treatments-xml/data/03/C8/D1/03C8D14B8C397507FF1B1CACFECE4A7E.xml
2024-06-21 12:22:17 +02:00

217 lines
27 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="09928BC8DCC9D8727597B79576801CF1" ID-CLB-Dataset="29962" ID-DOI="10.11646/zootaxa.4429.2.8" ID-GBIF-Dataset="796cc5ee-d357-4cd5-a7df-6bbaf52d0d8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1283817" ID-ZooBank="7AC25C78-BED6-471A-BEFF-88C657667540" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528271858807" checkinUser="plazi" docAuthor="Tabell, Jukka, Mutanen, Marko &amp; Sihvonen, Pasi" docDate="2018" docId="03C8D14B8C397507FF1B1CACFECE4A7E" docLanguage="en" docName="zootaxa.4429.2.8.pdf" docOrigin="Zootaxa 4429 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleophora embaensis Tabell &amp; Mutanen &amp; Sihvonen 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FFF1A9338C31750DFF8C1F08FFCB496A" masterDocTitle="Descriptions of five morphologically and genetically confirmed new species of the Coleophora poecilella Walsingham, 1907 species group (Lepidoptera, Coleophoridae) from the Palearctic Region" masterLastPageNumber="347" masterPageNumber="331" pageNumber="339" updateTime="1698547121334" updateUser="ExternalLinkService">
<mods:mods id="D855107D07600F8FD96AA6925525E766" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="341231D089B2790049D948E5773EB9D8">
<mods:title id="1287F888C5F41CD4ADC0246C4A712CE5">Descriptions of five morphologically and genetically confirmed new species of the Coleophora poecilella Walsingham, 1907 species group (Lepidoptera, Coleophoridae) from the Palearctic Region</mods:title>
</mods:titleInfo>
<mods:name id="665C6D461E0F6BECEFDB45AEA33DF548" type="personal">
<mods:role id="671DC3EE56E53978F79657C5AD2FE364">
<mods:roleTerm id="790A3A88912864405EB47C49DCEF10C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9298645400AF3E237657441062CB7F10">Tabell, Jukka</mods:namePart>
</mods:name>
<mods:name id="10D64BD4F0885C72353EBFBD15998C8F" type="personal">
<mods:role id="272091076F4B60375834BD5A2EFC89CA">
<mods:roleTerm id="79C56B2A3453717B550AEE897B7D6D2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE8F3FA7EBF19D0FBF94E104D1F818D2">Mutanen, Marko</mods:namePart>
</mods:name>
<mods:name id="F152520F1DB80082678A697EBF51E777" type="personal">
<mods:role id="85F2EB434D72B43FC5385F1254763DB0">
<mods:roleTerm id="04853833CC6FAFF8426F4BD0EE11AC8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="277D0ACBEE2179DE8C14052A9E89CB6B">Sihvonen, Pasi</mods:namePart>
</mods:name>
<mods:typeOfResource id="E70299FCF8524D4A603EF53B9C1F4E94">text</mods:typeOfResource>
<mods:relatedItem id="57EE57F3345589D33BD83C30B64D08F7" type="host">
<mods:titleInfo id="E807F7F6FC05A807E213B7565EBC771B">
<mods:title id="0C6DC7707A7AA522F7386988436729F4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6A6490251DB6D398CF9EB707D0D2E868">
<mods:date id="9DDA539E63BE424022D764FC3459A6A2">2018</mods:date>
<mods:detail id="0464523DC2EA78728A4F531EA0118269" type="pubDate">
<mods:number id="5E8DFA8DEA47196C91F67449A2ADCC2B">2018-06-06</mods:number>
</mods:detail>
<mods:detail id="ACA98FBD7C691DCFA8B8A1095F66C47C" type="volume">
<mods:number id="67929066B407B6086DCD7D4952ED1E21">4429</mods:number>
</mods:detail>
<mods:detail id="18FF28ED173394EFAF29D0BD1BBE84F6" type="issue">
<mods:number id="625447437B1C55FDEC9E5FC46D7C6248">2</mods:number>
</mods:detail>
<mods:extent id="B42237C31F91EED65A7288B5F023D76C" unit="page">
