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<document id="356469B26B2895336D46445DEB399BC4" ID-DOI="10.5281/zenodo.5360449" ID-ISSN="0024-4082" ID-Zenodo-Dep="5360449" ID-ZooBank="591555C6-EF37-495B-A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630232463663" checkinUser="marcus" docAuthor="Rajabizadeh, Mahdi, Nagy, Zoltán T., Adriaens, Dominique, Avci, Aziz, Masroor, Rafaqat, Schmidtler, Josef, Nazarov, Roman, Esmaeili, Hamid Reza &amp; Christiaens, Joachim" docDate="2016" docId="03D39A7AFFC19F36FC2BA41214D7683E" docLanguage="en" docName="zoj12342.pdf" docOrigin="Zoological Journal of the Linnean Society 176" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Contia persica Boulenger 1893" docType="treatment" docVersion="3" lastPageNumber="901" masterDocId="FFEAE202FFD79F21FFA5A128115C6F53" masterDocTitle="Alpine-Himalayan orogeny drove correlated morphological, molecular, and ecological diversification in the Persian dwarf snake (Squamata: Serpentes: Eirenis persicus)" masterLastPageNumber="913" masterPageNumber="878" pageNumber="900" updateTime="1699193229270" updateUser="plazi">
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<mods:title id="EE47972163FA6AB4282F05B11C55DF8F">Alpine-Himalayan orogeny drove correlated morphological, molecular, and ecological diversification in the Persian dwarf snake (Squamata: Serpentes: Eirenis persicus)</mods:title>
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<mods:namePart id="1A4EB0D0FC5FDE05FCF9769E75025237">Rajabizadeh, Mahdi</mods:namePart>
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<mods:namePart id="07751BEF4843C6A040C2FF0EE6FD395F">Nagy, Zoltán T.</mods:namePart>
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<mods:namePart id="CB3F751D7EBFC638A25328A9AF1E934E">Adriaens, Dominique</mods:namePart>
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<mods:namePart id="17B6EA7CDE1CD3343CADBC952DA2C051">Avci, Aziz</mods:namePart>
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<mods:namePart id="EA2ABCE19A242A303766E5C62F15CA5B">Masroor, Rafaqat</mods:namePart>
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<mods:date id="A95B0A60ACD1CEA500987ED2C044ECB6">2016</mods:date>
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<taxonomicName id="4C7A50EFFFC19F37FC2BA41215146A1E" authorityName="Boulenger" authorityYear="1893" box="[910,1096,1338,1359]" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="882" phylum="Chordata" rank="species" species="persica">CONTIA PERSICA</taxonomicName>
PERSICA
</emphasis>
<bibRefCitation id="EFEB569DFFC19F37FB10A41215946A3C" author="Terent ev PV &amp; Chernov SA" pageId="22" pageNumber="900" refId="ref17793" refString="Terent ev PV, Chernov SA. 1965. Key to amphibian and reptiles. Jerusalem: Israel Program for Scientific translation [English translation of Terent ev &amp; Chernov, 1949]." type="book" year="1965">TERENT ÉV &amp; CHERNOV, 1965</bibRefCitation>
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<materialsCitation id="3B122131FFC19F37FC99A4B7146C6A80" collectingDate="2005-05-01" collectionCode="ZDEU" collectorName="I. Baran &amp; Y. Kumlutas &amp; C. Ilgaz &amp; A. Avci" pageId="22" pageNumber="900" specimenCode="ZDEU 136" specimenCount="1" typeStatus="holotype">
Adult male,
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/2005, legit.
<collectorName id="268F4EBAFFC19F37FAAEA4B314266AE7" box="[1291,1402,1435,1460]" pageId="22" pageNumber="900">İ. Baran</collectorName>
,
<collectorName id="268F4EBAFFC19F37FA29A4B712F66A80" pageId="22" pageNumber="900">Y. Kumlutas</collectorName>
¸,
<collectorName id="268F4EBAFFC19F37FC1BA49515406A80" box="[958,1052,1469,1491]" pageId="22" pageNumber="900">Ç. Ilgaz</collectorName>
, and
<collectorName id="268F4EBAFFC19F37FBF8A49515ED6A81" box="[1117,1201,1469,1490]" pageId="22" pageNumber="900">A. Avcı</collectorName>
,
<date id="FFC40DACFFC19F37FB1AA49514706A80" box="[1215,1324,1469,1491]" pageId="22" pageNumber="900" value="2005-05-01">
<collectingDate id="EF80F444FFC19F37FB1AA49514706A80" box="[1215,1324,1469,1491]" pageId="22" pageNumber="900" value="2005-05-01">01.v.2005</collectingDate>
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.
