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<document id="065CD52987A8354D91B26AF169659DFA" ID-CLB-Dataset="59554" ID-DOI="10.5852/ejt.2023.885.2189" ID-GBIF-Dataset="f8509ab7-b884-41aa-a5ee-f8c425a46031" ID-ISSN="2118-9773" ID-Zenodo-Dep="8205500" ID-ZooBank="444CE1D2-8098-4CDE-9B97-927CA13FC39F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690898759111" checkinUser="felipe" docAuthor="Means, Jackson C., Bouzan, Rodrigo S., Iniesta, Luiz Felipe Moretti, Martínez-Torres, Daniela, Vasquez-Valverde, Luisa F., Brescovit, Antonio D. &amp; Ivanov, Kaloyan" docDate="2023" docId="03CC1013FFC93C7A9708FB2A36ECFDBC" docLanguage="en" docName="ejt-885_65-85.pdf" docOrigin="European Journal of Taxonomy 885 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.885.2189" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dibolostethus kattani Means, Bouzan, Martinez-Torres &amp; Ivanov 2023, sp. nov." docType="treatment" docUuid="E4B3174C-2DC2-4584-B664-D0E4FB12D6B7" docUuidSource="ZooBank" docVersion="4" lastPageNumber="78" masterDocId="FFF5686BFFC23C77966FFF973518FFDD" masterDocTitle="A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot" masterLastPageNumber="85" masterPageNumber="65" pageNumber="76" updateTime="1699550199751" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="06DDE11A8C95A8C9F897A998F1C5F843">A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot</mods:title>
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<mods:namePart id="3302C2523B8CB47C256229A738CA1D89">Means, Jackson C.</mods:namePart>
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<mods:affiliation id="3DA85E00BFBB82F11EA9C1C3666949DD">Virginia Museum of Natural History, 21 Starling Ave., Martinsville, Virginia 24112, USA.</mods:affiliation>
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<mods:namePart id="474D61A3B65073F586761F2A29054E7F">Bouzan, Rodrigo S.</mods:namePart>
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<mods:affiliation id="D3FB90EFE13902BE4B805524BB90CC40">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="FCE5A5CCEAC3F588D059D632B1954481" type="email">rodrigobouzan@ib.usp.br</mods:nameIdentifier>
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<mods:namePart id="B7DB2E566D40963A6D265D026F5D77E4">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:affiliation id="DE3E4AB6C0ECF8B05D8EE1EBB483F261">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="6CF56FF4F85724F2F2137B210480F698">Martínez-Torres, Daniela</mods:namePart>
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<mods:affiliation id="4E463CAEDFD370C4E62380CDCB568A6D">Grupo de Investigación en Aracnología &amp; Miriapodología (GAM-UN), National University of Colombia, Bogotá, Colombia.</mods:affiliation>
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<mods:affiliation id="75ACAA78792304AA863F594168F9FCDE">Department of Entomology, Virginia Tech, 216 A Price Hall, Blacksburg, VA 24061, USA.</mods:affiliation>
<mods:nameIdentifier id="1654438AD41206ADC0C0CF1A912DC19D" type="email">luisafvv@vt.edu</mods:nameIdentifier>
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<mods:namePart id="D2308825CF355C400570CEA13492AF58">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="CA367EB19870B6C874393755CB4EE982">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="73274889E1092734911DCB46D031CA17">Ivanov, Kaloyan</mods:namePart>
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<mods:affiliation id="56002838C9C2604D4717C835BFBC8D42">Virginia Museum of Natural History, 21 Starling Ave., Martinsville, Virginia 24112, USA.</mods:affiliation>
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<mods:title id="E312FF0A85B88DF8141B1FE0566A7859">European Journal of Taxonomy</mods:title>
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<mods:date id="C43EA951813042865940662FCBBF7F53">2023</mods:date>
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<mods:number id="37FA189AAAB9BFF468FC41A9BE560011">885</mods:number>