<mods:start id="53B70D6496CCA0182D75A7EE4D346FA6">331</mods:start>
<mods:end id="CF7279A68334A59471E68B1B16295AC1">347</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ADB222C3F20FFA78CDA50A7880DF19CF">journal article</mods:classification>
<mods:identifier id="F5A7F9203F6DE20C558E35132FB367FC" type="CLB-Dataset">29962</mods:identifier>
<mods:identifier id="2C528714FCAE89F70CF25B525C89D17A" type="DOI">10.11646/zootaxa.4429.2.8</mods:identifier>
<mods:identifier id="8370B86548760745BD8DA930C707DAD9" type="GBIF-Dataset">796cc5ee-d357-4cd5-a7df-6bbaf52d0d8d</mods:identifier>
<mods:identifier id="4337212663F623989571F00FB79CD35E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EB065DCD16D0EC395FBCB601423C6D2A" type="Zenodo-Dep">1283817</mods:identifier>
<mods:identifier id="2082C0ABA004B8FC3C1D9EE7B4D6FE49" type="ZooBank">7AC25C78-BED6-471A-BEFF-88C657667540</mods:identifier>
</mods:mods>
<treatment id="03C8D14B8C397507FF1B1CACFECE4A7E" ID-DOI="http://doi.org/10.5281/zenodo.5985234" ID-GBIF-Taxon="145907887" ID-Zenodo-Dep="5985234" LSID="urn:lsid:plazi:treatment:03C8D14B8C397507FF1B1CACFECE4A7E" httpUri="http://treatment.plazi.org/id/03C8D14B8C397507FF1B1CACFECE4A7E" lastPageId="10" lastPageNumber="341" pageId="8" pageNumber="339">
<subSubSection id="C37B33D68C397505FF1B1CACFDBA4A8A" pageId="8" pageNumber="339" type="nomenclature">
<paragraph id="8BDE605D8C397505FF1B1CACFDA14AD5" blockId="8.[151,625,932,1029]" box="[151,618,932,959]" pageId="8" pageNumber="339">
<heading id="D096D7318C397505FF1B1CACFDA14AD5" bold="true" box="[151,618,932,959]" fontSize="11" level="1" pageId="8" pageNumber="339" reason="1">
<emphasis id="B915BC4F8C397505FF1B1CACFDA14AD5" bold="true" box="[151,618,932,959]" pageId="8" pageNumber="339">
<taxonomicName id="4C611BDE8C397505FF1B1CACFE344AD5" authority="Tabell" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[151,511,932,959]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis" status="sp. nov.">
<emphasis id="B915BC4F8C397505FF1B1CACFE624AD4" bold="true" box="[151,425,932,958]" italics="true" pageId="8" pageNumber="339">Coleophora embaensis</emphasis>
Tabell
</taxonomicName>
,
<taxonomicNameLabel id="A22601348C397505FD801CADFDA14AD5" box="[524,618,933,959]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE605D8C397505FF1B1CCFFDBA4A8A" blockId="8.[151,625,932,1029]" box="[151,625,967,992]" pageId="8" pageNumber="339">Barcode Index Number: BOLD:ACT3652</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C397505FF1B1CE4FE024D6E" box="[151,457,1004,1029]" pageId="8" pageNumber="339" type="description">
<paragraph id="8BDE605D8C397505FF1B1CE4FE024D6E" blockId="8.[151,625,932,1029]" box="[151,457,1004,1029]" pageId="8" pageNumber="339">
<figureCitation id="135A7CD88C397505FF1B1CE4FF314D6E" box="[151,250,1004,1028]" captionStart="FIGURES 18" captionStartId="2.[151,264,1901,1924]" captionTargetBox="[152,1435,217,1853]" captionTargetId="figure@2.[151,1436,217,1853]" captionTargetPageId="2" captionText="FIGURES 18. Adults and larval cases of Coleophora spp. 16. Adults. 1. C. mirleftensis sp. nov., ♂ holotype. 2. C. nupponeni sp. nov., ♂ holotype. 3. C. embaensis sp. nov., ♂ paratype. 4. C. embaensis sp. nov., ♀ paratype. 5. C. charynensis sp. nov., ♂ holotype. 6. C. trichopterella Baldizzone, ♀. 78. Larval cases. 7. C. mirleftensis sp. nov. 8. C. trichopterella Baldizzone." httpUri="https://zenodo.org/record/1283819/files/figure.png" pageId="8" pageNumber="339">Figs. 3,4</figureCitation>
