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<document id="66129917BFB91DB9D5B3FA679EA99DF7" ID-DOI="10.5852/ejt.2020.615" ID-GBIF-Dataset="b2465730-6237-4a14-a98d-7fd2d1096cb0" ID-Zenodo-Dep="3711000" ID-ZooBank="637FC87F-13B5-4B32-BC52-11A9B30ECF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584210958022" checkinUser="plazi" docAuthor="Galea, Horia R." docDate="2020" docId="03B520481A53FFAF957D74F659DF83BC" docLanguage="en" docName="ejt-615_galea.pdf.imf" docOrigin="European Journal of Taxonomy 615" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cladocarpus partitus Galea 2020, sp. nov." docType="treatment" docUuid="498492D3-3B64-485D-9414-2BD05CF55103" docUuidSource="ZooBank" docVersion="6" lastPageNumber="13" masterDocId="FF8C58301A5AFFA3971777095A2A8450" masterDocTitle="Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program" masterLastPageNumber="47" masterPageNumber="1" pageNumber="10" updateTime="1698728323888" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="ABC3DB798A446BD7E83F181213E0D504" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CDAE83259E2A3059727961DB54F38D6B">
<mods:title id="FF283C84C3A5DDEC228B4D26B9E2C7E9">Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program</mods:title>
</mods:titleInfo>
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<mods:role id="51345215C41FBAFAE93337FF74B1B996">
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<mods:namePart id="30C47F48A2BEF2219D5D567A1DDC1334">Galea, Horia R.</mods:namePart>
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<mods:typeOfResource id="32EECCC7F67967405A470A936E87B291">text</mods:typeOfResource>
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<mods:title id="02750E7FC4BB4CF81E53DA4CF2423F93">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="98B7D44B4D1D39848C5E7F1953D7F310">
<mods:date id="859B5C6C6B45E98E41E23612F58D6B6C">2020</mods:date>
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<mods:number id="B38DBD820B9694F8F0BC2752D1B75C6E">2020-03-13</mods:number>
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<mods:detail id="ED465C337D4F4FF2E74160877C2E1282" type="volume">
<mods:number id="CA21A7B04BB473D8DB4E27842E6F7599">615</mods:number>
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<mods:classification id="A7DF2DCB40E686E6D16D1B750BFCBB02">journal article</mods:classification>
<mods:identifier id="E25EBEA6A1F4C096C9C657A2B9699F47" type="DOI">10.5852/ejt.2020.615</mods:identifier>
<mods:identifier id="0A96965E25183CDDFF7C88C99180F4EB" type="GBIF-Dataset">b2465730-6237-4a14-a98d-7fd2d1096cb0</mods:identifier>
<mods:identifier id="AF3511608B1FE6712E4536CAF8B69820" type="Zenodo-Dep">3711000</mods:identifier>
<mods:identifier id="3FCD99F5F367CFF7468F2899169C0B02" type="ZooBank">637FC87F-13B5-4B32-BC52-11A9B30ECF1D</mods:identifier>
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<treatment id="03B520481A53FFAF957D74F659DF83BC" ID-DOI="http://doi.org/10.5281/zenodo.3718315" ID-GBIF-Taxon="162505292" ID-Zenodo-Dep="3718315" LSID="urn:lsid:zoobank.org:act:498492D3-3B64-485D-9414-2BD05CF55103" httpUri="http://treatment.plazi.org/id/03B520481A53FFAF957D74F659DF83BC" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="C306C2D51A53FFAA957D74F65E97806D" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BA3915E1A53FFAA957D74F659E0804A" blockId="9.[375,1213,1023,1120]" box="[618,970,1023,1050]" pageId="9" pageNumber="10">
<heading id="D0EB26321A53FFAA957D74F659E0804A" box="[618,970,1023,1050]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="4C1CEADD1A53FFAA957D74F659428049" ID-CoL="847LS" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[618,872,1023,1050]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="9" pageNumber="10" phylum="Cnidaria" rank="species" species="partitus" status="sp. nov.">