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<treatment id="03CC1013FFC93C7A9708FB2A36ECFDBC" ID-DOI="http://doi.org/10.5281/zenodo.8206249" ID-GBIF-Taxon="212423779" ID-Zenodo-Dep="8206249" LSID="urn:lsid:zoobank.org:act:E4B3174C-2DC2-4584-B664-D0E4FB12D6B7" httpUri="http://treatment.plazi.org/id/03CC1013FFC93C7A9708FB2A36ECFDBC" lastPageId="13" lastPageNumber="78" pageId="11" pageNumber="76">
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<paragraph id="8BDAA105FFC93C7C9708FB2A31D5FB05" blockId="11.[359,1229,1213,1310]" box="[359,1229,1213,1240]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C9708FB2A31D5FB05" box="[359,1229,1213,1240]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="76" reason="2">
<taxonomicName id="4C65DA86FFC93C7C9708FB2A3172FB05" authority="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityName="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityYear="2023" box="[359,1130,1213,1240]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFC93C7C9708FB2A3772FB05" bold="true" box="[359,618,1213,1240]" italics="true" pageId="11" pageNumber="76">Dibolostethus kattani</emphasis>
Means, Bouzan, Martínez-Torres &amp; Ivanov
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<taxonomicNameLabel id="A222C06CFFC93C7C921DFB2931D5FB05" box="[1138,1229,1214,1240]" pageId="11" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BDAA105FFC93C7C9703FB7731DFFB26" blockId="11.[359,1229,1213,1310]" box="[364,1223,1248,1275]" pageId="11" pageNumber="76">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BDAA105FFC93C7C94F0FA93368DFAC3" blockId="11.[359,1229,1213,1310]" box="[671,917,1284,1310]" pageId="11" pageNumber="76">
<figureCitation id="135EBD80FFC93C7C94F0FA93361FFAC3" box="[671,775,1284,1310]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1963,1989]" captionStartId-1="5.[189,232,1822,1848]" captionTargetBox-0="[378,1032,415,1893]" captionTargetBox-1="[189,1398,642,1776]" captionTargetId-0="figure-14@4.[310,1277,265,1924]" captionTargetId-1="figure-157@5.[189,1398,642,1776]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 1. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm." captionText-1="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi-0="http://doi.org/10.5281/zenodo.8205502" figureDoi-1="http://doi.org/10.5281/zenodo.8205504" httpUri-0="https://zenodo.org/record/8205502/files/figure.png" httpUri-1="https://zenodo.org/record/8205504/files/figure.png" pageId="11" pageNumber="76">Figs 1–2</figureCitation>
,
<figureCitation id="135EBD80FFC93C7C957BFA933644FAC3" box="[788,860,1284,1310]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="11" pageNumber="76">3B–D</figureCitation>
,
<figureCitation id="135EBD80FFC93C7C9507FA933661FAC3" box="[872,889,1284,1310]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="11" pageNumber="76">7</figureCitation>
,
<figureCitation id="135EBD80FFC93C7C95E9FA93368DFAC3" box="[902,917,1284,1310]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="11" pageNumber="76">9</figureCitation>
</paragraph>
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<subSubSection id="C37FF28EFFC93C7C96D2FAD23477F9E3" pageId="11" pageNumber="76" type="diagnosis">
<paragraph id="8BDAA105FFC93C7C96D2FAD2342EFA82" blockId="11.[189,310,1349,1375]" box="[189,310,1349,1375]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C96D2FAD2342EFA82" bold="true" box="[189,310,1349,1375]" fontSize="11" level="3" pageId="11" pageNumber="76" reason="3">
<emphasis id="B9117D17FFC93C7C96D2FAD2342EFA82" bold="true" box="[189,310,1349,1375]" pageId="11" pageNumber="76">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2FAE43477F9E3" blockId="11.[188,1399,1395,1598]" pageId="11" pageNumber="76">
Adults males of
<taxonomicName id="4C65DA86FFC93C7C9710FAE334ECFA50" authority="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov, 2023" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[383,500,1395,1421]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFC93C7C9710FAE334ECFA50" box="[383,500,1395,1421]" italics="true" pageId="11" pageNumber="76">D. kattani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC93C7C9793FAE3374EFA53" box="[508,598,1396,1422]" pageId="11" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
