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<document id="F3E3FD200E00AE24AFF09E8E7A2B6D11" ID-CLB-Dataset="45869" ID-DOI="10.11646/zootaxa.3794.2.2" ID-GBIF-Dataset="4dab3d72-ed90-4061-959c-1a39618b69ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="230988" ID-ZooBank="E75F9E63-4785-49AB-AD6D-AC4838F39BAD" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460415592207" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="03D3CE22E3795850D4FB37A655542FF7" docLanguage="en" docName="zt03794p236.pdf" docOrigin="Zootaxa 3794 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maorinus" docType="treatment" docVersion="8" lastPageNumber="231" masterDocId="FFEAB65AE3715859D46C345E504E2E02" masterDocTitle="Comparative morphology and taxonomy of Sciacharis s. str., Chathamaenus Franz and Maorinus Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="236" masterPageNumber="222" pageNumber="230" updateTime="1698328690967" updateUser="ExternalLinkService">
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<mods:title id="C21A7F8F69B11838E3B792B657AFA54A">Comparative morphology and taxonomy of Sciacharis s. str., Chathamaenus Franz and Maorinus Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="B28C90E4BA9AA495E8F149986D1C0882" type="personal">
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<mods:namePart id="1F54C25FE3952B005CFB55007862558C">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="5CC47E445E701711673EB841FF0FBE53">2014</mods:date>
<mods:detail id="79788F2E359B4CC33AE8E9FDB2303661" type="volume">
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<mods:identifier id="D99960218418BC847A62B0F355E7C355" type="DOI">10.11646/zootaxa.3794.2.2</mods:identifier>
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<treatment id="03D3CE22E3795850D4FB37A655542FF7" ID-DOI="http://doi.org/10.5281/zenodo.6140177" ID-GBIF-Taxon="119424888" ID-Zenodo-Dep="6140177" LSID="urn:lsid:plazi:treatment:03D3CE22E3795850D4FB37A655542FF7" httpUri="http://treatment.plazi.org/id/03D3CE22E3795850D4FB37A655542FF7" lastPageId="9" lastPageNumber="231" pageId="8" pageNumber="230">
<subSubSection id="C3602CBFE3795851D4FB37A651AA2A10" box="[151,484,1015,1042]" pageId="8" pageNumber="230" type="nomenclature">
<paragraph id="8BC57F34E3795851D4FB37A651AA2A10" blockId="8.[151,484,1015,1042]" box="[151,484,1015,1042]" pageId="8" pageNumber="230">
<heading id="D08DC858E3795851D4FB37A651AA2A10" bold="true" box="[151,484,1015,1042]" fontSize="11" level="1" pageId="8" pageNumber="230" reason="1">
<emphasis id="B90EA326E3795851D4FB37A651AA2A10" bold="true" box="[151,484,1015,1042]" pageId="8" pageNumber="230">
subgenus
<taxonomicName id="4C7A04B7E3795851D57E37A951C72A13" box="[274,393,1015,1041]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D57E37A951C72A13" bold="true" box="[274,393,1015,1041]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
stat. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3602CBFE3795851D4FB301E51552B64" pageId="8" pageNumber="230" type="reference_group">
<paragraph id="8BC57F34E3795851D4FB301E53AE2A90" blockId="8.[151,1436,1086,1170]" pageId="8" pageNumber="230">
<treatmentCitationGroup id="AB6A581AE3795851D4FB301E53AE2A90" infered="true" pageId="8" pageNumber="230">
<taxonomicName id="4C7A04B7E3795851D4FB301E51F42A56" authority="Franz, 1980a: 291" authorityName="Franz" authorityPageNumber="291" authorityYear="1980" box="[151,442,1086,1109]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D4FB301E50B42A57" box="[151,250,1088,1109]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
<treatmentCitation id="0ADB5925E3795851D56C306151F42A56" author="Franz" box="[256,442,1086,1108]" page="291" pageId="8" pageNumber="230" year="1980">
<bibRefCitation id="EFEB02C5E3795851D56C306151C62A56" author="Franz" box="[256,392,1086,1108]" pageId="8" pageNumber="230" refString="Franz, H. (1980 a) Weiterer Beitrag zur Kenntnis der Scydmaenidenfauna Neuseelands. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 189, 249 - 313." type="journal article" year="1980" yearSuffix="a">Franz, 1980a</bibRefCitation>
: 291
</treatmentCitation>
</taxonomicName>
; replacement name for preoccupied
<taxonomicName id="4C7A04B7E3795851D745301E53F72A56" ID-CoL="8XMLX" authority="Franz" authorityName="Franz" box="[809,953,1087,1109]" class="Insecta" family="Staphylinidae" genus="Maoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D745301E533B2A57" box="[809,885,1088,1109]" italics="true" pageId="8" pageNumber="230">Maoria</emphasis>
Franz
</taxonomicName>
; elevated to the genus rank by
<bibRefCitation id="EFEB02C5E3795851D09D306055DF2A56" author="Kuschel" box="[1265,1425,1086,1108]" pageId="8" pageNumber="230" refString="Kuschel, G. (1990) Beetles in a suburban environment: a New Zealand case study. The identity and status of Coleoptera in the natural and modified habitats of Lynfield, Auckland 1974 - 1989. New Zealand Department of Scientific and Industrial Research. DSIR Plant Protection Report No. 3, 118 pp." type="book chapter" year="1990">Kuschel (1990)</bibRefCitation>
.