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<emphasis id="B90EF77EFFC19F37FC99A4D712F16947" box="[828,941,1535,1556]" italics="true" pageId="22" pageNumber="900">
<typeStatus id="54C195CEFFC19F37FC99A4D712F16947" box="[828,941,1535,1556]" pageId="22" pageNumber="900" type="paratype">Paratypes</typeStatus>
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<paragraph id="8BC52B6CFFC19F37FC99A73614566961" blockId="22.[828,1290,1535,1587]" box="[828,1290,1565,1587]" pageId="22" pageNumber="900">Twenty-two specimens. See Appendix 1.</paragraph>
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<taxonomicName id="4C7A50EFFFC19F37FC99A777129A6927" box="[828,966,1631,1652]" class="Insecta" family="Lycaenidae" genus="Terra" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="900" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B90EF77EFFC19F37FC99A777129A6927" box="[828,966,1631,1652]" italics="true" pageId="22" pageNumber="900">Terra typica</emphasis>
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and
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west of
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(
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), S¸ anlıurfa
<collectingMunicipality id="6BA1B116FFC19F37FACCA7B412316982" pageId="22" pageNumber="900">Province</collectingMunicipality>
,
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(locality indicated by circle
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<figureCitation id="134137E9FFC19F37FAF6A79314CA6982" box="[1363,1430,1723,1745]" captionStart="Figure 2" captionStartId="4.[46,125,1401,1422]" captionTargetBox="[104,1695,472,1368]" captionTargetId="figure-0@4.[97,1697,465,1373]" captionTargetPageId="4" captionText="Figure 2. All available distribution records of the Eirenis persicus species group. Circles indicate the E. persicus specimens, with the persicus morph bearing unicoloured dorsal scales; squires indicate specimens from eastern Iran, southern Turkmenistan, and southern and western Pakistan, having both persicus morph that base of their anterior dorsal scales are darker, and walteri morph; stars indicate specimens referred to Eirenis mcmahoni (Wall, 1911); plus symbols indicate specimens in north-eastern Pakistan, having both persicus morphs that base of their anterior dorsal scales are darker, and with walteri morph; asterisks indicate specimens with the novum pattern; triangles indicate specimens of the nigrofasciatus morph. Circle 29 indicates the type locality of Cyclophis persicus Anderson, 1872; squire 2 indicates the type locality of Pseudocyclophis walteri Boettger, 1888; squire 7 indicates the type locality of Contia zebrina Wall, 1923; triangle 6 indicates the type locality of Contia persica var. nigrofasciata Nikolsky, 1907; star 3 indicates the type locality of Contia angusticeps Boulenger, 1894; plus symbols 25 indicate the type series localities of Contia mcmahoni Wall, 1911. For more details, see Appendix 1." figureDoi="http://doi.org/10.5281/zenodo.5360455" httpUri="https://zenodo.org/record/5360455/files/figure.png" pageId="22" pageNumber="900">Fig. 2</figureCitation>
).
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a.s.l.
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<emphasis id="B90EF77EFFC19F37FC99A63312F06863" box="[828,940,1819,1840]" italics="true" pageId="22" pageNumber="900">Diagnosis</emphasis>
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<taxonomicName id="4C7A50EFFFC19F37FC99A6121541681D" authority="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens, 2016" authorityName="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens" authorityYear="2016" box="[828,1053,1849,1871]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="900" phylum="Chordata" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B90EF77EFFC19F37FC99A6121541681D" box="[828,1053,1849,1871]" italics="true" pageId="22" pageNumber="900">Eirenis occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D4A05FFC19F37FB83A6121522681C" box="[1062,1150,1850,1871]" pageId="22" pageNumber="900" rank="species">sp. nov.</taxonomicNameLabel>
is differentiated from all other congeners by having the unique combination of 186213 ventrals in
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and
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; 4659 subcaudals in
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and
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; ratio of the tail length over total length
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<specimenCount id="9D7CE0E5FFC09F36FDB7A02D13776E44" box="[530,555,261,279]" pageId="23" pageNumber="901" type="male"></specimenCount>
and
<quantity id="4C828689FFC09F36FDC9A02A11FA6E64" metricMagnitude="-1" metricUnit="m" metricValue="4.0690800000000005" metricValueMax="4.823459999999999" metricValueMin="3.3147000000000006" pageId="23" pageNumber="901" unit="in" value="16.02" valueMax="18.99" valueMin="13.05">13.0518.99 in</quantity>
<specimenCount id="9D7CE0E5FFC09F36FF10A00C11966E65" box="[181,202,292,310]" pageId="23" pageNumber="901" type="female"></specimenCount>
; usually seven, rarely eight, infralabials; unicoloured dorsum, anterior dorsal scales without dark base.