,
<figureCitation id="135A7CD88C397505FE8A1CE4FEDD4D6E" box="[262,278,1004,1028]" captionStart="FIGURE 9" captionStartId="4.[151,250,1987,2009]" captionTargetBox="[203,1358,259,1904]" captionTargetId="figure@4.[160,1427,247,1921]" captionTargetPageId="4" captionText="FIGURE 9. Neighbor-joining tree of the barcoded taxa of the C. poecilella species group, based on sequences of COI gene." httpUri="https://zenodo.org/record/1283821/files/figure.png" pageId="8" pageNumber="339">9</figureCitation>
,
<figureCitation id="135A7CD88C397505FEAE1CE4FEA04D6E" box="[290,363,1004,1028]" captionStart="FIGURES 1821" captionStartId="9.[151,264,1965,1988]" captionTargetBox="[209,1379,193,1948]" captionTargetId="figure@9.[208,1379,193,1948]" captionTargetPageId="9" captionText="FIGURES 1821. C. embaensis sp. nov. 18. ♂ genitalia, GP 4915 J. Tabell. 19. ♂ abdomen. 20. ♀ abdomen. 21. ♀ genitalia, GP 5361 J. Tabell." httpUri="https://zenodo.org/record/1283827/files/figure.png" pageId="8" pageNumber="339">1821</figureCitation>
, Table 1
</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C397505FF1B1B3CFE0C4C06" pageId="8" pageNumber="339" type="materials_examined">
<paragraph id="8BDE605D8C397505FF1B1B3CFE0C4C06" blockId="8.[151,1438,1076,2037]" pageId="8" pageNumber="339">
<emphasis id="B915BC4F8C397505FF1B1B3CFE814D27" bold="true" box="[151,330,1076,1101]" pageId="8" pageNumber="339">Type material.</emphasis>
<materialsCitation id="3B096A008C397505FEDA1B3CFBBC4D05" ID-GBIF-Occurrence="1848506014" collectingDate="2012-04-24" collectionCode="GP, DNA, NUPP" collectorName="K. Nupponen" country="Kazakhstan" elevation="280" latitude="48.924725" location="Emba vill. 17 km NE" longLatPrecision="18" longitude="58.31361" pageId="8" pageNumber="339" specimenCode="4968" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54DADEFF8C397505FEDA1B3CFE754D26" box="[342,446,1076,1100]" pageId="8" pageNumber="339" type="holotype">Holotype</typeStatus>
<specimenCount id="9D67ABD48C397505FE471B3DFE2B4D26" box="[459,480,1077,1100]" pageId="8" pageNumber="339" type="male"></specimenCount>
(
<collectionCode id="ED70F8988C397505FE781B3CFDD24D26" box="[500,537,1076,1100]" pageId="8" pageNumber="339">GP</collectionCode>
<specimenCode id="DBC7C8268C397505FDAA1B3CFD954D27" box="[550,606,1076,1101]" pageId="8" pageNumber="339">4968</specimenCode>
J. Tabell,
<collectionCode id="ED70F8988C397505FD6D1B3CFCD54D26" box="[737,798,1076,1100]" pageId="8" pageNumber="339">DNA</collectionCode>
sample 23575 Lepid Phyl):
<collectingCountry id="F37620CD8C397505FBFB1B3CFB354D27" box="[1143,1278,1076,1101]" name="Kazakhstan" pageId="8" pageNumber="339">Kazakhstan</collectingCountry>
,
<geoCoordinate id="EE55069A8C397505FA811B3CFA574D27" box="[1293,1436,1076,1101]" direction="north" orientation="latitude" pageId="8" pageNumber="339" precision="15" value="48.924725">48°55´29´´N</geoCoordinate>
<geoCoordinate id="EE55069A8C397505FF1B1B5FFEED4D1A" box="[151,294,1111,1136]" direction="east" orientation="longitude" pageId="8" pageNumber="339" precision="15" value="58.31361">58°18´49´´E</geoCoordinate>
,
<location id="8EBE36868C397505FEBE1B5FFDE24D05" LSID="urn:lsid:plazi:treatment:03C8D14B8C397507FF1B1CACFECE4A7E:8EBE36868C397505FEBE1B5FFDE24D05" box="[306,553,1111,1136]" country="Kazakhstan" latitude="48.924725" longLatPrecision="18" longitude="58.31361" name="Emba vill. 17 km NE" pageId="8" pageNumber="339">
Emba vill.