<taxonomicName id="4C7A04B7E3795851D4FB300051C02A71" ID-CoL="8XMLX" authority="Franz, 1975: 14" authorityName="Franz" authorityPageNumber="14" authorityYear="1975" box="[151,398,1117,1139]" class="Insecta" family="Staphylinidae" genus="Maoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D4FB300050AD2A71" box="[151,227,1118,1139]" italics="true" pageId="8" pageNumber="230">Maoria</emphasis>
<treatmentCitation id="0ADB5925E3795851D486300051C02A71" author="Franz" box="[234,398,1117,1139]" page="14" pageId="8" pageNumber="230" year="1975">
<bibRefCitation id="EFEB02C5E3795851D486300051262A71" author="Franz" box="[234,360,1117,1139]" pageId="8" pageNumber="230" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
: 14
</treatmentCitation>
</taxonomicName>
; as subgenus of
<taxonomicName id="4C7A04B7E3795851D65B300052D52A71" box="[567,667,1118,1139]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D65B300052D52A71" box="[567,667,1118,1139]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
, preoccupied,
<emphasis id="B90EA326E3795851D75E3000531B2A71" box="[818,853,1118,1139]" italics="true" pageId="8" pageNumber="230">nec</emphasis>
<bibRefCitation id="EFEB02C5E3795851D730300053A02A71" author="Laporte" box="[860,1006,1117,1139]" pageId="8" pageNumber="230" refString="Laporte, F. L. N. (1868) Notes on Australian Coleoptera. Part II, Second Family - Carabidae. Royal Society of Victoria, Melbourne, 139 pp." type="book" year="1868">Laporte, 1868</bibRefCitation>
,
<emphasis id="B90EA326E3795851D795300054522A71" box="[1017,1052,1118,1139]" italics="true" pageId="8" pageNumber="230">nec</emphasis>
<bibRefCitation id="EFEB02C5E3795851D04F300354E22A71" author="Pilsbry" box="[1059,1196,1117,1139]" pageId="8" pageNumber="230" refString="Pilsbry, H. A. (1892) Manual of Conchology, structural and systematic, with illustrations of the species, (2), 8, 87." type="journal article" year="1892">Pilsbry, 1892</bibRefCitation>
,
<emphasis id="B90EA326E3795851D0DB300054942A71" box="[1207,1242,1118,1139]" italics="true" pageId="8" pageNumber="230">nec</emphasis>
<bibRefCitation id="EFEB02C5E3795851D08D300055202A71" author="Warren" box="[1249,1390,1117,1139]" pageId="8" pageNumber="230" refString="Warren, W. (1912) Genus Maoria. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde, 11, pp. 76 - 82." type="journal article" year="1912">Warren, 1912</bibRefCitation>
,
<emphasis id="B90EA326E3795851D115300055D22A71" box="[1401,1436,1118,1139]" italics="true" pageId="8" pageNumber="230">nec</emphasis>
<bibRefCitation id="EFEB02C5E3795851D4AB302251282A90" author="Cameron" box="[199,358,1148,1170]" pageId="8" pageNumber="230" refString="Cameron, M. (1945) Some observations on the Staphylinidae of the Broun Collection of Coleoptera in the British Museum, with descriptions of new genera and species (continued). Annals and Magazine of Natural History, Series 11, 12, 158 - 180. http: // dx. doi. org / 10.1080 / 00222934508527501" type="journal article" year="1945">
<collectingCountry id="F36D3FA4E3795851D4AB302251682A90" box="[199,294,1148,1170]" name="Cameroon" pageId="8" pageNumber="230">Cameron</collectingCountry>
, 1945
</bibRefCitation>
.