differ from those of
<taxonomicName id="4C65DA86FFC93C7C9523FAE336D5FA50" authorityName="Hoffman" authorityYear="2009" box="[844,973,1396,1421]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFC93C7C9523FAE336D5FA50" box="[844,973,1396,1421]" italics="true" pageId="11" pageNumber="76">D. sicarius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C65DA86FFC93C7C9269FAE331BEFA50" authority="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov, 2023" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[1030,1190,1396,1421]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
<emphasis id="B9117D17FFC93C7C9269FAE331BEFA50" box="[1030,1190,1396,1421]" italics="true" pageId="11" pageNumber="76">D. inopinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC93C7C92C2FAE33010FA53" box="[1197,1288,1396,1422]" pageId="11" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
based on the presence of two pairs of short, blunt projections on the sternite of the 5
<superScript id="7C100C4DFFC93C7C9273FA013131FA79" attach="left" box="[1052,1065,1430,1444]" fontSize="6" pageId="11" pageNumber="76">th</superScript>
body ring (
<figureCitation id="135EBD80FFC93C7C92DFFA003014FA6C" box="[1200,1292,1431,1457]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="11" pageNumber="76">Fig. 3D</figureCitation>
), and the following combination of gonopodal characteristics: solenomere curving with distinct ridge at midway point, tip relatively straight (
<figureCitation id="135EBD80FFC93C7C947BFA4A378AFA2A" box="[532,658,1501,1527]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="11" pageNumber="76">Fig. 7A, S</figureCitation>
); prefemoral process slightly thinner basally, with shelf under linear lobe just before apex (
<figureCitation id="135EBD80FFC93C7C9464F9973794F9C6" box="[523,652,1536,1563]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="11" pageNumber="76">Figs 7A, C</figureCitation>
, PfP); apex with large, tongue-shaped lobe, best seen in anterior view (
<figureCitation id="135EBD80FFC93C7C976AF9B4347AF9E3" box="[261,354,1571,1598]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="11" pageNumber="76">Fig. 7B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC93C7C96D2F9F237C3F929" pageId="11" pageNumber="76" type="etymology">
<paragraph id="8BDAA105FFC93C7C96D2F9F2345CF9A2" blockId="11.[189,324,1637,1663]" box="[189,324,1637,1663]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C96D2F9F2345CF9A2" bold="true" box="[189,324,1637,1663]" fontSize="11" level="3" pageId="11" pageNumber="76" reason="3">
<emphasis id="B9117D17FFC93C7C96D2F9F2345CF9A2" bold="true" box="[189,324,1637,1663]" pageId="11" pageNumber="76">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2F90437C3F929" blockId="11.[189,1399,1683,1780]" pageId="11" pageNumber="76">
The species is named in honor of one of its first collectors, the late Dr Gustavo Kattan. Throughout his career, Dr Kattan collected myriapod specimens across
<collectingCountry id="F372E195FFC93C7C9520F92036DBF90C" box="[847,963,1719,1745]" name="Colombia" pageId="11" pageNumber="76">Colombia</collectingCountry>
and donated them to the Instituto de Ciencias Naturales. Noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC93C7B96D2F88B3146FDC2" lastPageId="12" lastPageNumber="77" pageId="11" pageNumber="76" type="materials_examined">
<paragraph id="8BDAA105FFC93C7C96D2F88B3474F8E8" blockId="11.[189,364,1819,1846]" box="[189,364,1819,1846]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C96D2F88B3474F8E8" bold="true" box="[189,364,1819,1846]" fontSize="11" level="3" pageId="11" pageNumber="76" reason="3">
<emphasis id="B9117D17FFC93C7C96D2F88B3474F8E8" bold="true" box="[189,364,1819,1846]" pageId="11" pageNumber="76">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC93C7C9685F8DD31DEF877" blockId="11.[189,1399,1866,1963]" pageId="11" pageNumber="76">
<materialsCitation id="3B0DAB58FFC93C7C9685F8DD31DEF877" ID-GBIF-Occurrence="4167732307" collectedFrom="under leaf litter in tropical forest" collectingDate="1990-12-10" collectorName="C. Murcia &amp; G. Kattan" country="Colombia" latitude="3.496" location="Cali-Buenaventura" longLatPrecision="78" longitude="-76.613" municipality="Cali" pageId="11" pageNumber="76" specimenCode="ICN-MD-1317" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="holotype">