<emphasis id="B9684D4C1A53FFAA957D74F659428049" bold="true" box="[618,872,1023,1050]" italics="true" pageId="9" pageNumber="10">Cladocarpus partitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A53FFAA9478730959E0804A" box="[879,970,1024,1050]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA3915E1A53FFAA9660732B5E97806D" blockId="9.[375,1213,1023,1120]" box="[375,1213,1058,1085]" pageId="9" pageNumber="10">
<uri id="FF8D9D5C1A53FFAA9660732B5E97806D" box="[375,1213,1058,1085]" pageId="9" pageNumber="10">
urn:lsid:zoobank.org:act:
<uuid id="FFBAAB8B1A53FFAA9582732B5E97806D" box="[661,1213,1058,1085]" pageId="9" pageNumber="10">498492D3-3B64-485D-9414-2BD05CF55103</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A53FFAA95DC734F59438030" box="[715,873,1093,1120]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BA3915E1A53FFAA95DC734F59438030" blockId="9.[375,1213,1023,1120]" box="[715,873,1093,1120]" pageId="9" pageNumber="10">
<figureCitation id="13278DDB1A53FFAA95DC734F5939800F" box="[715,787,1093,1120]" captionStart="Fig" captionStartId="10.[189,232,1860,1886]" captionTargetBox="[189,1399,659,1818]" captionTargetId="figure@10.[189,1399,659,1819]" captionTargetPageId="10" captionText="Fig. 6. A–B. Cladocarpus partitus sp. nov., fertile cormoid (A) and phylactocarp (B) from sample MNHN-IK-2015-529 (holotype). — C–E. Cladocarpus pennatus sp. nov., fertile colony (C) and phylactocarp with male gonothecae (D) from sample MNHN-IK-2015-534 (holotype); phylactocarp with female gonothecae from sample MNHN-IK-2015-537 (one paratype). Scale bars: A, C–E = 1 cm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3711010" httpUri="https://zenodo.org/record/3711010/files/figure.png" pageId="9" pageNumber="10">Figs 6</figureCitation>
A–B, 7
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A53FFAA97AA738E5833825C" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BA3915E1A53FFAA97AA738E5B1C80F1" blockId="9.[189,310,1159,1185]" box="[189,310,1159,1185]" pageId="9" pageNumber="10">
<heading id="D0EB26321A53FFAA97AA738E5B1C80F1" bold="true" box="[189,310,1159,1185]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9684D4C1A53FFAA97AA738E5B1C80F1" bold="true" box="[189,310,1159,1185]" pageId="9" pageNumber="10">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A53FFAA97AA73BF5833825C" blockId="9.[189,1399,1206,1549]" pageId="9" pageNumber="10">
<taxonomicName id="4C1CEADD1A53FFAA97AA73BF5B7F8080" authorityName="Allman" authorityYear="1874" box="[189,341,1206,1232]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="9" pageNumber="10" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A53FFAA97AA73BF5B7F8080" box="[189,341,1206,1232]" italics="true" pageId="9" pageNumber="10">Cladocarpus</emphasis>
</taxonomicName>
with lightly fascicled, unbranched stems, whose rather long internodes bear a frontal row of 2–3 nematothecae, a lateral apophysis with a basal mamelon, as well as an anterior axillar nematotheca. Cladia relatively spaced from one another. Cormidia elongated, with up to 8 internal septa, 2 abaxial up to 6 adaxial. Hydrotheca deep, mesial nematotheca short, rim with median abaxial cusp, nearly smooth elsewhere; an internal ridge projecting forwards and slightly upwards from the lower fourth of the adaxial wall of hydrotheca, as well as an arched, two-winged septum above, leaving a slit-like, median passage for the hydranth. Phylactocarp borne on stem internode by means of a short apophysis; segmented into moderately-long internodes, each with one frontal, triangular nematotheca; gonothecae on distalmost internodes; tubular, with a main, lateral, subterminal, slit-like aperture, and an inconspicuous, secondary gutter-shaped aperture above.