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF2BA09210E16E9C" blockId="23.[142,757,442,1657]" box="[142,445,442,463]" pageId="23" pageNumber="901">
<emphasis id="B90EF77EFFC09F36FF2BA09210E16E9C" box="[142,445,442,463]" italics="true" pageId="23" pageNumber="901">
Description of the
<typeStatus id="54C195CEFFC09F36FEFAA09210E16E9C" box="[351,445,442,463]" pageId="23" pageNumber="901" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF2BA0F010046B9F" blockId="23.[142,757,442,1657]" pageId="23" pageNumber="901">
Body cylindrical; head small and not distinct from the neck; head and body scales not keeled; rostrum
<quantity id="4C828689FFC09F36FD33A0DF13A86D5F" box="[662,756,503,524]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="23" pageNumber="901" unit="mm" value="0.98">0.98 mm</quantity>
high and
<quantity id="4C828689FFC09F36FF5DA33E10046D78" box="[248,344,534,555]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="901" unit="mm" value="2.1">2.10 mm</quantity>
wide, bordered by two upper labials, two nasals, and two internasals; rostrum slightly curved towards top of the head and does not quite intrude between the internasals; nostrils situated on the nasal scales at either side; distance between the nostrils
<quantity id="4C828689FFC09F36FF2BA38711AC6D97" box="[142,240,687,708]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="23" pageNumber="901" unit="mm" value="2.44">2.44 mm</quantity>
; loreal absent; single preocular and postocular on each side of the head; eyes small, with circular pupil of
<quantity id="4C828689FFC09F36FF0AA3C4104F6C52" box="[175,275,748,770]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="23" pageNumber="901" unit="mm" value="0.96">0.96 mm</quantity>
diameter; length of the narrow frontal
<quantity id="4C828689FFC09F36FF2BA22211AF6C73" box="[142,243,778,800]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="901" unit="mm" value="2.3">2.30 mm</quantity>
, width
<quantity id="4C828689FFC09F36FEEAA22210E86C73" box="[335,436,778,800]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="23" pageNumber="901" unit="mm" value="1.54">1.54 mm</quantity>
; seven upper labials, third and fourth in contact with the eyes on each side; seven infralabials on each side; three pairs of infralabials in contact with the anterior chin shields at the left side and four at the right side; one anterior temporal and one posterior temporal on each side; ten dorsals plus temporal scales surrounding the posterior margin of the parietals; 15 dorsal scale rows at mid body, 17 at the level of one head length posterior to the head, and 15 at one head length anterior to the level of anus; two anal plates; two gular scales in contact with the posterior chin shield; 209 ventrals, 46/46 + one subcaudals; pileus length
<quantity id="4C828689FFC09F36FE00A55113596BDC" box="[421,517,1145,1167]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="23" pageNumber="901" unit="mm" value="7.62">7.62 mm</quantity>
, width
<quantity id="4C828689FFC09F36FDF0A55113E96BDC" box="[597,693,1145,1167]" metricMagnitude="-3" metricUnit="m" metricValue="4.26" pageId="23" pageNumber="901" unit="mm" value="4.26">4.26 mm</quantity>
; head height
<quantity id="4C828689FFC09F36FF46A5B010166BFE" box="[227,330,1176,1198]" metricMagnitude="-3" metricUnit="m" metricValue="3.66" pageId="23" pageNumber="901" unit="mm" value="3.66">3.66 mm</quantity>
; snoutvent length
<quantity id="4C828689FFC09F36FD9EA5B013E26BFE" box="[571,702,1176,1198]" metricMagnitude="-1" metricUnit="m" metricValue="3.1368" pageId="23" pageNumber="901" unit="mm" value="313.68">313.68 mm</quantity>
, tail length
<quantity id="4C828689FFC09F36FF44A59E10086B9F" box="[225,340,1206,1228]" metricMagnitude="-2" metricUnit="m" metricValue="4.866" pageId="23" pageNumber="901" unit="mm" value="48.66">48.66 mm</quantity>
.