<typeStatus id="54C1C196E3795851D503302351EF2A90" box="[367,417,1149,1170]" pageId="8" pageNumber="230">Type</typeStatus>
species:
<taxonomicName id="4C7A04B7E3795851D5923022532B2A90" ID-CoL="777DH" authority="Broun, 1915" authorityName="Broun" authorityYear="1915" box="[510,869,1148,1170]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="species" species="alacer">
<emphasis id="B90EA326E3795851D592302252AF2A90" box="[510,737,1148,1170]" italics="true" pageId="8" pageNumber="230">Phagonophana alacer</emphasis>
<bibRefCitation id="EFEB02C5E3795851D68A3023532B2A90" author="Broun" box="[742,869,1148,1170]" pageId="8" pageNumber="230" refString="Broun, T. (1915) Descriptions of new genera and species of Coleoptera. Part IV. New Zealand Institute Bulletin, 1 (4), 267 - 346." type="journal article" year="1915">Broun, 1915</bibRefCitation>
</taxonomicName>
(des. orig.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC57F34E3795851D4FB30E051552B64" blockId="8.[151,1436,1214,2031]" pageId="8" pageNumber="230">
<emphasis id="B90EA326E3795851D4FB30E051382AD5" bold="true" box="[151,374,1214,1239]" pageId="8" pageNumber="230">Revised diagnosis.</emphasis>
A subgenus of
<taxonomicName id="4C7A04B7E3795851D65530E052E12AD5" ID-CoL="8XPS4" box="[569,687,1214,1239]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D65530E052E12AD5" box="[569,687,1214,1239]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
that differs from
<taxonomicName id="4C7A04B7E3795851D7EB30E053B32AD5" ID-CoL="8XPS4" box="[903,1021,1214,1239]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EA326E3795851D7EB30E053B32AD5" box="[903,1021,1214,1239]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D1E5DE3795851D06530E154072AD4" box="[1033,1097,1215,1238]" pageId="8" pageNumber="230" sensu="stricto">s. str.</taxonomicNameLabel>
in internal (adcoxal) part of prothoracic hypomeron anteriorly demarcated from prosternum and posteriorly fused with external part of hypomeron; and in mesoventral intercoxal process continuous, not interrupted in middle, distinctly visible in ventral view in intact specimens as moderately narrow carina with finely setose ventral surface clearly separating mesocoxae.
</paragraph>
</subSubSection>
<subSubSection id="C3602CBFE3795850D4AB312C55542FF7" lastPageId="9" lastPageNumber="231" pageId="8" pageNumber="230" type="discussion">
<paragraph id="8BC57F34E3795851D4AB312C522528A8" blockId="8.[151,1436,1214,2031]" pageId="8" pageNumber="230">
<emphasis id="B90EA326E3795851D4AB312C517A2B89" bold="true" box="[199,308,1394,1419]" pageId="8" pageNumber="230">Remarks</emphasis>
.
<taxonomicName id="4C7A04B7E3795851D52D312A51E02B89" box="[321,430,1396,1419]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D52D312A51E02B89" box="[321,430,1396,1419]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C7A04B7E3795851D5B0312A527E2B89" box="[476,560,1396,1419]" class="Insecta" family="Staphylinidae" genus="Maoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D5B0312A527E2B89" box="[476,560,1396,1419]" italics="true" pageId="8" pageNumber="230">Maoria</emphasis>
</taxonomicName>
) was described as a subgenus of
<taxonomicName id="4C7A04B7E3795851D7CE312A545E2B89" box="[930,1040,1396,1419]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D7CE312A545E2B89" box="[930,1040,1396,1419]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
characterized by a strongly convex body and usually long and at least in some places dense and erect setation (
<bibRefCitation id="EFEB02C5E3795851D7B331C9542A2BAD" author="Franz" box="[991,1124,1430,1455]" pageId="8" pageNumber="230" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz 1975</bibRefCitation>
). This diagnosis is certainly highly unclear and matches many species of
<taxonomicName id="4C7A04B7E3795851D6DE31E2536E2BD1" box="[690,800,1468,1491]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D6DE31E2536E2BD1" box="[690,800,1468,1491]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
-like
<taxonomicName id="4C7A04B7E3795851D70C31E454482BD1" box="[864,1030,1466,1491]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
from all continents.