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A53FFAA97AA713D58B782F0" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BA3915E1A53FFAA97AA713D5B6E821E" blockId="9.[189,1399,1588,1696]" box="[189,324,1588,1614]" pageId="9" pageNumber="10">
<heading id="D0EB26321A53FFAA97AA713D5B6E821E" bold="true" box="[189,324,1588,1614]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9684D4C1A53FFAA97AA713D5B6E821E" bold="true" box="[189,324,1588,1614]" pageId="9" pageNumber="10">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A53FFAA97AA716A58B782F0" blockId="9.[189,1399,1588,1696]" pageId="9" pageNumber="10">
From the Latin ‘
<emphasis id="B9684D4C1A53FFAA966C716A5BE4822D" box="[379,462,1635,1661]" italics="true" pageId="9" pageNumber="10">partĭor</emphasis>
,
<emphasis id="B9684D4C1A53FFAA96CB716A586C822C" box="[476,582,1635,1660]" italics="true" pageId="9" pageNumber="10">-ītus sum</emphasis>
,
<emphasis id="B9684D4C1A53FFAA9543716A5853822C" box="[596,633,1635,1660]" italics="true" pageId="9" pageNumber="10">-īri</emphasis>
’, meaning ‘partitioned’, with reference to its hydrothecae that are divided by a transverse, bilobate septum.
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A53FFAA97AA71CE5ECF8307" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BA3915E1A53FFAA97AA71CE5B8082B1" blockId="9.[189,426,1735,1761]" box="[189,426,1735,1761]" pageId="9" pageNumber="10">
<heading id="D0EB26321A53FFAA97AA71CE5B8082B1" bold="true" box="[189,426,1735,1761]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9684D4C1A53FFAA97AA71CE5B8082B1" bold="true" box="[189,426,1735,1761]" pageId="9" pageNumber="10">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A53FFAA97FD71FF5B768340" blockId="9.[189,1398,1782,1879]" box="[234,348,1782,1808]" pageId="9" pageNumber="10">
<emphasis id="B9684D4C1A53FFAA97FD71FF5B768340" bold="true" box="[234,348,1782,1808]" pageId="9" pageNumber="10">
<typeStatus id="54A72FFC1A53FFAA97FD71FF5B768340" box="[234,348,1782,1808]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA3915E1A53FFAA97AA70105ECF8307" blockId="9.[189,1398,1782,1879]" pageId="9" pageNumber="10">
<materialsCitation id="3B749B031A53FFAA97AA70105ECF8307" ID-GBIF-Occurrence="2575926616" collectingDate="2016-08-28" collectionCode="MNHN" collectorName="KANACONO" country="New Caledonia" elevation="463" latitude="-22.966667" location="stn DW 4770" longLatPrecision="1258" longitude="168.35" pageId="9" pageNumber="10" specimenCode="MNHN-IK-2015-529" specimenCount="1" stateProvince="PACIFIC OCEAN" typeStatus="holotype">
<collectingRegion id="49D85FBC1A53FFAA97AA70105BB38363" box="[189,409,1817,1843]" pageId="9" pageNumber="10">PACIFIC OCEAN</collectingRegion>
•
<specimenCount id="9D1A5AD71A53FFAA96A1701058358363" box="[438,543,1817,1843]" pageId="9" pageNumber="10">1 colony</specimenCount>
, ca
<quantity id="4CE43CBB1A53FFAA95587010588B8363" box="[591,673,1817,1844]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="cm" value="6.5">6.5 cm</quantity>
high, hydrorhiza missing; off
<collectingCountry id="F30BD1CE1A53FFAA931870105EE08363" box="[1039,1226,1817,1843]" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
,
<location id="8EC3C7851A53FFAA93C070105F588363" LSID="urn:lsid:plazi:treatment:03B520481A53FFAF957D74F659DF83BC:8EC3C7851A53FFAA93C070105F588363" box="[1239,1394,1817,1843]" country="New Caledonia" latitude="-22.966667" longLatPrecision="1258" longitude="168.35" name="stn DW 4770" pageId="9" pageNumber="10" stateProvince="PACIFIC OCEAN">stn DW4770</location>