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF03A5FD13A9692A" blockId="23.[142,757,442,1657]" pageId="23" pageNumber="901">
The background colour of the head and the dorsum in the living specimen is reddish brown, changing to yellowish or light brown in the preserved specimen, dorsum without pattern; blackish marking over the dorsal head, connected to a 56-scale-wide blackish band on the nape; dorsal marking extends to both sides of the head, reaching the nasals, four upper labials (1 4), around the eye, the postoculars, and the upper side of the temporals; a narrow black blotch extends from the upper part of the first sublabial to the contact of the third; a straight collar band can be seen over the lower part of the head; length of collar band 19 scales, not connected ventrally; the venter and the lower part of the tail uniformly cream, without maculation (
<figureCitation id="134137E9FFC09F36FD3BA74B13B5692A" box="[670,745,1635,1657]" captionStart="Figure 14" captionStartId="24.[166,245,1086,1105]" captionTargetBox="[325,1285,194,1056]" captionTargetId="figure-374@24.[325,1285,194,1056]" captionTargetPageId="24" captionText="Figure 14. Dorsal body (A), dorsal head (B), and lateral head (C) view of the holotype of Eirenis (Pseudocyclophis) occidentalis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5360473" httpUri="https://zenodo.org/record/5360473/files/figure.png" pageId="23" pageNumber="901">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF2BA78911A469E5" blockId="23.[142,757,1697,1902]" box="[142,248,1697,1718]" pageId="23" pageNumber="901">
<emphasis id="B90EF77EFFC09F36FF2BA78911A469E5" box="[142,248,1697,1718]" italics="true" pageId="23" pageNumber="901">Variation</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF2BA79711916863" blockId="23.[142,757,1697,1902]" pageId="23" pageNumber="901">
The
<typeStatus id="54C195CEFFC09F36FF67A797106F6987" box="[194,307,1727,1748]" pageId="23" pageNumber="901" type="paratype">paratypes</typeStatus>
agree in external morphology with the
<typeStatus id="54C195CEFFC09F36FF2BA7F511AD69A0" box="[142,241,1757,1779]" pageId="23" pageNumber="901" type="holotype">holotype</typeStatus>
. Detailed data on the
<typeStatus id="54C195CEFFC09F36FE4BA7F6130269A0" box="[494,606,1758,1779]" pageId="23" pageNumber="901" type="paratype">paratypes</typeStatus>
(
<emphasis id="B90EF77EFFC09F36FDCBA7F613DE69A0" box="[622,642,1758,1779]" italics="true" pageId="23" pageNumber="901">N</emphasis>
= 21;
<specimenCount id="9D7CE0E5FFC09F36FD6DA7F613B169A1" box="[712,749,1758,1779]" pageId="23" pageNumber="901" type="male">7♂</specimenCount>
,
<specimenCount id="9D7CE0E5FFC09F36FF2BA7D411E36842" box="[142,191,1788,1809]" pageId="23" pageNumber="901" type="female">14♀</specimenCount>
) and the single
<typeStatus id="54C195CEFFC09F36FE35A7D410A86841" box="[400,500,1788,1810]" pageId="23" pageNumber="901" type="holotype">holotype</typeStatus>
are given in Appendix 3.
</paragraph>
<paragraph id="8BC52B6CFFC09F36FF03A61115E16D5E" blockId="23.[142,757,1697,1902]" lastBlockId="23.[804,1420,197,525]" pageId="23" pageNumber="901">Dorsal background colour in preserved specimens yellowish or light brown, without dorsal pattern; black- ish marking on the dorsal head and a 47-scale-wide blackish band on the nape, usually separated from each other by a narrow light band crossing the tip of the parietals or connected to each other medially; the lateral head and the labials may be uniformly cream or the dorsal head marking may extend to the snout, the anterior or around eyes, adjacent supralabials or infralabials and upper margin of the temporal region; lower side of the head uniformly cream; the width of the napes band shrinks over the lateral sides of the neck and are detached over the ventral side.</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A31612E56D00" blockId="23.[804,1420,574,1545]" box="[804,953,574,595]" pageId="23" pageNumber="901">
<emphasis id="B90EF77EFFC09F36FC81A31612E56D00" box="[804,953,574,595]" italics="true" pageId="23" pageNumber="901">Comparisons</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A37514D0695A" blockId="23.[804,1420,574,1545]" pageId="23" pageNumber="901">
This new species is distinguished from
