DFA0C" authority="Laidlaw, 1912" authorityName="Laidlaw" authorityYear="1912" box="[512,812,1456,1481]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="remiger">
|
||
<emphasis id="B90BEAAD9809FFABFDE9FA64FDCAFA0C" box="[512,635,1457,1481]" italics="true" pageId="9" pageNumber="290">A. remiger</emphasis>
|
||
Laidlaw, 1912
|
||
</taxonomicName>
|
||
. Some populations of
|
||
<taxonomicName id="4C7F4D3C9809FFABFBDAFA64FB2BFA0C" authorityName="Lieftinck" authorityYear="1940" box="[1075,1178,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="annae">
|
||
<emphasis id="B90BEAAD9809FFABFBDAFA64FB2BFA0C" box="[1075,1178,1457,1481]" italics="true" pageId="9" pageNumber="290">A. annae</emphasis>
|
||
</taxonomicName>
|
||
are polymorphic with respect to the presence of the horn, although it is absent in most of them (R. Dow, pers. comm.). In females of the new species this horn, directed up and forward (
|
||
<figureCitation id="13442A3A9809FFABFD56FA2DFCA3F9D4" box="[703,786,1528,1553]" captionStart="FIGURE 6" captionStartId="8.[151,250,1878,1900]" captionTargetBox="[151,1436,193,1827]" captionTargetId="figure@8.[151,1436,193,1856]" captionTargetPageId="8" captionText="FIGURE 6. Amphicnemis valentini, sp. nov., details of the female paratype from the type locality in Cambodia: a—head and thorax, lateral view; b—forewing pterostigma; c—prothorax, lateral view; d—ditto, dorsal view; e—head, frontal view; f—end of abdomen, lateral view. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1283993/files/figure.png" pageId="9" pageNumber="290">Fig. 6c</figureCitation>
|
||
), is also most similar to that of
|
||
<taxonomicName id="4C7F4D3C9809FFABFB68FA2CFAB7F9D4" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[1153,1286,1528,1553]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="billitonis">
|
||
<emphasis id="B90BEAAD9809FFABFB68FA2CFAB7F9D4" box="[1153,1286,1528,1553]" italics="true" pageId="9" pageNumber="290">A. billitonis</emphasis>
|
||
</taxonomicName>
|
||
, as in males. Identification of
|
||
<taxonomicName id="4C7F4D3C9809FFABFEBBF9C9FE58F9F0" authorityName="Selys" authorityYear="1863" box="[338,489,1564,1589]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD9809FFABFEBBF9C9FE58F9F0" box="[338,489,1564,1589]" italics="true" pageId="9" pageNumber="290">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
females is mostly based on the prothorax structure (
|
||
<bibRefCitation id="EFEE4B4E9809FFABFBC2F9C9FB64F9F0" box="[1067,1237,1564,1589]" pageId="9" pageNumber="290" refId="ref7100" refString="Lieftinck, M. A. (1940) Descriptions and records of South-east Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article">Lieftinck 1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B4E9809FFABFB09F9C8FA24F9F0" box="[1248,1429,1564,1589]" pageId="9" pageNumber="290" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD9809FFABFAF4F9CBFAE5F9F0" box="[1309,1364,1564,1589]" italics="true" pageId="9" pageNumber="290">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) but we cannot differentiate the females of the new species from
|
||
<taxonomicName id="4C7F4D3C9809FFABFC8DF994FC56F99C" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[868,999,1600,1625]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="billitonis">