<quantity id="4C99CDB88C397505FE3D1B50FE324D1A" box="[433,505,1111,1136]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="339" unit="km" value="17.0">17 km</quantity>
NE
</location>
,
<elevation id="004C876E8C397505FDB91B50FD4B4D05" box="[565,640,1112,1136]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="339" unit="m" value="280.0">
<quantity id="4C99CDB88C397505FDB91B50FD4B4D05" box="[565,640,1112,1136]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="339" unit="m" value="280.0">280 m</quantity>
</elevation>
,
<collectingDate id="EF9BBF758C397505FD011B50FCC74D1A" box="[653,780,1112,1136]" pageId="8" pageNumber="339" value="2012-04-24">
<date id="FFDF469D8C397505FD011B50FCC74D1A" box="[653,780,1112,1136]" pageId="8" pageNumber="339" value="2012-04-24">24.IV.2012</date>
</collectingDate>
,
<collectorName id="2694058B8C397505FC941B50FC7A4D1A" box="[792,945,1112,1136]" pageId="8" pageNumber="339">K. Nupponen</collectorName>
leg., coll.
<collectionCode id="ED70F8988C397505FBA61B50FBB94D05" box="[1066,1138,1112,1135]" collectionName="NUPP" pageId="8" pageNumber="339">NUPP</collectionCode>
.
</materialsCitation>
Paratypes: 5 ♂, 2 ♀: 1 ♂ (DNA sample 23618 Lepid Phyl), 1 ♀ (GP 5361 J. Tabell) same collecting data as in holotype, colls. NUPP and TAB; 1 ♂ (GP 4915 J. Tabell, DNA sample 21786 Lepid Phyl) Kazakhstan, 47°12´02´´N 55°29´13´´E, Emba river, 45 m, Besbai village 2 km E, 11.V.2011, K. Nupponen leg., coll. NUPP; 2 ♂ same collecting data, but 19.IV.2013, coll. NUPP; 1 ♂ (GP 5161 J. Tabell, DNA sample 23570 Lepid Phyl) SW Kazakhstan, 44°01´04´´N 53°25´46´´E, 240 m, Sai-Utes vill. 40 km S, Syndy, 6.IV.2013, K. Nupponen leg., coll. TAB; 1 ♀ (DNA sample 21860 Lepid Phyl) Kazakhstan, 47°27´24´´N 55°06´45´´E, Aktolagai chalk hills, 95 m, Emba river 40 km NW, 15-V-2010, K. Nupponen leg, coll. NUPP.