<taxonomicName id="4C7A50EFFFC09F36FB49A375143B6D21" authorityName="Anderson" authorityYear="1872" box="[1260,1383,605,626]" class="Reptilia" family="Colubridae" genus="Cyclophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B90EF77EFFC09F36FB49A375143B6D21" box="[1260,1383,605,626]" italics="true" pageId="23" pageNumber="901">E. persicus</emphasis>
</taxonomicName>
by having a higher average number of ventrals, 186 213 (
<emphasis id="B90EF77EFFC09F36FCC1A3B2122C6DFC" box="[868,880,666,687]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 199.78) in
<specimenCount id="9D7CE0E5FFC09F36FBA2A3B5157C6DFC" box="[1031,1056,669,687]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 204224 (
<emphasis id="B90EF77EFFC09F36FB74A3B215816DFC" box="[1233,1245,666,687]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 211.29) in
<specimenCount id="9D7CE0E5FFC09F36FAD3A3B514D76DFC" box="[1398,1419,669,687]" pageId="23" pageNumber="901" type="female"></specimenCount>
[compared with 190203 (
<emphasis id="B90EF77EFFC09F36FB2DA39115C86D9D" box="[1160,1172,697,718]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 195.6) in
<specimenCount id="9D7CE0E5FFC09F36FA97A39414176D9D" box="[1330,1355,700,718]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 198214 (
<emphasis id="B90EF77EFFC09F36FC32A3FF12FF6DBF" box="[919,931,727,748]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 205.0) in
<specimenCount id="9D7CE0E5FFC09F36FB8FA3F215636DBF" box="[1066,1087,730,748]" pageId="23" pageNumber="901" type="female"></specimenCount>
in the latter species]; a lower average number of subcaudals, 4659 (
<emphasis id="B90EF77EFFC09F36FB5EA3DE145B6C58" box="[1275,1287,758,779]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 51.89) in
<specimenCount id="9D7CE0E5FFC09F36FC81A23F12616C7A" box="[804,829,791,809]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 4258 (
<emphasis id="B90EF77EFFC09F36FC73A23C12BE6C7A" box="[982,994,788,809]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 51.23) in
<specimenCount id="9D7CE0E5FFC09F36FBC8A23F15DE6C7A" box="[1133,1154,791,809]" pageId="23" pageNumber="901" type="female"></specimenCount>
[compared with 6483 (
<emphasis id="B90EF77EFFC09F36FC91A21B121C6C1B" box="[820,832,819,840]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 75.0) in
<specimenCount id="9D7CE0E5FFC09F36FC6AA21E12B46C1B" box="[975,1000,822,840]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 5477 (
<emphasis id="B90EF77EFFC09F36FB07A21B15F26C1B" box="[1186,1198,819,840]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 66.25) in
<specimenCount id="9D7CE0E5FFC09F36FAEEA21E143C6C1B" box="[1355,1376,822,840]" pageId="23" pageNumber="901" type="female"></specimenCount>
in
<taxonomicName id="4C7A50EFFFC09F36FC81A27A12C06C34" authorityName="Anderson" authorityYear="1872" box="[804,924,850,871]" class="Reptilia" family="Colubridae" genus="Cyclophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B90EF77EFFC09F36FC81A27A12C06C34" box="[804,924,850,871]" italics="true" pageId="23" pageNumber="901">E. persicus</emphasis>
</taxonomicName>
] and a lower average ratio of the tail length over the total length, 13.4319.49 (
<emphasis id="B90EF77EFFC09F36FB0AA25815E76CD6" box="[1199,1211,880,901]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 16.30) in
<specimenCount id="9D7CE0E5FFC09F36FAE7A25B14076CD6" box="[1346,1371,883,901]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 13.0518.99 (
<emphasis id="B90EF77EFFC09F36FC63A2A7128E6CF7" box="[966,978,911,932]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 15.65) in
<specimenCount id="9D7CE0E5FFC09F36FBC4A2BA152A6CF7" box="[1121,1142,914,932]" pageId="23" pageNumber="901" type="female"></specimenCount>
[compared with 21.93 29.09 (
<emphasis id="B90EF77EFFC09F36FCDDA28512D86C91" box="[888,900,941,962]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 23.80) in
<specimenCount id="9D7CE0E5FFC09F36FBA8A298157A6C91" box="[1037,1062,944,962]" pageId="23" pageNumber="901" type="male"></specimenCount>
and 18.9222.60 (
<emphasis id="B90EF77EFFC09F36FB5AA28514576C91" box="[1279,1291,941,962]" italics="true" pageId="23" pageNumber="901">x</emphasis>
= 20.42) in
<specimenCount id="9D7CE0E5FFC09F36FC81A2E712656CB2" box="[804,825,975,993]" pageId="23" pageNumber="901" type="female"></specimenCount>
in
<taxonomicName id="4C7A50EFFFC09F36FCC4A2E512806CB1" authorityName="Anderson" authorityYear="1872" box="[865,988,973,994]" class="Reptilia" family="Colubridae" genus="Cyclophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B90EF77EFFC09F36FCC4A2E512806CB1" box="[865,988,973,994]" italics="true" pageId="23" pageNumber="901">E. persicus</emphasis>