<emphasis id="B9117D17FFC93C7C9685F8DD3444F8B9" bold="true" box="[234,348,1866,1892]" pageId="11" pageNumber="76">
<typeStatus id="54DE1FA7FFC93C7C9685F8DD3444F8B9" box="[234,348,1866,1892]" pageId="11" pageNumber="76" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F372E195FFC93C7C96D2F8FA3440F85A" box="[189,344,1901,1927]" name="Colombia" pageId="11" pageNumber="76">COLOMBIA</collectingCountry>
•
<specimenCount id="9D636A8CFFC93C7C971FF8F93491F855" box="[368,393,1902,1928]" pageId="11" pageNumber="76" type="male">♂</specimenCount>
;
<collectingRegion id="49A16FE7FFC93C7C97FAF8FA3757F85A" box="[405,591,1901,1927]" country="Colombia" name="Valle del Cauca" pageId="11" pageNumber="76">Valle del Cauca</collectingRegion>
,
<collectingMunicipality id="6BBE3B7FFFC93C7C9435F8FA3796F85A" box="[602,654,1901,1927]" pageId="11" pageNumber="76">Cali</collectingMunicipality>
,
<quantity id="4C9D0CE0FFC93C7C94F5F8FA37FBF85A" box="[666,739,1901,1928]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="76" unit="km" value="15.0">
<locationDeviation id="9A68B827FFC93C7C94F5F8FA37FBF85A" box="[666,739,1901,1928]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="76" unit="km" value="15.0">15 km</locationDeviation>
</quantity>
vía
<location id="8EBAF7DEFFC93C7C957AF8FA36EEF85A" LSID="urn:lsid:plazi:treatment:03CC1013FFC93C7A9708FB2A36ECFDBC:8EBAF7DEFFC93C7C957AF8FA36EEF85A" box="[789,1014,1901,1927]" country="Colombia" latitude="3.496" longLatPrecision="78" longitude="-76.613" municipality="Cali" name="Cali-Buenaventura" pageId="11" pageNumber="76" stateProvince="Valle del Cauca">Cali-Buenaventura</location>
; [
<geoCoordinate id="EE51C7C2FFC93C7C9266F8FA3161F855" box="[1033,1145,1901,1928]" degrees="3.496" direction="north" orientation="latitude" pageId="11" pageNumber="76" precision="55" value="3.496">3.496° N</geoCoordinate>
,
<geoCoordinate id="EE51C7C2FFC93C7C92EBF8FA3010F855" box="[1156,1288,1901,1928]" degrees="76.613" direction="west" orientation="longitude" pageId="11" pageNumber="76" precision="55" value="-76.613">76.613° W</geoCoordinate>
];
<date id="FFDB87C5FFC93C7C9373F8FA35E3F877" pageId="11" pageNumber="76" value="1990-12-10">
<collectingDate id="EF9F7E2DFFC93C7C9373F8FA35E3F877" pageId="11" pageNumber="76" value="1990-12-10">10 Dec. 1990</collectingDate>
</date>
;
<collectorName id="2690C4D3FFC93C7C9768F8073466F877" box="[263,382,1936,1962]" pageId="11" pageNumber="76">C. Murcia</collectorName>
and
<collectorName id="2690C4D3FFC93C7C97D8F8073731F876" box="[439,553,1936,1963]" pageId="11" pageNumber="76">G. Kattan</collectorName>
leg.;
<collectedFrom id="3879E2F8FFC93C7C9404F80736F5F877" box="[619,1005,1936,1962]" pageId="11" pageNumber="76">under leaf litter in tropical forest</collectedFrom>
;
<specimenCode id="DBC3097EFFC93C7C9596F80731B4F877" box="[1017,1196,1936,1962]" pageId="11" pageNumber="76">ICN-MD-1317</specimenCode>
A.
</materialsCitation>
</paragraph>
<paragraph id="8BDAA105FFCE3C7B9685FE9E315CFE9B" blockId="12.[189,1092,265,326]" pageId="12" pageNumber="77">
<materialsCitation id="3B0DAB58FFCE3C7B9685FE9E3158FE9B" ID-GBIF-Occurrence="4167732305" collectedFrom="under leaf litter in tropical forest" collectingDate="1990-12-10" collectorName="C. Murcia &amp; G. Kattan" country="Colombia" latitude="3.496" location="COLOMBIA" longLatPrecision="78" longitude="-76.613" municipality="Cali" pageId="12" pageNumber="77" specimenCode="ICN-MD-1317B" specimenCount="2" specimenCount-male="2" stateProvince="Valle del Cauca" typeStatus="paratype">
<emphasis id="B9117D17FFCE3C7B9685FE9E347FFEFE" bold="true" box="[234,359,265,291]" pageId="12" pageNumber="77">
<typeStatus id="54DE1FA7FFCE3C7B9685FE9E347FFEFE" box="[234,359,265,291]" pageId="12" pageNumber="77" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F372E195FFCE3C7B96D2FEBB3440FE9B" box="[189,344,300,326]" name="Colombia" pageId="12" pageNumber="77">COLOMBIA</collectingCountry>