</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C397505FF4B1A7FFE5E4FBE" pageId="8" pageNumber="339" type="diagnosis">
<paragraph id="8BDE605D8C397505FF4B1A7FFDC54F2E" blockId="8.[151,1438,1076,2037]" pageId="8" pageNumber="339">
<emphasis id="B915BC4F8C397505FF4B1A7FFE8A4CFA" bold="true" box="[199,321,1399,1424]" pageId="8" pageNumber="339">Diagnosis.</emphasis>
In genitalia
<taxonomicName id="4C611BDE8C397505FE5D1A70FDA34CFA" authority="Tabell &amp; Mutanen &amp; Sihvonen, 2018" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[465,616,1400,1424]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis" status="sp. nov.">
<emphasis id="B915BC4F8C397505FE5D1A70FDA34CFA" box="[465,616,1400,1424]" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22601348C397505FDE31A7FFD004CE5" box="[623,715,1399,1424]" pageId="8" pageNumber="339" rank="species">
<emphasis id="B915BC4F8C397505FDE31A7FFD004CE5" bold="true" box="[623,715,1399,1424]" pageId="8" pageNumber="339">sp. nov.</emphasis>
</taxonomicNameLabel>
is close to
<taxonomicName id="4C611BDE8C397505FCC11A70FBAD4CFA" authority="Toll, 1961" authorityName="Toll" authorityYear="1961" box="[845,1126,1399,1424]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="ochroflava">
<emphasis id="B915BC4F8C397505FCC11A70FC2C4CFA" box="[845,999,1400,1424]" italics="true" pageId="8" pageNumber="339">C. ochroflava</emphasis>
Toll, 1961
</taxonomicName>
. In the male genitalia of
<taxonomicName id="4C611BDE8C397505FA0E1A70FEC74CDE" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis">
<emphasis id="B915BC4F8C397505FA0E1A70FEC74CDE" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
cornuti are absent, the valvula is rounded and the cucullus is conical, whereas in
<taxonomicName id="4C611BDE8C397505FB5E1A94FAA44CDE" authorityName="Toll" authorityYear="1961" box="[1234,1391,1436,1460]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="ochroflava">
<emphasis id="B915BC4F8C397505FB5E1A94FAA44CDE" box="[1234,1391,1436,1460]" italics="true" pageId="8" pageNumber="339">C. ochroflava</emphasis>
</taxonomicName>
the cornuti are grouped into robust bundle, the valvula is triangular and the cucullus is parallel-sided. In the female genitalia of
<taxonomicName id="4C611BDE8C397505FE921AECFE7E4C96" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[286,437,1508,1532]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis">
<emphasis id="B915BC4F8C397505FE921AECFE7E4C96" box="[286,437,1508,1532]" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
the sclerotization of the sterigma in the proximal part is larger, the apophyses anteriores are thicker and the ductus bursae has two twists and several conical dark spines; in
<taxonomicName id="4C611BDE8C397505FBB61900FB184F4A" authorityName="Toll" authorityYear="1961" box="[1082,1235,1544,1568]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="ochroflava">
<emphasis id="B915BC4F8C397505FBB61900FB864F75" box="[1082,1101,1544,1567]" italics="true" pageId="8" pageNumber="339">C</emphasis>
.
<emphasis id="B915BC4F8C397505FBD71900FB184F4A" box="[1115,1235,1544,1568]" italics="true" pageId="8" pageNumber="339">ochroflava</emphasis>
</taxonomicName>
the ductus bursae is straight without conical spines.
</paragraph>
<paragraph id="8BDE605D8C397505FF4B1947FE5E4FBE" blockId="8.[151,1438,1076,2037]" pageId="8" pageNumber="339">
<emphasis id="B915BC4F8C397505FF4B1947FE094F02" bold="true" box="[199,450,1615,1640]" pageId="8" pageNumber="339">Molecular diagnosis.</emphasis>
Five specimens of
<taxonomicName id="4C611BDE8C397505FD251958FC894F02" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[681,834,1616,1640]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis">
<emphasis id="B915BC4F8C397505FD251958FC894F02" box="[681,834,1616,1640]" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
were sequenced, resulting in 658 bp (n=2), 636 bp (n=2) and 611 bp (n=1) DNA barcode fragments. The resultant sequences revealed 7.74-12.67 % divergences between
<taxonomicName id="4C611BDE8C397505FF701990FE5F4FDA" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[252,404,1688,1712]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis">
<emphasis id="B915BC4F8C397505FF701990FE5F4FDA" box="[252,404,1688,1712]" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
and the other barcoded species (Table 1). The barcodes of
<taxonomicName id="4C611BDE8C397505FBB41990FB044FDA" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[1080,1231,1688,1712]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="339" phylum="Tracheophyta" rank="species" species="embaensis">
<emphasis id="B915BC4F8C397505FBB41990FB044FDA" box="[1080,1231,1688,1712]" italics="true" pageId="8" pageNumber="339">C. embaensis</emphasis>
</taxonomicName>
displayed 0.16 % intraspecific variation.
