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<mods:title id="9B0622538615E8A277A12F2145C827B4">An annotated and illustrated checklist of the porcelain crabs of Panama (Decapoda: Anomura)</mods:title>
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<paragraph id="8BC09D7D775AFFFE4EF95AF6FD82AA9A" blockId="57.[151,557,1231,1258]" box="[151,557,1231,1258]" pageId="57" pageNumber="58">
<heading id="D0882A11775AFFFE4EF95AF6FD82AA9A" bold="true" box="[151,557,1231,1258]" fontSize="11" level="2" pageId="57" pageNumber="58" reason="4">
<taxonomicName id="4C7FE6FE775AFFFE4EF95AF6FD82AA9A" ID-CoL="26cd09f2-6c53-4a83-8e97-8c2a3b626450" authority="Haig, 1960" authorityName="Haig" authorityYear="1960" box="[151,557,1231,1258]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4EF95AF6FD82AA9A" bold="true" box="[151,557,1231,1258]" pageId="57" pageNumber="58">
<emphasis id="B90B416F775AFFFE4EF95AF6FE0FAA99" bold="true" box="[151,416,1231,1258]" italics="true" pageId="57" pageNumber="58">Pachycheles trichotus</emphasis>
<bibRefCitation id="EFEEE08C775AFFFE4FC95AF6FD82AA9A" author="Haig, J." box="[423,557,1231,1257]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig, 1960</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BC09D7D775AFFFE4EF95B2AFBFBAB5F" blockId="57.[151,1108,1299,1324]" box="[151,1108,1299,1324]" pageId="57" pageNumber="58">
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<treatmentCitation id="0ADEBB6C775AFFFE4EF95B2AFDB3AB58" author="Haig, J." box="[151,540,1299,1323]" page="157" pageId="57" pageNumber="58" year="1960">
<taxonomicName id="4C7FE6FE775AFFFE4EF95B2AFDB3AB58" ID-CoL="26cd09f2-6c53-4a83-8e97-8c2a3b626450" authority="Haig 1960: 157" authorityName="Haig" authorityPageNumber="157" authorityYear="1960" box="[151,540,1299,1323]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4EF95B2AFED9AB58" box="[151,374,1299,1323]" italics="true" pageId="57" pageNumber="58">Pachycheles trichotus</emphasis>
<bibRefCitation id="EFEEE08C775AFFFE4F135B2AFDB3AB58" author="Haig, J." box="[381,540,1299,1323]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960: 157</bibRefCitation>
</taxonomicName>
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, pl. 12, pl. 32, fig. 3;
<treatmentCitation id="0ADEBB6C775AFFFE4C985B2AFC26AB58" author="Haig, J." box="[758,905,1299,1323]" page="57" pageId="57" pageNumber="58" year="1968">
<bibRefCitation id="EFEEE08C775AFFFE4C985B2AFC26AB58" author="Haig, J." box="[758,905,1299,1323]" pageId="57" pageNumber="58" pagination="57 - 74" refId="ref72890" refString="Haig, J. (1968) Eastern Pacific expeditions of the New York Zoological Society. Porcellanid crabs (Crustacea: Anomura) from the west coast of tropical America. Zoologica, New York, 53 (2), 57 - 74." type="journal article" year="1968">Haig 1968: 57</bibRefCitation>
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, 69;
<treatmentCitation id="0ADEBB6C775AFFFE4DD55B2DFBFFAB5F" author="Gore, R. H." box="[955,1104,1300,1324]" page="10" pageId="57" pageNumber="58" year="1982">
<bibRefCitation id="EFEEE08C775AFFFE4DD55B2DFBFFAB5F" author="Gore, R. H." box="[955,1104,1300,1324]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore 1982: 10</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B90B416F775AFFFE4EF95B60FED5AB00" bold="true" box="[151,378,1369,1395]" pageId="57" pageNumber="58">Material examined</emphasis>
.