•
<specimenCount id="9D636A8CFFCE3C7B971EFEBB34ABFE9B" box="[369,435,300,326]" pageId="12" pageNumber="77" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBC3097EFFCE3C7B9516FEBB3158FE9B" box="[889,1088,300,326]" pageId="12" pageNumber="77">ICN-MD-1317B</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDAA105FFCE3C7B96D2FEFA3461FE5A" blockId="12.[189,377,365,391]" box="[189,377,365,391]" pageId="12" pageNumber="77">
<heading id="D0921669FFCE3C7B96D2FEFA3461FE5A" bold="true" box="[189,377,365,391]" fontSize="11" level="3" pageId="12" pageNumber="77" reason="3">
<emphasis id="B9117D17FFCE3C7B96D2FEFA3461FE5A" bold="true" box="[189,377,365,391]" pageId="12" pageNumber="77">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFCE3C7B96D2FE0B3146FDC2" blockId="12.[189,1399,412,544]" pageId="12" pageNumber="77">
<materialsCitation id="3B0DAB58FFCE3C7B96D2FE0B307DFE04" ID-GBIF-Occurrence="4167732301" collectedFrom="under log in tropical forest" collectingDate="1979-10-11" collectorName="C. Murcia &amp; G. Kattan" country="Colombia" elevation="1800" latitude="3.493" location="Cali-Buenaventura" longLatPrecision="78" longitude="-76.613" municipality="Cali" pageId="12" pageNumber="77" specimenCode="ICN-MD-1219" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca">
<collectingCountry id="F372E195FFCE3C7B96D2FE0B3440FE6B" box="[189,344,412,438]" name="Colombia" pageId="12" pageNumber="77">COLOMBIA</collectingCountry>
•
<specimenCount id="9D636A8CFFCE3C7B9715FE0B34B3FE6B" box="[378,427,412,438]" pageId="12" pageNumber="77" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49A16FE7FFCE3C7B97D0FE0B379BFE6B" box="[447,643,412,438]" country="Colombia" name="Valle del Cauca" pageId="12" pageNumber="77">Valle del Cauca</collectingRegion>
,
<collectingMunicipality id="6BBE3B7FFFCE3C7B94FBFE0B37D0FE6B" box="[660,712,412,438]" pageId="12" pageNumber="77">Cali</collectingMunicipality>
,
<quantity id="4C9D0CE0FFCE3C7B94B7FE0B363EFE6B" box="[728,806,412,438]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="77" unit="km" value="16.0">
<locationDeviation id="9A68B827FFCE3C7B94B7FE0B363EFE6B" box="[728,806,412,438]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="77" unit="km" value="16.0">16 km</locationDeviation>
</quantity>
vía
<location id="8EBAF7DEFFCE3C7B950CFE0B315CFE6B" LSID="urn:lsid:plazi:treatment:03CC1013FFC93C7A9708FB2A36ECFDBC:8EBAF7DEFFCE3C7B950CFE0B315CFE6B" box="[867,1092,412,438]" country="Colombia" latitude="3.493" longLatPrecision="78" longitude="-76.613" municipality="Cali" name="Cali-Buenaventura" pageId="12" pageNumber="77" stateProvince="Valle del Cauca">Cali-Buenaventura</location>
; [
<geoCoordinate id="EE51C7C2FFCE3C7B9233FE0B31C8FE6B" box="[1116,1232,412,438]" degrees="3.493" direction="north" orientation="latitude" pageId="12" pageNumber="77" precision="55" value="3.493">3.493° N</geoCoordinate>
,
<geoCoordinate id="EE51C7C2FFCE3C7B928EFE0B3072FE6B" box="[1249,1386,412,438]" degrees="76.613" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="55" value="-76.613">76.613° W</geoCoordinate>
];
<elevation id="00484636FFCE3C7B96D2FE28344EFE04" box="[189,342,447,473]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="m" value="1800.0">
<quantity id="4C9D0CE0FFCE3C7B96D2FE28340EFE04" box="[189,278,447,473]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFDB87C5FFCE3C7B970CFE2834E5FE04" box="[355,509,447,473]" pageId="12" pageNumber="77" value="1979-10-11">
<collectingDate id="EF9F7E2DFFCE3C7B970CFE2834E5FE04" box="[355,509,447,473]" pageId="12" pageNumber="77" value="1979-10-11">11 Oct. 1979</collectingDate>
</date>
;
<collectorName id="2690C4D3FFCE3C7B9466FE283799FE04" box="[521,641,447,473]" pageId="12" pageNumber="77">C. Murcia</collectorName>
and
<collectorName id="2690C4D3FFCE3C7B94D6FE283633FE04" box="[697,811,447,473]" pageId="12" pageNumber="77">G. Kattan</collectorName>
leg.;
<collectedFrom id="3879E2F8FFCE3C7B9502FE2831B2FE04" box="[877,1194,447,473]" pageId="12" pageNumber="77">under log in tropical forest</collectedFrom>
;