;
<geoCoordinate id="EE28F7991A53FFAA97AA70335B098306" box="[189,291,1850,1878]" degrees="22" direction="south" minutes="58" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="-22.966667">22°58ʹ S</geoCoordinate>
,
<geoCoordinate id="EE28F7991A53FFAA962770335B828306" box="[304,424,1850,1878]" degrees="168" direction="east" minutes="21" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="168.35">168°21ʹ E</geoCoordinate>
;
<quantity id="4CE43CBB1A53FFAA96A3703558168306" box="[436,572,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="4.7" metricValueMin="4.55" pageId="9" pageNumber="10" unit="m" value="462.5" valueMax="470.0" valueMin="455.0">
<elevation id="0031766D1A53FFAA96A3703558168306" box="[436,572,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="4.7" metricValueMin="4.55" pageId="9" pageNumber="10" unit="m" value="462.5" valueMax="470.0" valueMin="455.0">455–470 m</elevation>
</quantity>
;
<date id="FFA2B79E1A53FFAA955E703558C18306" box="[585,747,1852,1878]" pageId="9" pageNumber="10" value="2016-08-28">
<collectingDate id="EFE64E761A53FFAA955E703558C18306" box="[585,747,1852,1878]" pageId="9" pageNumber="10" value="2016-08-28">28 Aug. 2016</collectingDate>
</date>
;
<collectorName id="26E9F4881A53FFAA95E0703559B58306" box="[759,927,1852,1878]" pageId="9" pageNumber="10">KANACONO</collectorName>
leg.;
<specimenCode id="DBBA39251A53FFAA94F670355ECB8307" box="[993,1249,1852,1879]" pageId="9" pageNumber="10">MNHN-IK-2015-529</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A53FFAF97AA7074582880A1" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="8BA3915E1A53FFAA97AA70745B6483C7" blockId="9.[189,1399,1917,2026]" box="[189,334,1917,1943]" pageId="9" pageNumber="10">
<heading id="D0EB26321A53FFAA97AA70745B6483C7" bold="true" box="[189,334,1917,1943]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9684D4C1A53FFAA97AA70745B6483C7" bold="true" box="[189,334,1917,1943]" pageId="9" pageNumber="10">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A53FFAF97AA70A5582880A1" blockId="9.[189,1399,1917,2026]" lastBlockId="12.[189,1399,267,1265]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
Colony erect, feather shaped, delicate, ca
<quantity id="4CE43CBB1A53FFAA95D470A559328396" box="[707,792,1964,1991]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="cm" value="6.5">6.5 cm</quantity>
high, hydrorhiza not observed. Stem unbranched although, in the present specimen, the distal part had been partly broken off, but still remained attached, and was consequently displaced laterally by a new, regenerative colony tip that has grown up vertically, giving the impression that the old apex is merely a lateral branch; stem fascicled proximally, grading to monosiphonic distally; every additional auxiliary tube adhering to the dorsal side of the preceding one, not forming a beam around the main tube, but resulting in a laterally-flattened structure (
<figureCitation id="13278DDB1A50FFA9921C767C5F4D85DF" box="[1291,1383,373,399]" captionStart="Fig" captionStartId="11.[189,232,1827,1853]" captionTargetBox="[202,1398,265,1768]" captionTargetId="figure@11.[189,1398,265,1777]" captionTargetPageId="11" captionText="Fig. 7. Cladocarpus partitus sp. nov., from sample MNHN-IK-2015-529. Portions from proximal (A, seen frontally; B, seen laterally) and distal (C, with insertion of phylactocarp) parts of stem. Hydrotheca seen laterally (D) and frontally (E), and details of its basal (F) and distal (G) parts; mesial nematotheca (H) seen frontally (left) and apically (right); internal septa in an apical view of the hydrotheca (I). Distal end of phylactocarp with two gonothecae (J). Scale bars: A–C = 500 µm; D–G, J = 200 µm; H, I = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3711012" httpUri="https://zenodo.org/record/3711012/files/figure.png" pageId="10" pageNumber="11">Fig. 7B</figureCitation>