<bibRefCitation id="EFEB02C5E3795851D16931E555D22BD1" author="Franz" box="[1285,1436,1466,1491]" pageId="8" pageNumber="230" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz (1975)</bibRefCitation>
compared
<taxonomicName id="4C7A04B7E3795851D561318151342BF4" box="[269,378,1503,1526]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D561318151342BF4" box="[269,378,1503,1526]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C7A04B7E3795851D5C6318151B02BF4" box="[426,510,1503,1526]" class="Insecta" family="Staphylinidae" genus="Maoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D5C6318151B02BF4" box="[426,510,1503,1526]" italics="true" pageId="8" pageNumber="230">Maoria</emphasis>
</taxonomicName>
) to
<taxonomicName id="4C7A04B7E3795851D640318152D42BF4" box="[556,666,1503,1526]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D640318152D42BF4" box="[556,666,1503,1526]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C7A04B7E3795851D77F318054252BF5" authority="Motschulsky, 1870" authorityName="Motschulsky" authorityYear="1870" box="[787,1131,1502,1527]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D77F318053C12BF5" box="[787,911,1502,1527]" italics="true" pageId="8" pageNumber="230">Tetramelus</emphasis>
<bibRefCitation id="EFEB02C5E3795851D7F9318054252BF5" author="Motschulsky" box="[917,1131,1502,1527]" pageId="8" pageNumber="230" refString="Motschulsky, V. (1870) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. 7 - ieme Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42, 252 - 272." type="journal article" year="1870">Motschulsky, 1870</bibRefCitation>
</taxonomicName>
and listed some characters to distinguish these taxa. In my opinion similarities between
<taxonomicName id="4C7A04B7E3795851D721325A53F42819" box="[845,954,1540,1563]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D721325A53F42819" box="[845,954,1540,1563]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A04B7E3795851D798325C543E2819" box="[1012,1136,1538,1563]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D798325C543E2819" box="[1012,1136,1538,1563]" italics="true" pageId="8" pageNumber="230">Tetramelus</emphasis>
</taxonomicName>
are only superficial and it remains unclear why Franz associated the morphology of both genera. It is also unclear why
<bibRefCitation id="EFEB02C5E3795851D086327855D2283D" author="Kuschel" box="[1258,1436,1574,1599]" pageId="8" pageNumber="230" refString="Kuschel, G. (1990) Beetles in a suburban environment: a New Zealand case study. The identity and status of Coleoptera in the natural and modified habitats of Lynfield, Auckland 1974 - 1989. New Zealand Department of Scientific and Industrial Research. DSIR Plant Protection Report No. 3, 118 pp." type="book chapter" year="1990">Kuschel (1990)</bibRefCitation>
elevated
<taxonomicName id="4C7A04B7E3795851D492321251252861" box="[254,363,1612,1635]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D492321251252861" box="[254,363,1612,1635]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
to the genus rank, and why Franz accepted this decision, in opposition to
<taxonomicName id="4C7A04B7E3795851D0A0321455DB2861" box="[1228,1429,1610,1635]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D0A0321455DB2861" box="[1228,1429,1610,1635]" italics="true" pageId="8" pageNumber="230">Magellanoconnus</emphasis>
</taxonomicName>
, which was also treated by
<bibRefCitation id="EFEB02C5E3795851D5A0323052322885" author="Kuschel" box="[460,636,1646,1671]" pageId="8" pageNumber="230" refString="Kuschel, G. (1990) Beetles in a suburban environment: a New Zealand case study. The identity and status of Coleoptera in the natural and modified habitats of Lynfield, Auckland 1974 - 1989. New Zealand Department of Scientific and Industrial Research. DSIR Plant Protection Report No. 3, 118 pp." type="book chapter" year="1990">Kuschel (1990)</bibRefCitation>
as a genus (again, without giving any reasons) but retained by
<bibRefCitation id="EFEB02C5E3795851D1303231509428A9" author="Franz" pageId="8" pageNumber="230" refString="Franz, H. (1993) Nachtrage zur Kenntnis der Scydmaenidenfauna des magellanisch-valdivianischen Faunengebietes in Chile (Coleoptera, Scydmaenidae). Zeitschrift der Arbeitsgemeinchaft Osterreichischer Entomologen, 44, 99 - 108." type="journal article" year="1993">Franz (1993</bibRefCitation>