<specimenCode id="DBC3097EFFCE3C7B92DAFE28307DFE04" box="[1205,1381,447,473]" pageId="12" pageNumber="77">ICN-MD-1219</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B0DAB58FFCE3C7B96D2FE753142FDC2" ID-GBIF-Occurrence="4167732303" collectedFrom="collected at night at the edge of a road" collectingDate="2014-11-10" collectorName="D. Martinez" country="Colombia" elevation="1900" latitude="3.517" location="Cali-Buenaventura" longLatPrecision="78" longitude="-76.621" municipality="Cali" pageId="12" pageNumber="77" specimenCode="ICN-MD-1367" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca">
<specimenCount id="9D636A8CFFCE3C7B96D2FE7535F0FE20" box="[189,232,482,509]" pageId="12" pageNumber="77" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49A16FE7FFCE3C7B9699FE7534B7FE21" box="[246,431,482,508]" country="Colombia" name="Valle del Cauca" pageId="12" pageNumber="77">Valle del Cauca</collectingRegion>
,
<collectingMunicipality id="6BBE3B7FFFCE3C7B97D5FE7534F6FE21" box="[442,494,482,508]" pageId="12" pageNumber="77">Cali</collectingMunicipality>
,
<quantity id="4C9D0CE0FFCE3C7B9796FE75375AFE21" box="[505,578,482,508]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="km" value="18.0">
<locationDeviation id="9A68B827FFCE3C7B9796FE75375AFE21" box="[505,578,482,508]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="km" value="18.0">18 km</locationDeviation>
</quantity>
vía
<location id="8EBAF7DEFFCE3C7B941CFE75364CFE21" LSID="urn:lsid:plazi:treatment:03CC1013FFC93C7A9708FB2A36ECFDBC:8EBAF7DEFFCE3C7B941CFE75364CFE21" box="[627,852,482,508]" country="Colombia" latitude="3.517" longLatPrecision="78" longitude="-76.621" municipality="Cali" name="Cali-Buenaventura" pageId="12" pageNumber="77" stateProvince="Valle del Cauca">Cali-Buenaventura</location>
; [
<geoCoordinate id="EE51C7C2FFCE3C7B9509FE7536CDFE20" box="[870,981,482,509]" degrees="3.517" direction="north" orientation="latitude" pageId="12" pageNumber="77" precision="55" value="3.517">3.517° N</geoCoordinate>
,
<geoCoordinate id="EE51C7C2FFCE3C7B958EFE75317CFE20" box="[993,1124,482,509]" degrees="76.621" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="55" value="-76.621">76.621° W</geoCoordinate>
];
<elevation id="00484636FFCE3C7B9218FE753016FE21" box="[1143,1294,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="77" unit="m" value="1900.0">
<quantity id="4C9D0CE0FFCE3C7B9218FE7531D7FE20" box="[1143,1231,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="77" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
;
<date id="FFDB87C5FFCE3C7B9374FE7535E3FDC2" pageId="12" pageNumber="77" value="2014-11-10">
<collectingDate id="EF9F7E2DFFCE3C7B9374FE7535E3FDC2" pageId="12" pageNumber="77" value="2014-11-10">10 Nov. 2014</collectingDate>
</date>
;
<collectorName id="2690C4D3FFCE3C7B9768FD91348DFDC2" box="[263,405,517,544]" pageId="12" pageNumber="77">D. Martinez</collectorName>
leg.;
<collectedFrom id="3879E2F8FFCE3C7B97B8FD923683FDC2" box="[471,923,517,544]" pageId="12" pageNumber="77">collected at night at the edge of a road</collectedFrom>
;
<specimenCode id="DBC3097EFFCE3C7B95C9FD923142FDC2" box="[934,1114,517,543]" pageId="12" pageNumber="77">ICN-MD-1367</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFCE3C7A96D2FDD13420FE2F" lastPageId="13" lastPageNumber="78" pageId="12" pageNumber="77" type="description">
<paragraph id="8BDAA105FFCE3C7B96D2FDD13456FDBD" blockId="12.[189,334,582,608]" box="[189,334,582,608]" pageId="12" pageNumber="77">
<heading id="D0921669FFCE3C7B96D2FDD13456FDBD" bold="true" box="[189,334,582,608]" fontSize="11" level="3" pageId="12" pageNumber="77" reason="3">
<emphasis id="B9117D17FFCE3C7B96D2FDD13456FDBD" bold="true" box="[189,334,582,608]" pageId="12" pageNumber="77">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFCE3C7B9685FDE23767FD52" blockId="12.[188,1400,629,937]" box="[234,639,629,655]" pageId="12" pageNumber="77">
<emphasis id="B9117D17FFCE3C7B9685FDE23432FD52" bold="true" box="[234,298,629,655]" pageId="12" pageNumber="77">Male</emphasis>
(
<typeStatus id="54DE1FA7FFCE3C7B9756FDE234BCFD52" box="[313,420,629,655]" pageId="12" pageNumber="77" type="holotype">holotype</typeStatus>
, ICN-MD-1317A)
</paragraph>