); division into internodes indistinct, except for 3–4 oblique, deeply-cut nodes towards the proximal part of hydrocaulus, passing through all cauline tubes at a time. Equivalents of internodes
<quantity id="4CE43CBB1A50FFA9939976B25F198586" box="[1166,1331,443,470]" metricMagnitude="-1" metricUnit="m" metricValue="8.75" metricValueMax="10.5" metricValueMin="7.0" pageId="10" pageNumber="11" unit="mm" value="875.0" valueMax="1050.0" valueMin="700.0">700–1050 µm</quantity>
long,
<quantity id="4CE43CBB1A50FFA997AA76D75B7F85A9" box="[189,341,478,505]" metricMagnitude="-1" metricUnit="m" metricValue="2.125" metricValueMax="2.25" metricValueMin="2.0" pageId="10" pageNumber="11" unit="mm" value="212.5" valueMax="225.0" valueMin="200.0">200–225 µm</quantity>
wide, composed of 2–3 frontal nematothecae in a row, a short lateral cladial apophysis, a reduced nematotheca (with small, rounded aperture) on an inconspicuous mamelon at its base, as well as an axillar, anterior nematotheca rather lateral to it; there is no dorsal axillar nematotheca. Stem nematothecae large, inverted-triangular, with thick perisarc, apertures distal, wide, with inwardly- rolled abaxial wall, giving the impression that two circular apertures are present laterally; apophyses slightly shifted onto the anterior side of the colony, the two rows of hydrocladia forming an obtuse angle. Cladia relatively spaced from one another, up to
<quantity id="4CE43CBB1A56FFAF9440765A59B9853E" box="[855,915,339,366]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="cm" value="1.0">1 cm</quantity>
long, unbranched, divided into up to 12 cormidia by means of transverse, though not clearly demarcated nodes (
<quantity id="4CE43CBB1A56FFAF94EB767E5E5C85C2" box="[1020,1142,375,402]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="12" pageNumber="13" unit="mm" value="85.0" valueMax="95.0" valueMin="75.0">75–95 µm</quantity>
wide), each internode
<quantity id="4CE43CBB1A56FFAF97AA76925B7F85E6" box="[189,341,411,438]" metricMagnitude="-1" metricUnit="m" metricValue="8.2" metricValueMax="8.65" metricValueMin="7.75" pageId="12" pageNumber="13" unit="mm" value="820.0" valueMax="865.0" valueMin="775.0">775–865 µm</quantity>
long, accommodating a hydrotheca and its 3 associated nematothecae: one mesial and a pair of laterals; up to 8 intranodal projections of the perisarc, of which 2 are given off from the abaxial side below the hydrothecal base, and up to 6 arise from the dorsal wall of the hydrotheca and project for a short distance into the lumen of the internode. Hydrotheca elongated,
<quantity id="4CE43CBB1A56FFAF933D750E5EEA8672" box="[1066,1216,519,546]" metricMagnitude="-1" metricUnit="m" metricValue="6.175" metricValueMax="6.35" metricValueMin="6.0" pageId="12" pageNumber="13" unit="mm" value="617.5" valueMax="635.0" valueMin="600.0">600–635 µm</quantity>