</taxonomicName>
]; usually seven (78.3%), rarely eight, infralabials [compared with usually eight (88.9%), rarely seven]. The new species has a unicoloured dorsum and anterior dorsal scales without dark bases, which is different to
<taxonomicName id="4C7A50EFFFC09F36FC2DA56F15606B08" box="[904,1084,1094,1116]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B90EF77EFFC09F36FC2DA56F15606B08" box="[904,1084,1094,1116]" italics="true" pageId="23" pageNumber="901">E. nigrofasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7A50EFFFC09F36FBEFA56F15EE6B08" authorityName="Boettger" authorityYear="1888" box="[1098,1202,1094,1116]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="walteri">
<emphasis id="B90EF77EFFC09F36FBEFA56F15EE6B08" box="[1098,1202,1094,1116]" italics="true" pageId="23" pageNumber="901">E. walteri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7A50EFFFC09F36FB54A56F14D76B0F" authorityName="Boulenger" authorityYear="1894" box="[1265,1419,1095,1116]" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="angusticeps">
<emphasis id="B90EF77EFFC09F36FB54A56F14D76B0F" box="[1265,1419,1095,1116]" italics="true" pageId="23" pageNumber="901">E. angusticeps</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7A50EFFFC09F36FC89A54D12916B29" box="[812,973,1125,1146]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B90EF77EFFC09F36FC89A54D12916B29" box="[812,973,1125,1146]" italics="true" pageId="23" pageNumber="901">nigrofasciatus</emphasis>
</taxonomicName>
morph in
<taxonomicName id="4C7A50EFFFC09F36FBF5A54E144C6B29" box="[1104,1296,1125,1147]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B90EF77EFFC09F36FBF5A54E144C6B29" box="[1104,1296,1125,1147]" italics="true" pageId="23" pageNumber="901">E. nigrofasciatus</emphasis>
</taxonomicName>
, and both
<taxonomicName id="4C7A50EFFFC09F36FC81A5AC12226BCA" authorityName="Anderson" authorityYear="1872" box="[804,894,1156,1177]" class="Reptilia" family="Colubridae" genus="Cyclophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B90EF77EFFC09F36FC81A5AC12226BCA" box="[804,894,1156,1177]" italics="true" pageId="23" pageNumber="901">persicus</emphasis>
</taxonomicName>
morph, the anterior dorsal scales having dark bases, and the
<taxonomicName id="4C7A50EFFFC09F36FC45A58A156E6BE4" authorityName="Boettger" authorityYear="1888" box="[992,1074,1186,1207]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="walteri">
<emphasis id="B90EF77EFFC09F36FC45A58A156E6BE4" box="[992,1074,1186,1207]" italics="true" pageId="23" pageNumber="901">walteri</emphasis>
</taxonomicName>
morph in
<taxonomicName id="4C7A50EFFFC09F36FB1DA58B14736BE4" authorityName="Boettger" authorityYear="1888" box="[1208,1327,1186,1208]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="walteri">
<emphasis id="B90EF77EFFC09F36FB1DA58B14736BE4" box="[1208,1327,1186,1208]" italics="true" pageId="23" pageNumber="901">E. walteri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A50EFFFC09F36FAD7A58B12E96B85" authorityName="Boulenger" authorityYear="1894" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="angusticeps">
<emphasis id="B90EF77EFFC09F36FAD7A58B12E96B85" italics="true" pageId="23" pageNumber="901">E. angusticeps</emphasis>
</taxonomicName>
). The new species also differs from
<taxonomicName id="4C7A50EFFFC09F36FC81A5C812B66BA7" box="[804,1002,1247,1269]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B90EF77EFFC09F36FC81A5C812B66BA7" box="[804,1002,1247,1269]" italics="true" pageId="23" pageNumber="901">E. nigrofasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7A50EFFFC09F36FC58A5C8152C6BA7" authorityName="Boettger" authorityYear="1888" box="[1021,1136,1247,1269]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="walteri">
<emphasis id="B90EF77EFFC09F36FC58A5C8152C6BA7" box="[1021,1136,1247,1269]" italics="true" pageId="23" pageNumber="901">E. walteri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7A50EFFFC09F36FB1EA5C814346BA6" authorityName="Boulenger" authorityYear="1894" box="[1211,1384,1248,1269]" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="angusticeps">
<emphasis id="B90EF77EFFC09F36FB1EA5C814346BA6" box="[1211,1384,1248,1269]" italics="true" pageId="23" pageNumber="901">E. angusticeps</emphasis>