<paragraph id="8BDAA105FFCE3C7B96D2FD0F340FFC74" blockId="12.[188,1400,629,937]" pageId="12" pageNumber="77">
With characteristics typical for the genus. Coloration (recently preserved in 70% ethanol): body rings dark, reddish brown (nearly purple); head light brown; legs yellowish ocher; telson reddish, with posterior margin (= spinnerets region) yellowish (
<figureCitation id="135EBD80FFCE3C7B9492FD483659FD24" box="[765,833,735,761]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[378,1032,415,1893]" captionTargetId="figure-14@4.[310,1277,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8205502" httpUri="https://zenodo.org/record/8205502/files/figure.png" pageId="12" pageNumber="77">Fig. 1</figureCitation>
). Antennae broken. Head: epicranial macrosetae 2–2, interantennal macrosetae 1–1, frontal macrosetae 2–2. Sternites: 4
<superScript id="7C100C4DFFCE3C7B9276FC96313EFCD2" attach="left" box="[1049,1062,769,783]" fontSize="6" pageId="12" pageNumber="77">th</superScript>
body ring with two robust, conical, blunt projections, separated along entire length (
<figureCitation id="135EBD80FFCE3C7B9538FCB236ABFCE2" box="[855,947,805,831]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="12" pageNumber="77">Fig. 2B</figureCitation>
); 5
<superScript id="7C100C4DFFCE3C7B95B7FCB336FDFCEF" attach="left" box="[984,997,804,818]" fontSize="6" pageId="12" pageNumber="77">th</superScript>
body ring with two pairs of short, blunt projections, the anterior pair more conspicuous (
<figureCitation id="135EBD80FFCE3C7B955CFCDF3689FCBE" box="[819,913,840,867]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="12" pageNumber="77">Fig. 3D</figureCitation>
). Gonopod aperture on 7
<superScript id="7C100C4DFFCE3C7B92DDFCDF31A7FC8B" attach="left" box="[1202,1215,840,854]" fontSize="6" pageId="12" pageNumber="77">th</superScript>
body ring oval, rim notably projecting above sternite (
<quantity id="4C9D0CE0FFCE3C7B9416FCFB37FFFC5B" box="[633,743,876,902]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="12" pageNumber="77" unit="mm" value="0.67">0.67 mm</quantity>
), with a deep U-shaped concavity (
<quantity id="4C9D0CE0FFCE3C7B9210FCFB31C3FC5B" box="[1151,1243,876,902]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="77" unit="mm" value="0.3">0.3 mm</quantity>
) on posterior margin.
</paragraph>
<paragraph id="8BDAA105FFCE3C7A96D2FC473651FE50" blockId="12.[188,1400,976,1144]" lastBlockId="13.[189,1399,265,397]" lastPageId="13" lastPageNumber="78" pageId="12" pageNumber="77">
<smallCapsWord id="8D3C37D9FFCE3C7B96D2FC47342CFC34" baselines="995,996" box="[189,308,976,1002]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="12" pageNumber="77">GONOPODS</smallCapsWord>
(
<figureCitation id="135EBD80FFCE3C7B9724FC473480FC37" box="[331,408,976,1003]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77">Fig. 7</figureCitation>
). Gonocoxae rounded, equivalent to about half the length of the telopodite; not prominent in ectal view, with two macrosetae on dorsum. Cannula hook-shaped, broken in the
<typeStatus id="54DE1FA7FFCE3C7B9367FC64306BFBD0" box="[1288,1395,1011,1037]" pageId="12" pageNumber="77" type="holotype">holotype</typeStatus>
. Solenomere long, slender, unbranched, curving with distinct ridge at midway point, falciform at apex, tip straight (
<figureCitation id="135EBD80FFCE3C7B9720FBAD348FFB89" box="[335,407,1082,1108]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77">Fig. 7</figureCitation>
, S). Prefemoral process: thin at base, with shelf (
<figureCitation id="135EBD80FFCE3C7B958FFBAD3126FB89" box="[992,1086,1082,1108]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77">Fig. 7A</figureCitation>
, Sh) under linear lobe just before apex (
<figureCitation id="135EBD80FFCE3C7B973AFBCA34A9FBAA" box="[341,433,1117,1143]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77">Fig. 7A</figureCitation>
, Lb); apex with large, tongue-shaped lobe, best seen in anterior view (
<figureCitation id="135EBD80FFCE3C7B92B8FBCA3029FBA5" box="[1239,1329,1117,1144]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77">Fig. 7B</figureCitation>
, PfP).