|
||
<emphasis id="B90BEAAD9809FFABFC8DF994FC56F99C" box="[868,999,1600,1625]" italics="true" pageId="9" pageNumber="290">A. billitonis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9809FFABFF2EF9B1FCA4F890" blockId="9.[151,1437,1024,2021]" pageId="9" pageNumber="290">
|
||
<emphasis id="B90BEAAD9809FFABFF2EF9B1FE9BF9B8" bold="true" box="[199,298,1636,1661]" pageId="9" pageNumber="290">Habitat.</emphasis>
|
||
In
|
||
<collectingCountry id="F368762F9809FFABFEB9F9B1FE77F9B8" box="[336,454,1636,1661]" name="Cambodia" pageId="9" pageNumber="290">Cambodia</collectingCountry>
|
||
, individuals of the new species were flushed from herbs on shady ground bluffs
|
||
<quantity id="4C879B5A9809FFABFABCF9B0FA2DF9B9" box="[1365,1436,1637,1661]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="290" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
||
high at the banks of a small and shallow permanent nameless (?) rivulet with a dark bottom (
|
||
<figureCitation id="13442A3A9809FFABFB0DF95DFA9DF964" box="[1252,1324,1672,1697]" captionStart="FIGURE 9" captionStartId="11.[151,250,1956,1978]" captionTargetBox="[175,1411,283,1923]" captionTargetId="figure@11.[175,1411,283,1923]" captionTargetPageId="11" captionText="FIGURE 9. The type locality of Amphicnemis valentini, sp. nov., a small rivulet in lowland evergreen forest near the coast of Ream Peninsula in Preah Sihanouk Province of Cambodia, 5 iii 2017. Photo by O.K." httpUri="https://zenodo.org/record/1283999/files/figure.png" pageId="9" pageNumber="290">Fig. 9</figureCitation>
|
||
) crossing lowland evergreen forest on a coastal flatland
|
||
<quantity id="4C879B5A9809FFABFD72F978FD58F900" box="[667,745,1708,1733]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="290" unit="km" value="1.7">1.7 km</quantity>
|
||
from its fall to the sea. The individuals hung from leaf tips or perched on twigs over a muddy ground or at moist ground bluffs of rivulet banks. No individual of this species was found during earlier visits to the same locality on
|
||
<date id="FFC1107F9809FFABFD2BF920FC80F8C8" box="[706,817,1781,1805]" pageId="9" pageNumber="290" value="2013-05-27">27 v 2013</date>
|
||
(Kosterin 2014) and
|
||
<date id="FFC1107F9809FFABFBC9F920FB28F8C8" box="[1056,1177,1780,1805]" pageId="9" pageNumber="290" value="2015-03-27">27 iii 2015</date>
|
||
(
|
||
<bibRefCitation id="EFEE4B4E9809FFABFB4EF921FAE8F8C8" box="[1191,1369,1780,1805]" pageId="9" pageNumber="290" refId="ref6888" refString="Kosterin, O. E. (2015 a) Dry season Odonata of the Cardamonean coast (Cambodia and Thailand) revisited. International Dragonfly Fund Report, 89, 1 - 36." type="journal article">Kosterin 2015a</bibRefCitation>
|
||
); note the latter date to be in the same season as when the new species was found (
|
||
<date id="FFC1107F9809FFABFBF7F8CDFB03F8F4" box="[1054,1202,1816,1841]" pageId="9" pageNumber="290" value="2017-03-04" valueMax="2017-03-05" valueMin="2017-03-04">4–5 iii 2017</date>
|
||
). For other
|
||
<taxonomicName id="4C7F4D3C9809FFABFAD4F8CDFA2DF8F4" authorityName="Fabricius" authorityYear="1793" box="[1341,1436,1816,1841]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="290" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
species found in this locality see Kosterin (2014, 2015a).