</taxonomicName>
in having a lower number of subcaudal scales,
<quantity id="4C828689FFC09F36FA82A5D614D76A47" box="[1319,1419,1278,1300]" metricMagnitude="0" metricUnit="m" metricValue="1.2827000000000002" metricValueMax="1.4986000000000002" metricValueMin="1.0668000000000002" pageId="23" pageNumber="901" unit="in" value="50.5" valueMax="59.0" valueMin="42.0">4259 in</quantity>
both sexes (compared with
<quantity id="4C828689FFC09F36FBFEA43415E26A60" box="[1115,1214,1308,1331]" metricMagnitude="0" metricUnit="m" metricValue="2.0701" metricValueMax="2.4384" metricValueMin="1.7018" pageId="23" pageNumber="901" unit="in" value="81.5" valueMax="96.0" valueMin="67.0">6796 in</quantity>
<taxonomicName id="4C7A50EFFFC09F36FB63A43514D86A61" box="[1222,1412,1309,1330]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B90EF77EFFC09F36FB63A43514D86A61" box="[1222,1412,1309,1330]" italics="true" pageId="23" pageNumber="901">E. nigrofasciatus</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C7A50EFFFC09F36FC03A414154B6A03" authorityName="Boettger" authorityYear="1888" box="[934,1047,1339,1361]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="walteri">
<emphasis id="B90EF77EFFC09F36FC03A414154B6A03" box="[934,1047,1339,1361]" italics="true" pageId="23" pageNumber="901">E. walteri</emphasis>
</taxonomicName>
, and
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<taxonomicName id="4C7A50EFFFC09F36FB70A41414276A02" authorityName="Boulenger" authorityYear="1894" box="[1237,1403,1340,1361]" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="angusticeps">
<emphasis id="B90EF77EFFC09F36FB70A41414276A02" box="[1237,1403,1340,1361]" italics="true" pageId="23" pageNumber="901">E. angusticeps</emphasis>
</taxonomicName>
).
<taxonomicName id="4C7A50EFFFC09F36FC81A472155F6A3C" authority="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens, 2016" authorityName="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens" authorityYear="2016" box="[804,1027,1370,1391]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B90EF77EFFC09F36FC81A472155F6A3C" box="[804,1027,1370,1391]" italics="true" pageId="23" pageNumber="901">Eirenis occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D4A05FFC09F36FBAEA472153D6A3C" box="[1035,1121,1370,1391]" pageId="23" pageNumber="901" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C7A50EFFFC09F36FB50A47214D76A3C" baseAuthorityName="Wall" baseAuthorityYear="1911" box="[1269,1419,1370,1391]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="mcmahoni">
<emphasis id="B90EF77EFFC09F36FB50A47214D76A3C" box="[1269,1419,1370,1391]" italics="true" pageId="23" pageNumber="901">E. mcmahoni</emphasis>
</taxonomicName>
in having 15 anterior dorsal scales (compared with
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<taxonomicName id="4C7A50EFFFC09F36FCE0A4B012876AFF" baseAuthorityName="Wall" baseAuthorityYear="1911" box="[837,987,1431,1453]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="mcmahoni">
<emphasis id="B90EF77EFFC09F36FCE0A4B012876AFF" box="[837,987,1431,1453]" italics="true" pageId="23" pageNumber="901">E. mcmahoni</emphasis>
</taxonomicName>
).
<taxonomicName id="4C7A50EFFFC09F36FC51A4B015896AFF" authority="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens, 2016" authorityName="Rajabizadeh &amp; Nagy &amp; Adriaens &amp; Avci &amp; Masroor &amp; Schmidtler &amp; Nazarov &amp; Esmaeili &amp; Christiaens" authorityYear="2016" box="[1012,1237,1431,1453]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B90EF77EFFC09F36FC51A4B015896AFF" box="[1012,1237,1431,1453]" italics="true" pageId="23" pageNumber="901">Eirenis occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D4A05FFC09F36FB7AA4BF146B6AFF" box="[1247,1335,1431,1452]" pageId="23" pageNumber="901" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the other species of the genus
<taxonomicName id="4C7A50EFFFC09F36FA9BA49E14D76A98" authorityName="Jan" authorityYear="1863" box="[1342,1419,1462,1483]" class="Reptilia" family="Colubridae" genus="Eirenis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="genus">
<emphasis id="B90EF77EFFC09F36FA9BA49E14D76A98" box="[1342,1419,1462,1483]" italics="true" pageId="23" pageNumber="901">Eirenis</emphasis>
</taxonomicName>
by having 15 rows of dorsal scales at the mid body and usually no loreal scales (see