<materialsCitation id="3B179720775AFFFE4FE65B63FD28ABCE" collectingDate="2015-04-20" collectionCode="MZUSP" collectorName="A. Anker &amp; Anker, J. F &amp; T. Kaji" country="Panama" location="Rio Mar" pageId="57" pageNumber="58" specimenCode="MZUSP 40072" specimenCount="2" specimenCount-male="2">
<collectingCountry id="F368DDED775AFFFE4FE65B63FE4EAB07" box="[392,481,1370,1396]" name="Panama" pageId="57" pageNumber="58">Panama</collectingCountry>
[Pacific]:
<specimenCount id="9D7956F4775AFFFE4C365B63FD1EAB07" box="[600,689,1370,1396]" pageId="57" pageNumber="58" type="male">2 males</specimenCount>
, cl 4.7, cw 5.0 and cl 4.5, cw 4.8, 2 ov. females, cl 5.2, cw 5.6 and cl 3.9, cw 4.3 (
<specimenCode id="DBD93506775AFFFE4F555B47FE45ABEB" box="[315,490,1406,1432]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="57" pageNumber="58">MZUSP 40072</specimenCode>
),
<location id="8EA0CBA6775AFFFE4F935B47FDF2ABEB" LSID="urn:lsid:plazi:treatment:03D62C6B775AFFFE4EF95AF6FED5A9E2:8EA0CBA6775AFFFE4F935B47FDF2ABEB" box="[509,605,1406,1432]" country="Panama" name="Rio Mar" pageId="57" pageNumber="58">Río Mar</location>
, rocky intertidal, low tide, under rocks and in rock crevices, leg.
<collectorName id="268AF8AB775AFFFE4B5E5B47FA37ABEB" box="[1328,1432,1406,1432]" pageId="57" pageNumber="58">A. Anker</collectorName>
, J.F. Lazarus-Agudelo &amp;
<collectorName id="268AF8AB775AFFFE4FC45B9BFE55ABCF" box="[426,506,1442,1468]" pageId="57" pageNumber="58">T. Kaji</collectorName>
,
<date id="FFC1BBBD775AFFFE4C6B5B9BFD28ABCE" box="[517,647,1442,1469]" pageId="57" pageNumber="58" value="2015-04-20">
<collectingDate id="EF854255775AFFFE4C6B5B9BFD28ABCE" box="[517,647,1442,1469]" pageId="57" pageNumber="58" value="2015-04-20">20.04.2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC09D7D775AFFFE4EA95BFCFCDFAB93" blockId="57.[151,1436,1369,1937]" box="[199,880,1477,1505]" pageId="57" pageNumber="58">
<emphasis id="B90B416F775AFFFE4EA95BFCFD95AB93" bold="true" box="[199,570,1477,1504]" pageId="57" pageNumber="58">
Previous records from
<collectingCountry id="F368DDED775AFFFE4FB65BFFFD95AB93" box="[472,570,1478,1504]" name="Panama" pageId="57" pageNumber="58">Panama</collectingCountry>
</emphasis>
.
<bibRefCitation id="EFEEE08C775AFFFE4C265BFFFD7CAB92" author="Haig, J." box="[584,723,1478,1505]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
;
<bibRefCitation id="EFEEE08C775AFFFE4CB15BFFFCC3AB93" author="Gore, R. H." box="[735,876,1478,1504]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore (1982)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365CEF6775AFFFE4EA95BD0FB9FA85A" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BC09D7D775AFFFE4EA95BD0FB9FA85A" blockId="57.[151,1436,1369,1937]" pageId="57" pageNumber="58">
<emphasis id="B90B416F775AFFFE4EA95BD0FEF6A870" bold="true" box="[199,345,1513,1539]" pageId="57" pageNumber="58">Distribution</emphasis>
. East Pacific:
<collectingCountry id="F368DDED775AFFFE4F935BD3FD2DA877" box="[509,642,1514,1540]" name="El Salvador" pageId="57" pageNumber="58">El Salvador</collectingCountry>
,