|
||
</paragraph>
|
||
<paragraph id="8BC036BF9809FFABFF2EF8B4FDCBF820" blockId="9.[151,1437,1024,2021]" pageId="9" pageNumber="290">
|
||
On Phú Quốc Island of
|
||
<collectingCountry id="F368762F9809FFABFE34F8B5FD8EF8BC" box="[477,575,1888,1913]" name="Vietnam" pageId="9" pageNumber="290">Vietnam</collectingCountry>
|
||
, the species was commonly observed in
|
||
<date id="FFC1107F9809FFABFBFFF8B4FB18F8BC" box="[1046,1193,1888,1913]" pageId="9" pageNumber="290" value="2016-08">
|
||
August
|
||
<quantity id="4C879B5A9809FFABFB98F8B5FB18F8BC" box="[1137,1193,1888,1913]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="9" pageNumber="290" unit="in" value="2016.0">2016</quantity>
|
||
</date>
|
||
in, likely year-round wet, swamp forest. Individuals of both sexes perched on leaves and twigs on the ground, or hung from the tips of nearby leaves. Towards dusk, they moved higher up into the vegetation. Although the same habitat was surveyed in the winter, no individuals could be located.
|
||
</paragraph>
|
||
<caption id="DF006637980AFFA8FF7EFAEEFB54FAB5" httpUri="https://zenodo.org/record/1283997/files/figure.png" pageId="10" pageNumber="291" startId="10.[151,250,1339,1361]" targetBox="[159,1423,193,1315]" targetPageId="10">
|
||
<paragraph id="8BC036BF980AFFA8FF7EFAEEFB54FAB5" blockId="10.[151,1436,1339,1392]" pageId="10" pageNumber="291">
|
||
<emphasis id="B90BEAAD980AFFA8FF7EFAEEFEA2FA95" bold="true" box="[151,275,1339,1361]" pageId="10" pageNumber="291">FIGURE 8.</emphasis>
|
||
Details of paratypes of
|
||
<taxonomicName id="4C7F4D3C980AFFA8FDE0FAEEFD40FA94" authority="Kosterin & Kompier, 2018" authorityName="Kosterin & Kompier" authorityYear="2018" box="[521,753,1339,1361]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="valentini" status="sp. nov.">
|
||
<emphasis id="B90BEAAD980AFFA8FDE0FAEEFD40FA94" box="[521,753,1339,1361]" italics="true" pageId="10" pageNumber="291">Amphicnemis valentini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23857D6980AFFA8FD17FAEEFCE1FA95" box="[766,848,1339,1360]" pageId="10" pageNumber="291" rank="species">
|
||
<emphasis id="B90BEAAD980AFFA8FD17FAEEFCE1FA95" bold="true" box="[766,848,1339,1360]" pageId="10" pageNumber="291">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Phú Quốc Island, 20 Viii 2016: a—head and thorax of an immature female; b—anal appendages of a fresh male, lateral view; c—
|
||
<emphasis id="B90BEAAD980AFFA8FC40FA8FFC66FAB5" box="[937,983,1370,1392]" italics="true" pageId="10" pageNumber="291">ditto</emphasis>
|
||
, dorsal view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036BF980AFFA8FF2EFA49FCDAF9E4" blockId="10.[151,1437,1436,2037]" pageId="10" pageNumber="291">
|
||
<taxonomicName id="4C7F4D3C980AFFA8FF2EFA49FEEFFA70" authorityName="Selys" authorityYear="1863" box="[199,350,1436,1461]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD980AFFA8FF2EFA49FEEFFA70" box="[199,350,1436,1461]" italics="true" pageId="10" pageNumber="291">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
species are known to usually inhabit swampy forests (
|
||
<bibRefCitation id="EFEE4B4E980AFFA8FC0DFA48FB16FA70" box="[996,1191,1436,1461]" pageId="10" pageNumber="291" refId="ref6511" refString="Dow, R. A., Choong C. E. & Ng, Y. F. (2010) A review of the genus Amphicnemis in Peninsular Malaysia and Singapore, with descriptions of two new species (Odonata: Zygoptera: Coenagrionidae). Zootaxa, 2605, 45 - 55. https: // doi. org / 10.5281 / zenodo. 197652" type="journal article">
|
||
Dow
|
||
<emphasis id="B90BEAAD980AFFA8FBCEFA48FBD2FA70" box="[1063,1123,1436,1461]" italics="true" pageId="10" pageNumber="291">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) and so did the new species on Phú Quốc. At the same time, although its Cambodian
|
||
<typeStatus id="54C4881D980AFFA8FC6CFA14FC04FA1C" box="[901,949,1473,1497]" pageId="10" pageNumber="291">type</typeStatus>
|
||
locality was in a moist lowland evergreen forest, it was well drained by the rivulet. There were areas of swampy forest elsewhere in this forest, e.g. as closely as
|
||
<quantity id="4C879B5A980AFFA8FF5CF9DDFEB2F9E4" box="[181,259,1544,1569]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="291" unit="km" value="0.6">0.6 km</quantity>
|
||
to the northeast, but no
|
||
<taxonomicName id="4C7F4D3C980AFFA8FDFBF9DDFD18F9E4" authorityName="Selys" authorityYear="1863" box="[530,681,1544,1569]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAAD980AFFA8FDFBF9DDFD18F9E4" box="[530,681,1544,1569]" italics="true" pageId="10" pageNumber="291">Amphicnemis</emphasis>
|
||
</taxonomicName>
|
||
was found there.