,
<bibRefCitation id="EFEB02C5E3795851D48A32CD515128A9" author="Franz" box="[230,287,1683,1707]" pageId="8" pageNumber="230" refString="Franz, H. (1997) Weitere von T. Cekalovic in Chile gesammelte Scydmaeniden (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 67, 159 - 168." type="journal article" year="1997">1997</bibRefCitation>
) as a subgenus of
<taxonomicName id="4C7A04B7E3795851D58232CC522A28A9" box="[494,612,1682,1707]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D58232CC522A28A9" box="[494,612,1682,1707]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC57F34E3795851D4AB32E8545B29ED" blockId="8.[151,1436,1214,2031]" pageId="8" pageNumber="230">
The morphology of
<taxonomicName id="4C7A04B7E3795851D5C732E9525628CC" box="[427,536,1719,1742]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D5C732E9525628CC" box="[427,536,1719,1742]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
, although the general appearance (Figs. 4–5) seems distinct from
<taxonomicName id="4C7A04B7E3795851D16832E8553428CD" box="[1284,1402,1718,1743]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D16832E8553428CD" box="[1284,1402,1718,1743]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
, is highly similar to structures of
<taxonomicName id="4C7A04B7E3795851D5833284522B28F1" box="[495,613,1754,1779]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EA326E3795851D5833284522B28F1" box="[495,613,1754,1779]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D1E5DE3795851D601328552E728F0" box="[621,681,1755,1778]" pageId="8" pageNumber="230" sensu="stricto">s. str.</taxonomicNameLabel>
(as shown in
<figureCitation id="134163B1E3795851D7253284539E28F1" box="[841,976,1754,1779]" captionStart="FIGURES 6 – 11" captionStartId="4.[151,264,1516,1539]" captionTargetBox="[159,1427,195,1484]" captionTargetId="figure@4.[151,1436,193,1495]" captionTargetPageId="4" captionText="FIGURES 6 – 11. Head capsule of males in dorsal (6, 8, 10) and ventral (7, 9, 11) views: Sciacharis (s. str.) fulva Broun (6 – 7); Sciacharis (s. str.) chathamensis (Franz), comb. n. (8 – 9); Sciacharis (Maorinus) sp. (10 – 11). Abbreviations: bst, basistipes; cd, cardo; clp, clypeal projection; fo, foramen occipitale; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lbr, labrum; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; oc, occiput; occ, occipital constriction; pp, palpifer; prm, prementum; ptp, posterior tentorial pit; sat, supraantennal tubercle; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/230989/files/figure.png" pageId="8" pageNumber="230">Figs. 10–11</figureCitation>
,
<figureCitation id="134163B1E3795851D7B13284546D28F1" box="[989,1059,1754,1779]" captionStart="FIGURES 12 – 17" captionStartId="5.[151,264,1878,1901]" captionTargetBox="[156,1432,851,1848]" captionTargetId="figure@5.[151,1436,841,1857]" captionTargetPageId="5" captionText="FIGURES 12 – 17. Prothorax in dorsal (12, 14, 16) and ventral (13, 15, 17) views: Sciacharis (s. str.) fulva Broun (12 – 13); Sciacharis (s. str.) chathamensis (Franz), comb. n. (14 – 15); Sciacharis (Maorinus) sp. (16 – 17). Abbreviations: bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; mp, median antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pf, profurca; slc, sublateral carina." httpUri="https://zenodo.org/record/230990/files/figure.png" pageId="8" pageNumber="230">16–17</figureCitation>
and
<figureCitation id="134163B1E3795851D0303285543728F1" box="[1116,1145,1755,1779]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,964,987]" captionTargetBox="[151,1430,196,942]" captionTargetId="figure@6.[151,1436,193,943]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Pterothorax in ventral view: Sciacharis (s. str.) fulva Broun (18); Sciacharis (s. str.) chathamensis (Franz), comb. n. with (19) and without (20) setation; Sciacharis (Maorinus) sp. (21). Abbreviations: ar, anterior ridge; lmfa, lateral mesofurcal arm; mcp, mesocoxal projection; mff, mesofurcal fovea; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pl, posterior lobe; pre, prepectus; si, setose impression; v 3, metaventrite; vlf, ventrolateral foveae." httpUri="https://zenodo.org/record/230991/files/figure.png" pageId="8" pageNumber="230">21</figureCitation>