<bibRefCitation id="EFEB569DFFC09F36FB38A4DB14DC695A" author="Mahlow K &amp; Tillack F &amp; Schmidtler JF &amp; Muller J" box="[1181,1408,1523,1545]" pageId="23" pageNumber="901" pagination="41 - 85" refId="ref16487" refString="Mahlow K, Tillack F, Schmidtler JF, Muller J. 2013. An annotated checklist, description and key to the dwarf snakes of the genus Eirenis Jan, 1863 (Reptilia: Squamata: Colubridae), with special emphasis on the dentition. Vertebrate Zoology 63: 41 - 85." type="journal article" year="2013">
Mahlow
<emphasis id="B90EF77EFFC09F36FAA5A4DB146C695B" box="[1280,1328,1523,1544]" italics="true" pageId="23" pageNumber="901">et al</emphasis>
., 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A71212F3691C" blockId="23.[804,1419,1594,1739]" box="[804,943,1594,1615]" pageId="23" pageNumber="901">
<emphasis id="B90EF77EFFC09F36FC81A71212F3691C" box="[804,943,1594,1615]" italics="true" pageId="23" pageNumber="901">Distribution</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A77112286998" blockId="23.[804,1419,1594,1739]" pageId="23" pageNumber="901">
Occurs in south-eastern
<collectingCountry id="F36D6BFCFFC09F36FB90A77115D8693C" box="[1077,1156,1625,1647]" name="Turkey" pageId="23" pageNumber="901">Turkey</collectingCountry>
(eastern Taurus Mountains and foothills) up to
<collectingRegion id="49BEE58EFFC09F36FBC7A75F143469DE" box="[1122,1384,1655,1677]" country="Iran" name="Kermanshah" pageId="23" pageNumber="901">Kermanshah province</collectingRegion>
in western
<collectingCountry id="F36D6BFCFFC09F36FC22A7BE12E569F8" box="[903,953,1686,1707]" name="Iran" pageId="23" pageNumber="901">Iran</collectingCountry>
(localities indicated by circles
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<figureCitation id="134137E9FFC09F36FC81A79C123B6998" box="[804,871,1716,1739]" captionStart="Figure 2" captionStartId="4.[46,125,1401,1422]" captionTargetBox="[104,1695,472,1368]" captionTargetId="figure-0@4.[97,1697,465,1373]" captionTargetPageId="4" captionText="Figure 2. All available distribution records of the Eirenis persicus species group. Circles indicate the E. persicus specimens, with the persicus morph bearing unicoloured dorsal scales; squires indicate specimens from eastern Iran, southern Turkmenistan, and southern and western Pakistan, having both persicus morph that base of their anterior dorsal scales are darker, and walteri morph; stars indicate specimens referred to Eirenis mcmahoni (Wall, 1911); plus symbols indicate specimens in north-eastern Pakistan, having both persicus morphs that base of their anterior dorsal scales are darker, and with walteri morph; asterisks indicate specimens with the novum pattern; triangles indicate specimens of the nigrofasciatus morph. Circle 29 indicates the type locality of Cyclophis persicus Anderson, 1872; squire 2 indicates the type locality of Pseudocyclophis walteri Boettger, 1888; squire 7 indicates the type locality of Contia zebrina Wall, 1923; triangle 6 indicates the type locality of Contia persica var. nigrofasciata Nikolsky, 1907; star 3 indicates the type locality of Contia angusticeps Boulenger, 1894; plus symbols 25 indicate the type series localities of Contia mcmahoni Wall, 1911. For more details, see Appendix 1." figureDoi="http://doi.org/10.5281/zenodo.5360455" httpUri="https://zenodo.org/record/5360455/files/figure.png" pageId="23" pageNumber="901">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A7D412C76842" blockId="23.[804,1420,1788,1902]" box="[804,923,1788,1809]" pageId="23" pageNumber="901">
<emphasis id="B90EF77EFFC09F36FC81A7D412C76842" box="[804,923,1788,1809]" italics="true" pageId="23" pageNumber="901">Etymology</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC09F36FC81A63214D7683E" blockId="23.[804,1420,1788,1902]" pageId="23" pageNumber="901">
The name occidentalis (Latin: western) indicates the westernmost known distribution of the taxon amongst the known species within the subgenus
<taxonomicName id="4C7A50EFFFC09F36FB72A67014D8683E" authorityName="Boettger" authorityYear="1888" box="[1239,1412,1880,1901]" class="Reptilia" family="Colubridae" genus="Pseudocyclophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="901" phylum="Chordata" rank="genus">
<emphasis id="B90EF77EFFC09F36FB72A67014D8683E" box="[1239,1412,1880,1901]" italics="true" pageId="23" pageNumber="901">Pseudocyclophis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>