|
||
</paragraph>
|
||
<paragraph id="8BC036BF980AFFA8FF2EF9F8FB35F910" blockId="10.[151,1437,1436,2037]" pageId="10" pageNumber="291">
|
||
In
|
||
<collectingRegion id="49BBF85D980AFFA8FF01F9F9FE61F980" box="[232,464,1580,1605]" country="Cambodia" name="Koh Kong" pageId="10" pageNumber="291">Koh Kong Province</collectingRegion>
|
||
of
|
||
<collectingCountry id="F368762F980AFFA8FE12F9F9FDC0F980" box="[507,625,1580,1605]" name="Cambodia" pageId="10" pageNumber="291">Cambodia</collectingCountry>
|
||
, G. Chartier (pers. comm.) photographed an immature (red) female at thickets, which included
|
||
<taxonomicName id="4C7F4D3C980AFFA8FE46F985FC8CF9AD" authority="(Forssk.) Kunze" authorityName="(Forssk.) Kunze" box="[431,829,1616,1641]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="10" pageNumber="291" phylum="Tracheophyta" rank="species" species="odorifer">
|
||
<emphasis id="B90BEAAD980AFFA8FE46F985FD33F9AC" box="[431,642,1616,1641]" italics="true" pageId="10" pageNumber="291">Pandanus odorifer</emphasis>
|
||
(Forssk.) Kunze
|
||
</taxonomicName>
|
||
and some mangroves, overhanging the lower reach of a small river (a tributary of the large Tatai River estuary), at sea level, with brackish water under tidal influence. This stream crossed a marshy coastal flatland transitory to mangroves, with abundant spikerush (
|
||
<taxonomicName id="4C7F4D3C980AFFA8FB19F94DFAD8F974" box="[1264,1385,1688,1713]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="10" pageNumber="291" phylum="Liliopsida" rank="genus">
|
||
<emphasis id="B90BEAAD980AFFA8FB19F94DFAD8F974" box="[1264,1385,1688,1713]" italics="true" pageId="10" pageNumber="291">Eleocharis</emphasis>
|
||
</taxonomicName>
|
||
sp.) and stands of
|
||
<taxonomicName id="4C7F4D3C980AFFA8FEDBF969FDD1F910" authority="Powell" authorityName="Powell" authorityYear="1809" box="[306,608,1724,1749]" class="Insecta" family="Tachinidae" genus="Melaleuca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="cajuputi">
|
||
<emphasis id="B90BEAAD980AFFA8FEDBF969FDBAF911" box="[306,523,1724,1749]" italics="true" pageId="10" pageNumber="291">Melaleuca cajuputi</emphasis>
|
||
Powell
|
||
</taxonomicName>
|
||
and a submangrove palm
|
||
<taxonomicName id="4C7F4D3C980AFFA8FC6FF969FB35F910" authority="Roxb." authorityName="Roxb." box="[902,1156,1724,1749]" class="Liliopsida" family="Arecaceae" genus="Licuala" higherTaxonomySource="GBIF" kingdom="Plantae" order="Arecales" pageId="10" pageNumber="291" phylum="Tracheophyta" rank="species" species="spinosa">
|
||
<emphasis id="B90BEAAD980AFFA8FC6FF969FB89F911" box="[902,1080,1724,1749]" italics="true" pageId="10" pageNumber="291">Licuala spinosa</emphasis>
|
||
Roxb.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656534980AFFA9FF2EF935FE56FF10" lastPageId="11" lastPageNumber="292" pageId="10" pageNumber="291" type="distribution">
|
||