) and differs in two easily observable details (indicated in
<figureCitation id="134163B1E3795851D67E32A0523C2915" box="[530,626,1790,1815]" captionStart="FIGURES 12 – 17" captionStartId="5.[151,264,1878,1901]" captionTargetBox="[156,1432,851,1848]" captionTargetId="figure@5.[151,1436,841,1857]" captionTargetPageId="5" captionText="FIGURES 12 – 17. Prothorax in dorsal (12, 14, 16) and ventral (13, 15, 17) views: Sciacharis (s. str.) fulva Broun (12 – 13); Sciacharis (s. str.) chathamensis (Franz), comb. n. (14 – 15); Sciacharis (Maorinus) sp. (16 – 17). Abbreviations: bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; lp, lateral antebasal pit; mp, median antebasal pit; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pf, profurca; slc, sublateral carina." httpUri="https://zenodo.org/record/230990/files/figure.png" pageId="8" pageNumber="230">Figs. 17</figureCitation>
and
<figureCitation id="134163B1E3795851D6D832A1529E2915" box="[692,720,1791,1815]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,964,987]" captionTargetBox="[151,1430,196,942]" captionTargetId="figure@6.[151,1436,193,943]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Pterothorax in ventral view: Sciacharis (s. str.) fulva Broun (18); Sciacharis (s. str.) chathamensis (Franz), comb. n. with (19) and without (20) setation; Sciacharis (Maorinus) sp. (21). Abbreviations: ar, anterior ridge; lmfa, lateral mesofurcal arm; mcp, mesocoxal projection; mff, mesofurcal fovea; mscc, mesocoxal cavity; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pl, posterior lobe; pre, prepectus; si, setose impression; v 3, metaventrite; vlf, ventrolateral foveae." httpUri="https://zenodo.org/record/230991/files/figure.png" pageId="8" pageNumber="230">21</figureCitation>
by arrows). The internal (adcoxal) part of each prothoracic hypomeron is anteriorly strongly narrowing and completely demarcated from prosternum (anteriorly indistinctly separated or even fused with prosternum in
<taxonomicName id="4C7A04B7E3795851D6F33318535B295D" box="[671,789,1862,1887]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EA326E3795851D6F33318535B295D" box="[671,789,1862,1887]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D1E5DE3795851D74C3319532E295D" box="[800,864,1863,1887]" pageId="8" pageNumber="230" sensu="stricto">s. str.</taxonomicNameLabel>
) and the posterior parts of hypomeral ridges are obliterated, so that adcoxal and external parts of hypomera are fused posteriorly (demarcated in
<taxonomicName id="4C7A04B7E3795851D0BF333455072981" box="[1235,1353,1898,1923]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EA326E3795851D0BF333455072981" box="[1235,1353,1898,1923]" italics="true" pageId="8" pageNumber="230">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D1E5DE3795851D13D333555C12981" box="[1361,1423,1899,1923]" pageId="8" pageNumber="230" sensu="stricto">s. str.</taxonomicNameLabel>
). The mesoventral intercoxal process in
<taxonomicName id="4C7A04B7E3795851D63B33D1528A29A4" box="[599,708,1935,1958]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D63B33D1528A29A4" box="[599,708,1935,1958]" italics="true" pageId="8" pageNumber="230">Maorinus</emphasis>
</taxonomicName>
is not interrupted in middle and forms a long carina distinctly separating mesocoxae. The carina, in contrast to
<taxonomicName id="4C7A04B7E3795851D68033EA531429C9" box="[748,858,1972,1995]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D68033EA531429C9" box="[748,858,1972,1995]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
, is not expanded ventrally beyond the level of mesocoxae, and the metacoxae are contiguous (broadly separated in
<taxonomicName id="4C7A04B7E3795851D7FB3389544B29EC" box="[919,1029,2007,2030]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3795851D7FB3389544B29EC" box="[919,1029,2007,2030]" italics="true" pageId="8" pageNumber="230">Euconnus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC57F34E3785850D4AB34C952272F66" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="231">
Species of
<taxonomicName id="4C7A04B7E3785850D52E34C751E02EB2" box="[322,430,153,176]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D52E34C751E02EB2" box="[322,430,153,176]" italics="true" pageId="9" pageNumber="231">Maorinus</emphasis>