</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C397507FF4B19D7FBF74B57" lastPageId="10" lastPageNumber="341" pageId="8" pageNumber="339" type="description">
<paragraph id="8BDE605D8C397505FF4B19D7FB494EE2" blockId="8.[151,1438,1076,2037]" pageId="8" pageNumber="339">
<emphasis id="B915BC4F8C397505FF4B19D7FE924F92" bold="true" box="[199,345,1759,1784]" pageId="8" pageNumber="339">Description.</emphasis>
Wingspan 911 mm. Head pale fuscous. Antenna white, annulated with pale brown. Scape pale fuscous. Labial palpus pale fuscous mixed with white; tufted second segment 1.5 x longer than third segment. Thorax pale fuscous; tegula mixed with whitish grey scales. Forewing mottled, creamy white, densely covered with ochreous brown-tipped, pale fuscous scales, in female scales paler and more sparse, fringe whitish grey to pale brown. Hindwing pale grey, fringe whitish grey. Abdomen whitish grey, slightly lustrous.
</paragraph>
<paragraph id="8BDE605D8C397505FF4B189CFC154E9E" blockId="8.[151,1438,1076,2037]" pageId="8" pageNumber="339">Abdominal structures. Latero-posterior bar short, 2/52/7x length of latero-anterior one. Transverse bar straight, proximal edge thin, medially slightly convex, distal edge thick, medially strongly constricted. Tergal sclerite 2.53.5x longer than wide, covered with 40 conical spines (on T3).</paragraph>
<caption id="DF1E30D58C387504FF1B18A5FE984E89" httpUri="https://zenodo.org/record/1283827/files/figure.png" pageId="9" pageNumber="340" startId="9.[151,264,1965,1988]" targetBox="[209,1379,193,1948]" targetPageId="9">
<paragraph id="8BDE605D8C387504FF1B18A5FE984E89" blockId="9.[151,1436,1965,2019]" pageId="9" pageNumber="340">
<emphasis id="B915BC4F8C387504FF1B18A5FE9F4EA9" bold="true" box="[151,340,1965,1988]" pageId="9" pageNumber="340">FIGURES 1821.</emphasis>
<taxonomicName id="4C611BDE8C387504FED718A7FE284EAE" authority="Tabell &amp; Mutanen &amp; Sihvonen, 2018" authorityName="Tabell &amp; Mutanen &amp; Sihvonen" authorityYear="2018" box="[347,483,1966,1988]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="340" phylum="Tracheophyta" rank="species" species="embaensis" status="sp. nov.">
<emphasis id="B915BC4F8C387504FED718A7FE284EAE" box="[347,483,1966,1988]" italics="true" pageId="9" pageNumber="340">C. embaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22601348C387504FE6518A7FDF14EAE" box="[489,570,1967,1988]" pageId="9" pageNumber="340" rank="species">
<emphasis id="B915BC4F8C387504FE6518A7FDF14EAE" bold="true" box="[489,570,1967,1988]" pageId="9" pageNumber="340">sp. nov.</emphasis>
</taxonomicNameLabel>
18. ♂ genitalia, GP 4915 J. Tabell. 19. ♂ abdomen. 20. ♀ abdomen. 21. ♀ genitalia, GP 5361 J. Tabell.
</paragraph>
</caption>
<paragraph id="8BDE605D8C3B7507FF4B1F90FC20482A" blockId="10.[151,1438,151,788]" pageId="10" pageNumber="341">Male genitalia. Gnathos knob transversely oval, broad. Tegumen conical, pedunculus short. Transtilla transversely straight, stubby. Cucullus conical. Costa basally slightly bulged. Valvula narrower than cucullus, outer margin distinctly delineated and evenly rounded, lined with several bristles. Sacculus simple; ventral margin slightly convex, terminating in acute angle, dorsal margin slightly concave, horizontal. Phallotheca narrow, well sclerotized, tubular, apical half conical. Vesica membranous, cornuti absent.</paragraph>
<paragraph id="8BDE605D8C3B7507FF4B1E44FBF74B57" blockId="10.[151,1438,151,788]" pageId="10" pageNumber="341">Female genitalia. Papillae anales conical, covered with long bristles. Anterior apophyses thick, half as long as sterigma, posterior apophyses twice as long as anterior ones. Sterigma quadrate, distal part covered with bristles; proximal margin concave; caudal margin rounded; medial excavation as long as sterigma, distally trough-shaped, proximally rounded. Ostium bursae and colliculum membranous. Ductus bursae transparent; posterior section twisted once, with a medial sclerotized strip about as long as anterior apophysis; section around ductus seminalis twisted, with about 40 small, conical spines; anterior section with one large coil and scattered minute spinules. Corpus bursae oval. Signum elongate, fluke-shaped, spine long with serrated ridge.</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C3B7507FF4B1D4FFA5C4BC2" pageId="10" pageNumber="341" type="biology_ecology">
<paragraph id="8BDE605D8C3B7507FF4B1D4FFA5C4BC2" blockId="10.[151,1438,151,788]" pageId="10" pageNumber="341">
<emphasis id="B915BC4F8C3B7507FF4B1D4FFEEC4B0A" bold="true" box="[199,295,583,608]" pageId="10" pageNumber="341">Biology.</emphasis>
The immature stages and larval host plant are unknown.The species inhabits chalk steppes and cliffs in western Kazakhstan. The habitat of the holotype and seven paratypes is illustrated by
<bibRefCitation id="EFF01DAC8C3B7507FB0D1D64FF284BC2" author="Nupponen, K." journalOrPublisher="SHILAP Revista de lepidopterologia" pageId="10" pageNumber="341" pagination="237 - 255" part="42" publicationUrl="http://www.redalyc.org" refId="ref6388" refString="Nupponen, K. (2014) Notes on Scythrididae observed during 2012 from southern and western Kazakhstan, with descriptions of six new species. SHILAP Revista de lepidopterologia, 42 (166), 237 - 255. Available from: http: // www. redalyc. org / articulo. oa? id = 45532157004 (Accessed 5 Jun. 2018)" title="Notes on Scythrididae observed during 2012 from southern and western Kazakhstan, with descriptions of six new species" type="journal article" year="2014">
Nupponen (2014: p. 250,
<figureCitation id="135A7CD88C3B7507FF1B1D98FF1C4BC2" box="[151,215,656,680]" captionStart="FIGURES 18" captionStartId="2.[151,264,1901,1924]" captionTargetBox="[152,1435,217,1853]" captionTargetId="figure@2.[151,1436,217,1853]" captionTargetPageId="2" captionText="FIGURES 18. Adults and larval cases of Coleophora spp. 16. Adults. 1. C. mirleftensis sp. nov., ♂ holotype. 2. C. nupponeni sp. nov., ♂ holotype. 3. C. embaensis sp. nov., ♂ paratype. 4. C. embaensis sp. nov., ♀ paratype. 5. C. charynensis sp. nov., ♂ holotype. 6. C. trichopterella Baldizzone, ♀. 78. Larval cases. 7. C. mirleftensis sp. nov. 8. C. trichopterella Baldizzone." httpUri="https://zenodo.org/record/1283819/files/figure.png" pageId="10" pageNumber="341">Fig. 1</figureCitation>
)
</bibRefCitation>
. The flight period starts in early April and extends into mid-May. The moth is nocturnal and comes to light.
</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C3B7507FF4B1DBCFABC4BA7" box="[199,1399,692,717]" pageId="10" pageNumber="341" type="distribution">
<paragraph id="8BDE605D8C3B7507FF4B1DBCFABC4BA7" blockId="10.[151,1438,151,788]" box="[199,1399,692,717]" pageId="10" pageNumber="341">
<emphasis id="B915BC4F8C3B7507FF4B1DBCFEAB4BA7" bold="true" box="[199,352,692,717]" pageId="10" pageNumber="341">Distribution.</emphasis>
So far the new taxon is known from few localities in western and south-western Kazakhstan.
</paragraph>
</subSubSection>
<subSubSection id="C37B33D68C3B7507FF4B1DDFFECE4A7E" pageId="10" pageNumber="341" type="etymology">
<paragraph id="8BDE605D8C3B7507FF4B1DDFFECE4A7E" blockId="10.[151,1438,151,788]" pageId="10" pageNumber="341">
<emphasis id="B915BC4F8C3B7507FF4B1DDFFE7E4B9A" bold="true" box="[199,437,727,752]" pageId="10" pageNumber="341">Derivation of name.</emphasis>
The specific name refers to the river Emba, which flows in the vicinity of the collecting localities.
</paragraph>
</subSubSection>
</treatment>
</document>