<collectingCountry id="F368DDED775AFFFE4CE05BD3FCAEA877" box="[654,769,1514,1540]" name="Nicaragua" pageId="57" pageNumber="58">Nicaragua</collectingCountry>
and
<collectingCountry id="F368DDED775AFFFE4D565BD3FC3EA877" box="[824,913,1514,1540]" name="Panama" pageId="57" pageNumber="58">Panama</collectingCountry>
(
<collectingCountry id="F368DDED775AFFFE4DCE5BD3FC65A877" box="[928,970,1514,1540]" name="Iceland" pageId="57" pageNumber="58">Isla</collectingCountry>
Verde inside Golfo de Montijo, Río Mar,
<collectingCountry id="F368DDED775AFFFE4EF95837FF10A85B" box="[151,191,1550,1576]" name="Iceland" pageId="57" pageNumber="58">Isla</collectingCountry>
Venao, vicinity of Ciudad de Panamá) (
<bibRefCitation id="EFEEE08C775AFFFE4C115837FD56A85B" author="Haig, J." box="[639,761,1550,1576]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960</bibRefCitation>
;
<bibRefCitation id="EFEEE08C775AFFFE4D6B5837FCD0A85B" author="Gore, R. H." box="[773,895,1550,1576]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore 1982</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF6775AFFFE4EA9580BFB6CA83E" box="[199,1219,1586,1613]" pageId="57" pageNumber="58" type="biology_ecology">
<paragraph id="8BC09D7D775AFFFE4EA9580BFB6CA83E" blockId="57.[151,1436,1369,1937]" box="[199,1219,1586,1613]" pageId="57" pageNumber="58">
<emphasis id="B90B416F775AFFFE4EA9580BFE89A83F" bold="true" box="[199,294,1586,1612]" pageId="57" pageNumber="58">Ecology</emphasis>
. Intertidal; under rocks or in rock crevices (
<bibRefCitation id="EFEEE08C775AFFFE4D7C580BFC23A83F" author="Haig, J." box="[786,908,1586,1612]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960</bibRefCitation>
;
<bibRefCitation id="EFEEE08C775AFFFE4DF6580BFBBDA83F" author="Gore, R. H." box="[920,1042,1586,1612]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore 1982</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF6775AFFFE4EA9586FFED5A9E2" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BC09D7D775AFFFE4EA9586FFED5A9E2" blockId="57.[151,1436,1369,1937]" pageId="57" pageNumber="58">
<emphasis id="B90B416F775AFFFE4EA9586FFE9BA803" bold="true" box="[199,308,1622,1648]" pageId="57" pageNumber="58">Remarks</emphasis>
. According to
<bibRefCitation id="EFEEE08C775AFFFE4F8F586FFDC0A803" author="Gore, R. H." box="[481,623,1622,1648]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore (1982)</bibRefCitation>
,
<taxonomicName id="4C7FE6FE775AFFFE4C13586EFD50A803" authorityName="Haig" authorityYear="1960" box="[637,767,1622,1648]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4C13586EFD50A803" box="[637,767,1622,1648]" italics="true" pageId="57" pageNumber="58">P. trichotus</emphasis>
</taxonomicName>
is an extremely rare species, previously known from only
<specimenCount id="9D7956F4775AFFFE4EF95843FEEAA8E7" box="[151,325,1658,1684]" pageId="57" pageNumber="58" type="generic">four specimens</specimenCount>
, in addition to
<specimenCount id="9D7956F4775AFFFE4F835843FD3CA8E7" box="[493,659,1658,1684]" pageId="57" pageNumber="58" type="generic">two specimens</specimenCount>
examined by this author.
<bibRefCitation id="EFEEE08C775AFFFE4DDB5843FB91A8E6" author="Haig, J." box="[949,1086,1658,1685]" pageId="57" pageNumber="58" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
separated
<taxonomicName id="4C7FE6FE775AFFFE4AD85842FA99A8E7" authorityName="Haig" authorityYear="1960" box="[1206,1334,1658,1684]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4AD85842FA99A8E7" box="[1206,1334,1658,1684]" italics="true" pageId="57" pageNumber="58">P. trichotus</emphasis>
</taxonomicName>
from the very similar
<taxonomicName id="4C7FE6FE775AFFFE4F4B58A6FE6AA8CB" authorityName="Faxon" authorityYear="1893" box="[293,453,1695,1720]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90B416F775AFFFE4F4B58A6FE6AA8CB" box="[293,453,1695,1720]" italics="true" pageId="57" pageNumber="58">P. panamensis</emphasis>
</taxonomicName>
by the shape of the P1 fingers; the presence of plumose setae on the carapace front and P1P4; and the absence of pleopods in males. However,
<bibRefCitation id="EFEEE08C775AFFFE4D6658FBFC3FA8AF" author="Gore, R. H." box="[776,912,1730,1756]" pageId="57" pageNumber="58" pagination="1 - 34" refId="ref72204" refString="Gore, R. H. (1982) Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contributions to Zoology, 363, 1 - 34. https: // doi. org / 10.5479 / si. 00810282.363" type="journal article" year="1982">Gore (1982)</bibRefCitation>
separated
<taxonomicName id="4C7FE6FE775AFFFE4A6858FAFB2BA8AF" authorityName="Haig" authorityYear="1960" box="[1030,1156,1730,1756]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4A6858FAFB2BA8AF" box="[1030,1156,1730,1756]" italics="true" pageId="57" pageNumber="58">P. trichotus</emphasis>
</taxonomicName>
from both
<taxonomicName id="4C7FE6FE775AFFFE4A9058FAFA33A8AF" authorityName="Faxon" authorityYear="1893" box="[1278,1436,1731,1756]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90B416F775AFFFE4A9058FAFA33A8AF" box="[1278,1436,1731,1756]" italics="true" pageId="57" pageNumber="58">P. panamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7FE6FE775AFFFE4EA658DEFED6A973" authorityName="Haig" authorityYear="1957" box="[200,377,1766,1792]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="spinidactylus">
<emphasis id="B90B416F775AFFFE4EA658DEFED6A973" box="[200,377,1766,1792]" italics="true" pageId="57" pageNumber="58">P. spinidactylus</emphasis>
</taxonomicName>
by the epimeral plate formula, which the author found to be “both distinctive and consistent, comprising a single large plate in
<taxonomicName id="4C7FE6FE775AFFFE4C795932FD38A957" authorityName="Haig" authorityYear="1960" box="[535,663,1802,1828]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4C795932FD38A957" box="[535,663,1802,1828]" italics="true" pageId="57" pageNumber="58">P. trichotus</emphasis>
</taxonomicName>
, one large and one small plate in
<taxonomicName id="4C7FE6FE775AFFFE4A7B5932FB69A957" authorityName="Haig" authorityYear="1957" box="[1045,1222,1802,1828]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="spinidactylus">
<emphasis id="B90B416F775AFFFE4A7B5932FB69A957" box="[1045,1222,1802,1828]" italics="true" pageId="57" pageNumber="58">P. spinidactylus</emphasis>
</taxonomicName>
, and one large and several smaller fragments in
<taxonomicName id="4C7FE6FE775AFFFE4FB35916FDD1A93B" authorityName="Faxon" authorityYear="1893" box="[477,638,1839,1864]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90B416F775AFFFE4FB35916FDD1A93B" box="[477,638,1839,1864]" italics="true" pageId="57" pageNumber="58">P. panamensis</emphasis>
</taxonomicName>
”. Unfortunately, the
<specimenCount id="9D7956F4775AFFFE4D035917FBBCA93B" box="[877,1043,1838,1864]" pageId="57" pageNumber="58" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C7FE6FE775AFFFE4A545916FB15A93B" authorityName="Haig" authorityYear="1960" box="[1082,1210,1838,1864]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trichotus">
<emphasis id="B90B416F775AFFFE4A545916FB15A93B" box="[1082,1210,1838,1864]" italics="true" pageId="57" pageNumber="58">P. trichotus</emphasis>
</taxonomicName>
from Río Mar were not photographed alive, likely due to the species resemblance with
<taxonomicName id="4C7FE6FE775AFFFE4DCC596AFBEBA91F" authorityName="Faxon" authorityYear="1893" box="[930,1092,1875,1900]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90B416F775AFFFE4DCC596AFBEBA91F" box="[930,1092,1875,1900]" italics="true" pageId="57" pageNumber="58">P. panamensis</emphasis>
</taxonomicName>
, which was also found at this locality (see above).
</paragraph>
</subSubSection>
</treatment>
</document>