<paragraph id="8BC036BF980AFFA9FF2EF935FE56FF10" blockId="10.[151,1437,1436,2037]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="292" pageId="10" pageNumber="291">
|
||
<emphasis id="B90BEAAD980AFFA8FF2EF935FED1F93C" bold="true" box="[199,352,1760,1785]" pageId="10" pageNumber="291">Distribution.</emphasis>
|
||
The species is reliably known from two localities, on Ream Peninsula, which is a part of the larger Kampongsaom Peninsula, in
|
||
<collectingCountry id="F368762F980AFFA8FDCFF8D1FD28F8D8" box="[550,665,1796,1821]" name="Cambodia" pageId="10" pageNumber="291">Cambodia</collectingCountry>
|
||
and from Phú Quốc
|
||
<collectingCountry id="F368762F980AFFA8FC6AF8D1FC76F8D8" box="[899,967,1796,1821]" name="Iceland" pageId="10" pageNumber="291">Island</collectingCountry>
|
||
in
|
||
<collectingCountry id="F368762F980AFFA8FC02F8D1FBFFF8D8" box="[1003,1102,1796,1821]" name="Vietnam" pageId="10" pageNumber="291">Vietnam</collectingCountry>
|
||
. Although located in different countries and separated by sea, these localities are just
|
||
<quantity id="4C879B5A980AFFA8FCE4F8FCFCE7F884" box="[781,854,1832,1857]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="291" unit="km" value="40.0">40 km</quantity>
|
||
apart. Two smaller Cambodian islands, Thmei and Sês, lie between Ream Peninsula and Phú Quốc
|
||
<collectingCountry id="F368762F980AFFA8FD3CF899FCACF8A0" box="[725,797,1868,1893]" name="Iceland" pageId="10" pageNumber="291">Island</collectingCountry>
|
||
. So, in addition to the rather short distance, these two localities are also separated by three narrow straits, 0.4, 1.4 and
|
||
<quantity id="4C879B5A980AFFA8FC8EF8A4FC04F84C" box="[871,949,1904,1929]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="291" unit="km" value="4.3">4.3 km</quantity>
|
||
wide at their narrowest points. The species can thus be expected to also inhabit at least the two intermediate islands. The immature female with a prothoracic horn photographed in
|
||
<collectingRegion id="49BBF85D980AFFA8FE78F86DFDC2F814" box="[401,627,1976,2001]" country="Cambodia" name="Koh Kong" pageId="10" pageNumber="291">Koh Kong Province</collectingRegion>
|
||
could not be positively identified, but is unlikely not to be this species. Its location was
|
||
<quantity id="4C879B5A980AFFA8FEB1F808FE01F830" box="[344,432,2012,2037]" metricMagnitude="5" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="291" unit="km" value="127.0">127 km</quantity>
|
||
NW from the
|
||
<typeStatus id="54C4881D980AFFA8FD89F808FD21F830" box="[608,656,2013,2037]" pageId="10" pageNumber="291">type</typeStatus>
|
||
locality, separated by the large Botum Sakor Peninsula. Hence the species may be widespread in lowland evergreen forests and even in forests transitory to mangroves in the southeastern Cardamom ecoregion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |