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<document id="29DA62C1F94A1542EBC1742119D5DE99" ID-DOI="10.5281/zenodo.180818" ID-GBIF-Dataset="850fc9b3-67a6-4ced-b77b-3cc2bebe8857" ID-ISSN="1175-5326" ID-Zenodo-Dep="180818" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816636239" checkinUser="plazi" docAuthor="Ivie, Michael A. &amp; Philips, Keith" docDate="2008" docId="03BC87DCAF3CFFB3FF68FA12FB114909" docLanguage="en" docName="zt01701p014.pdf" docOrigin="Zootaxa 1701" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Canthonella jarmilae Ivie &amp; Philips, new species" docType="treatment" docVersion="8" lastPageNumber="3" masterDocId="FF85FFA4AF3DFFB1FFFFFFE8FFEC4D74" masterDocTitle="Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1698221501712" updateUser="plazi">
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<mods:title id="F4151C89DFAD0F4D98CB691CA1E76F2B">Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="E99F4E35AB982BE2CF2926C1457BEAD4" type="personal">
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<mods:namePart id="94855830F7866A1BD203A7A2FD9A6DE5">Ivie, Michael A.</mods:namePart>
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<mods:namePart id="B449C7368CCF5A31C94E433321169105">Philips, Keith</mods:namePart>
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<mods:title id="B23575D9A1E285F0ADDEACCB0A275B5F">Zootaxa</mods:title>
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<mods:date id="867F0E3EF9B1465C6FB69B311CAA8025">2008</mods:date>
<mods:detail id="E561EA722F89C3E2B2CCCF09384D728A" type="volume">
<mods:number id="ACF648213335AC7D4747238EC696E76E">1701</mods:number>
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<mods:classification id="6501C42984C65E88A64A32F224D11D7C">journal article</mods:classification>
<mods:identifier id="0AC254D101E6D72B3538611D605A36C7" type="DOI">10.5281/zenodo.180818</mods:identifier>
<mods:identifier id="C28DD8904D76FAC5A1D349ADBA22D176" type="GBIF-Dataset">850fc9b3-67a6-4ced-b77b-3cc2bebe8857</mods:identifier>
<mods:identifier id="133137063BD0913999E47BCA2BBAB9D3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BC87DCAF3CFFB3FF68FA12FB114909" ID-DOI="http://doi.org/10.5281/zenodo.6233219" ID-GBIF-Taxon="119354875" ID-Zenodo-Dep="6233219" LSID="urn:lsid:plazi:treatment:03BC87DCAF3CFFB3FF68FA12FB114909" httpUri="http://treatment.plazi.org/id/03BC87DCAF3CFFB3FF68FA12FB114909" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="C30F6541AF3CFFB0FF68FA12FE3C4B43" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BAA36CAAF3CFFB0FF68FA12FD074B60" blockId="1.[151,747,1530,1591]" box="[151,747,1530,1556]" pageId="1" pageNumber="2">
<heading id="D0E281A6AF3CFFB0FF68FA12FD074B60" bold="true" box="[151,747,1530,1556]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B961EAD8AF3CFFB0FF68FA12FD074B60" bold="true" box="[151,747,1530,1556]" pageId="1" pageNumber="2">
<taxonomicName id="4C154D49AF3CFFB0FF68FA12FDA24B60" ID-CoL="QPFP" authority="Ivie &amp; Philips" authorityName="Ivie &amp; Philips" box="[151,590,1530,1556]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jarmilae" status="sp. nov.">
<emphasis id="B961EAD8AF3CFFB0FF68FA12FE754B60" bold="true" box="[151,409,1530,1556]" italics="true" pageId="1" pageNumber="2">Canthonella jarmilae</emphasis>
Ivie &amp; Philips
</taxonomicName>
,
<taxonomicNameLabel id="A25257A3AF3CFFB0FDA4FA12FD074B60" box="[603,747,1530,1556]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36CAAF3CFFB0FF68F9F5FE3C4B43" blockId="1.[151,747,1530,1591]" box="[151,464,1565,1591]" pageId="1" pageNumber="2">
<figureCitation id="132E2A4FAF3CFFB0FF68F9F5FECF4B43" box="[151,291,1565,1591]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="1" pageNumber="2">Figs. 1, 4, 7</figureCitation>
, 10, 13, 16, 19
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF3CFFB0FF68F985FB494A03" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BAA36CAAF3CFFB0FF68F985FB494A03" blockId="1.[151,1436,1645,2034]" pageId="1" pageNumber="2">
<emphasis id="B961EAD8AF3CFFB0FF68F985FEFB4BF3" bold="true" box="[151,279,1645,1671]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
The distinct contrasting color pattern on the elytron and pronotum, as well as the shape of the elytral spot (
<figureCitation id="132E2A4FAF3CFFB0FEF4F97DFE8E4BDB" box="[267,354,1685,1711]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="1" pageNumber="2">Fig. 19</figureCitation>
) makes this species very easily distinguished from the other species of
<taxonomicName id="4C154D49AF3CFFB0FB39F97DFABB4BDA" box="[1222,1367,1685,1710]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD8AF3CFFB0FB39F97DFABB4BDA" box="[1222,1367,1685,1710]" italics="true" pageId="1" pageNumber="2">Canthonella</emphasis>
</taxonomicName>
. This species is unique in usually having the pronotal disc and always the apical half of the elytron a chestnut reddish-brown, which contrasts with the pitchy margins of the pronotal disc and the elytral area anterior to the mid-point but behind and between the whitish-yellow humeral spot. This whitish-yellow spot is narrowly separated from the pronotal base, and does not extend mesad onto the second interstria. The unique shape of the male apex of the male genitalia (Fig. 10) will further confirm the identity of this species.
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF3CFFB3FF3AF86DFCE54EF9" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8BAA36CAAF3CFFB3FF3AF86DFACA4F39" blockId="1.[151,1436,1645,2034]" lastBlockId="2.[151,1436,152,1149]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="B961EAD8AF3CFFB0FF3AF86DFE5B4AEB" bold="true" box="[197,439,1925,1951]" pageId="1" pageNumber="2">Description (male).</emphasis>
Body oval (
<figureCitation id="132E2A4FAF3CFFB0FDAEF86DFD464AEB" box="[593,682,1925,1951]" captionStart="FIGURES 19 – 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 – 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="1" pageNumber="2">Fig. 19</figureCitation>
); finely, sparsely pilose; castaneous to dark castaneous, pitchy around and between elytral maculae, along suture, and along base of pronotum; whitish-yellow spot on each elytron from 8th stria to slightly mesad of 3rd, narrowly separated from anterior margin of elytron, rounded behind, very slightly indented at intersection with 4th stria (
<figureCitation id="132E2A4FAF3FFFB3FCB1FF72FC7A4DC0" box="[846,918,154,180]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
); epipleuron broadly whitish-yellow laterad of elytral spot for basal 2/3rds of distance to metacoxa; apical half of elytron without whitish-yellow spot. Eyes dorsally separated by
<date id="FFAB100AAF3FFFB3FE66FF05FE514C73" box="[409,445,237,263]" pageId="2" pageNumber="3">7X</date>
maximum width of dorsal lobe (
<figureCitation id="132E2A4FAF3FFFB3FCBCFF05FC614C73" box="[835,909,237,263]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Labium with apex truncate, palpi with palpomere 2 bearded internally with long setae; base of mentum bearing 2 very long setae (
<figureCitation id="132E2A4FAF3FFFB3FB56FEFDFB034C5B" box="[1193,1263,277,303]" captionStart="FIGURES 1 – 9. 1 – 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 – 9. 1 – 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 – 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 – 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Pronotum as wide as base of elytra; lateral margins nearly parallel for 4/5ths length from base, rounded inwardly in apical 1/ 5th; basal margin distinct laterad 2nd elytral striae. Elytral striae 1–6 impunctate, consisting of 2 parallel lines, stria 7 and 8 very faintly subpunctate, 7 very weakly impressed, obsolete anteriorly on elytral spot; interstria flat, conforming to shape of elytron. Foretibia transversely truncate apically, with 2 subacute teeth externally; serrulate between and proximal to teeth, apical spur stout, wide toward apex (Fig. 13). Metatibia weakly serrulate on internal margin (Fig. 16). Adeagus with shape of phallobase apex and parameres unique, as in Fig. 10. Length: 2.3–2.5 mm, measured along midline from anterior margin of pronotum to tip of elytra.
</paragraph>
<paragraph id="8BAA36CAAF3FFFB3FF39FDB3FEAE4FB1" blockId="2.[151,1436,152,1149]" pageId="2" pageNumber="3">Female: differs from male in lacking serrulations on internal margin of metatibia, the tapering apical spur of the protibia, and the more brightly contrasting difference in the castaneous versus pitchy areas on elytron and pronotum.</paragraph>
<paragraph id="8BAA36CAAF3FFFB3FF3AFD3BFCE54EF9" blockId="2.[151,1436,152,1149]" pageId="2" pageNumber="3">
<emphasis id="B961EAD8AF3FFFB3FF3AFD3BFEC74F99" bold="true" box="[197,299,723,749]" pageId="2" pageNumber="3">
<typeStatus id="54AE8868AF3FFFB3FF3AFD3BFECA4F99" box="[197,294,723,749]" pageId="2" pageNumber="3">TYPES</typeStatus>
.
</emphasis>
<typeStatus id="54AE8868AF3FFFB3FEC8FD3BFE3E4F99" box="[311,466,723,749]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
MALE: DOM:REP.: Independencia;
<quantity id="4CED9B2FAF3FFFB3FC62FD3BFC004F99" box="[925,1004,723,749]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="km" value="30.0">30 km</quantity>
NW La Descubierta; Sabana Real,
<quantity id="4CED9B2FAF3FFFB3FF68FD13FF064E61" box="[151,234,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="2" pageNumber="3" unit="m" value="1646.0">1646m</quantity>
, cloud; forest dung tps,
<date id="FFAB100AAF3FFFB3FDFFFD13FDD34E61" box="[512,575,763,789]" pageId="2" pageNumber="3" value="1925-11">25XI</date>
–;
<date id="FFAB100AAF3FFFB3FDA5FD13FD2C4E61" box="[602,704,763,789]" pageId="2" pageNumber="3" value="1991-12-05">5.XII.91</date>
, S and J Peck, 91-332 [CMNC]. ALLOTYPE FEMALE: same data as
<typeStatus id="54AE8868AF3FFFB3FF0FFCCBFEBA4E49" box="[240,342,803,829]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
<typeStatus id="54AE8868AF3FFFB3FEA0FCCBFDEA4E49" box="[351,518,803,829]" pageId="2" pageNumber="3" type="paratype">PARATYPES</typeStatus>
: 1 MALE, 2 FEMALES — same data as
<typeStatus id="54AE8868AF3FFFB3FC0DFCCBFBB44E49" box="[1010,1112,803,829]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
[CMNC, TKPC, WIBF]. 1 FEMALE — DOM:REP.: Independencia;
<quantity id="4CED9B2FAF3FFFB3FD77FCA3FD3E4E11" box="[648,722,843,869]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="km" value="30.0">30 km</quantity>
NW La Descubierta; Sabana Real,
<quantity id="4CED9B2FAF3FFFB3FB8CFCA3FB244E11" box="[1139,1224,843,869]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="2" pageNumber="3" unit="m" value="1646.0">1646m</quantity>
, cloud; forest carrion,
<date id="FFAB100AAF3FFFB3FF2BFC9BFEFB4EF9" box="[212,279,883,909]" pageId="2" pageNumber="3" value="1925-11">25XI</date>
–;
<date id="FFAB100AAF3FFFB3FED3FC9BFE624EF9" box="[300,398,883,909]" pageId="2" pageNumber="3" value="1991-12-05">5.XII.91</date>
, S and J Peck, 91-331 [CMNC].
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF3FFFB3FF3AFC73FB114909" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BAA36CAAF3FFFB3FF3AFC73FDA44EA9" blockId="2.[151,1436,152,1149]" pageId="2" pageNumber="3">
<emphasis id="B961EAD8AF3FFFB3FF3AFC73FEBE4EC1" bold="true" box="[197,338,923,949]" pageId="2" pageNumber="3">Etymology.</emphasis>
Named in honor of one of the collectors, our valued friend and colleague, Jarmila Kukalová- Peck of Carleton University, Ottawa.
</paragraph>
<paragraph id="8BAA36CAAF3FFFB3FF39FC03FB114909" blockId="2.[151,1436,152,1149]" pageId="2" pageNumber="3">
<emphasis id="B961EAD8AF3FFFB3FF39FC03FEF84971" bold="true" box="[198,276,1003,1029]" pageId="2" pageNumber="3">Notes.</emphasis>
This is a pretty little species, with some individuals quite strikingly marked with the pronotal disc and apico-lateral areas of elytron contrastingly lighter that surrounding areas. It is known only from high elevation cloud forests in the south facing slopes of the Sierra de Neiba, an isolated massif between the Cordillera Central and Lago Enriquillo depression. Specimens were taken in dung traps and at carrion.
</paragraph>
</subSubSection>
</treatment>
</document>
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