</taxonomicName>
are usually (but not always) larger, more robust, more convex and darker than
<taxonomicName id="4C7A04B7E3785850D14A34C955D22EB2" box="[1318,1436,151,176]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EA326E3785850D14A34C955D22EB2" box="[1318,1436,151,176]" italics="true" pageId="9" pageNumber="231">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D1E5DE3785850D4FB34E350952ED6" box="[151,219,189,212]" pageId="9" pageNumber="231" sensu="stricto">s. str.</taxonomicNameLabel>
Additionally, the vertex in
<taxonomicName id="4C7A04B7E3785850D65034E352E72ED6" box="[572,681,189,212]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D65034E352E72ED6" box="[572,681,189,212]" italics="true" pageId="9" pageNumber="231">Maorinus</emphasis>
</taxonomicName>
can be slightly expanded dorsocaudad, but not strongly and subconically, as in some species of
<taxonomicName id="4C7A04B7E3785850D64B34BF52DB2EFA" box="[551,661,225,248]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D64B34BF52DB2EFA" box="[551,661,225,248]" italics="true" pageId="9" pageNumber="231">Euconnus</emphasis>
</taxonomicName>
; also the metaventral intercoxal process can be shallowly notched in middle. However, only minor differences in the important structures as the head venter, prosternum and pterothorax were found and they do not justify a separate position of
<taxonomicName id="4C7A04B7E3785850D7CD357754402F42" box="[929,1038,297,320]" class="Insecta" family="Staphylinidae" genus="Maorinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D7CD357754402F42" box="[929,1038,297,320]" italics="true" pageId="9" pageNumber="231">Maorinus</emphasis>
</taxonomicName>
as a genus. Therefore, this taxon is here reduced to a subgenus of
<taxonomicName id="4C7A04B7E3785850D5873512522F2F67" box="[491,609,332,357]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D5873512522F2F67" box="[491,609,332,357]" italics="true" pageId="9" pageNumber="231">Sciacharis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC57F34E3785850D4AB352E55542FF7" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="231">
A preliminary study revealed that at least some Australian,
<collectingCountry id="F36D3FA4E3785850D7E9352E54512F8A" box="[901,1055,367,392]" name="New Zealand" pageId="9" pageNumber="231">New Zealand</collectingCountry>
and New Caledonian species of
<taxonomicName id="4C7A04B7E3785850D4FB35CA51A02FAE" authority="(Magellanoconnus)" baseAuthorityName="Magellanoconnus" box="[151,494,404,429]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D4FB35CA51432FAF" box="[151,269,404,429]" italics="true" pageId="9" pageNumber="231">Sciacharis</emphasis>
(
<emphasis id="B90EA326E3785850D57135CA51A82FAF" box="[285,486,404,429]" italics="true" pageId="9" pageNumber="231">Magellanoconnus</emphasis>
)
</taxonomicName>
do not differ in any important characters from
<taxonomicName id="4C7A04B7E3785850D79535CA54B92FAE" authority="(Maorinus)" baseAuthorityName="Maorinus" box="[1017,1271,404,429]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D79535CA54212FAF" box="[1017,1135,404,429]" italics="true" pageId="9" pageNumber="231">Sciacharis</emphasis>
(
<emphasis id="B90EA326E3785850D01335CB54A22FAE" box="[1151,1260,405,428]" italics="true" pageId="9" pageNumber="231">Maorinus</emphasis>
)
</taxonomicName>
, but also some of them have slender, and not expanded maxillary palpomere III. However,
<taxonomicName id="4C7A04B7E3785850D78535E954FC2FD2" box="[1001,1202,439,464]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B90EA326E3785850D78535E954FC2FD2" box="[1001,1202,439,464]" italics="true" pageId="9" pageNumber="231">Magellanoconnus</emphasis>
</taxonomicName>
is also distributed in South
<collectingCountry id="F36D3FA4E3785850D4B3358251712FF7" box="[223,319,476,501]" name="United States of America" pageId="9" pageNumber="231">America</collectingCountry>
and
<collectingCountry id="F36D3FA4E3785850D51A3582524B2FF7" box="[374,517,476,501]" name="South Africa" pageId="9" pageNumber="231">South Africa</collectingCountry>
and its morphology and taxonomic status requires a separate analysis.
</paragraph>
</subSubSection>
</treatment>
</document>
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