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<smallCapsWord id="8D3C37D9FFCF3C7A9765FE9934ABFEFC" baselines="284" box="[266,435,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="13" pageNumber="78">MEASUREMENTS</smallCapsWord>
(mm). Total length 29.33, total width 2.10. Collum: length 0.77, width 1.55. Podomere length of 10
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body ring (1–7): 0.29, 0.56, 1.15, 0.52, 0.66, 1.05, 0.11. Gonopod aperture: length 0.50, width 0.86. Gonopod: length 1.21, width 0.96. Gonocoxae: length 0.54, width 0.59. Telopodite: length 1.19, width 0.42. Telson length 0.98.
</paragraph>
<caption id="DF1AF18DFFCE3C7B96D3F8F33424F835" ID-DOI="http://doi.org/10.5281/zenodo.8205514" ID-Zenodo-Dep="8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="12" pageNumber="77" startId="12.[188,231,1892,1918]" targetBox="[190,1398,1234,1849]" targetPageId="12">
<paragraph id="8BDAA105FFCE3C7B96D3F8F33424F835" blockId="12.[188,1399,1892,2025]" pageId="12" pageNumber="77">
<emphasis id="B9117D17FFCE3C7B96D3F8F33414F8A2" bold="true" box="[188,268,1892,1919]" pageId="12" pageNumber="77">Fig. 7.</emphasis>
<taxonomicName id="4C65DA86FFCE3C7B9775F8F2313CF8A2" authority="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityName="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityYear="2023" box="[282,1060,1893,1919]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="77" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFCE3C7B9775F8F2370FF8A2" box="[282,535,1893,1919]" italics="true" pageId="12" pageNumber="77">Dibolostethus kattani</emphasis>
Means, Bouzan, Martínez-Torres &amp; Ivanov
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFCE3C7B9241F8F23195F8A2" box="[1070,1165,1893,1919]" pageId="12" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ICN- MD-1317A), right gonopod.
<emphasis id="B9117D17FFCE3C7B947EF81F373EF87F" bold="true" box="[529,550,1928,1954]" pageId="12" pageNumber="77">A</emphasis>
. Mesal view.
<emphasis id="B9117D17FFCE3C7B94A6F81F37C5F87F" bold="true" box="[713,733,1928,1954]" pageId="12" pageNumber="77">B</emphasis>
. Anterior view.
<emphasis id="B9117D17FFCE3C7B95F4F81F36A8F87F" bold="true" box="[923,944,1928,1954]" pageId="12" pageNumber="77">C</emphasis>
. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf.
</paragraph>
</caption>
<paragraph id="8BDAA105FFCF3C7A9685FE23345DFE13" blockId="13.[189,325,436,498]" box="[234,325,436,462]" pageId="13" pageNumber="78">
<emphasis id="B9117D17FFCF3C7A9685FE23345DFE13" bold="true" box="[234,325,436,462]" pageId="13" pageNumber="78">Female</emphasis>
</paragraph>
<paragraph id="8BDAA105FFCF3C7A96D2FE403420FE2F" blockId="13.[189,325,436,498]" box="[189,312,471,498]" pageId="13" pageNumber="78">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFCF3C7A96D2FD8F36ECFDBC" box="[189,1012,536,610]" pageId="13" pageNumber="78" type="distribution">
<paragraph id="8BDAA105FFCF3C7A96D2FD8F344EFDEF" blockId="13.[189,342,536,562]" box="[189,342,536,562]" pageId="13" pageNumber="78">
<heading id="D0921669FFCF3C7A96D2FD8F344EFDEF" bold="true" box="[189,342,536,562]" fontSize="11" level="3" pageId="13" pageNumber="78" reason="3">
<emphasis id="B9117D17FFCF3C7A96D2FD8F344EFDEF" bold="true" box="[189,342,536,562]" pageId="13" pageNumber="78">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFCF3C7A96D2FDD036ECFDBC" blockId="13.[189,1012,583,610]" box="[189,1012,583,610]" pageId="13" pageNumber="78">
Only known from the
<collectingRegion id="49A16FE7FFCF3C7A97AEFDD03617FDBF" box="[449,783,583,610]" country="Colombia" name="Valle del Cauca" pageId="13" pageNumber="78">Valle del Cauca Department</collectingRegion>
,
<collectingCountry id="F372E195FFCF3C7A9575FDD03696FDBC" box="[794,910,583,609]" name="Colombia" pageId="13" pageNumber="78">Colombia</collectingCountry>
(
<figureCitation id="135EBD80FFCF3C7A95F1FDD036FDFDBC" box="[926,997,583,609]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="13" pageNumber="78">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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