deep, laterallyflattened, ad- and abaxial walls with slight basal bulges; a septum projecting slightly upwards from the lower portion of the adaxial wall reaches up to the middle of the thecal lumen, and is provided anteriorly with a distal swelling scattered with granular projections; a second, lamellar, two-winged septum is given off upwardly from the lateral sides of the hydrotheca at a rather acute angle, then changes its orientation and slightly plunges downwards, attaching its anterior border to the abaxial wall of the theca, leaving a slit-like, median passage between its two lateral ‘wings’; hydrothecal aperture
<quantity id="4CE43CBB1A56FFAF922B75D65B30874E" metricMagnitude="-1" metricUnit="m" metricValue="2.275" metricValueMax="2.3" metricValueMin="2.25" pageId="12" pageNumber="13" unit="mm" value="227.5" valueMax="230.0" valueMin="225.0">225– 230 µm</quantity>
wide, rim with a prominent, central, well-developed, abaxial cusp, nearly smooth elsewhere. Mesial nematothecae elongated, rather broad in frontal view, adnate for most of their length, leaving a short (
<quantity id="4CE43CBB1A56FFAF961F74425BAF8736" box="[264,389,843,870]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="12" pageNumber="13" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 µm</quantity>
) portion free from the base of the corresponding hydrotheca; aperture broad, abaxial wall inwardly-rolled, rim crenelate at corners; mesial nematotheca of first cormidium short, guttershaped and not coalescent with the hydrothecal base. Lateral nematothecae
<quantity id="4CE43CBB1A56FFAF932C749A5EF887FE" box="[1083,1234,915,942]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="12" pageNumber="13" unit="mm" value="150.0" valueMax="155.0" valueMin="145.0">145–155 µm</quantity>
long, broadly cylindrical, reaching the level of the hydrothecal rim; adaxial wall deeply scooped out, abaxial wall crenelate to wavy. Phylactocarp mounted on a short stem apophysis arising laterally, next to a cladial apophysis, on opposite side; divided into several moderately-long (
<quantity id="4CE43CBB1A56FFAF94CF74F65E58804A" box="[984,1138,1023,1050]" metricMagnitude="-1" metricUnit="m" metricValue="4.225" metricValueMax="4.85" metricValueMin="3.6" pageId="12" pageNumber="13" unit="mm" value="422.5" valueMax="485.0" valueMin="360.0">360–485 µm</quantity>
) internodes by means of transverse to slightly oblique nodes, each bearing a frontal nematotheca similar to those of stem, with large apical apertures and inwardly-rolled abaxial walls, giving the impression that the nematothecae are bifid; last couple of internodes additionally provided with a basally concrescent gonotheca; the latter long, tubular (ca 1630 ×
<quantity id="4CE43CBB1A56FFAF96F97386586580FA" box="[494,591,1167,1194]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="mm" value="400.0">400 µm</quantity>
), tapering below, distally with a main, broad, subterminal, slit-like aperture with flimsy, wavy rim, above which a second, short, slender, gutter-shaped aperture is to be found. Sex undeterminable.
</paragraph>
</subSubSection>
<caption id="DF63C1D61A50FFA997AA704D5B1183BC" ID-DOI="http://doi.org/10.5281/zenodo.3711010" ID-Zenodo-Dep="3711010" httpUri="https://zenodo.org/record/3711010/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1860,1886]" targetBox="[189,1399,659,1818]" targetPageId="10">
<paragraph id="8BA3915E1A50FFA997AA704D5B1183BC" blockId="10.[189,1399,1860,2028]" pageId="10" pageNumber="11">
<emphasis id="B9684D4C1A50FFA997AA704D5B7B830E" bold="true" box="[189,337,1860,1886]" pageId="10" pageNumber="11">Fig. 6. A–B</emphasis>
.
<taxonomicName id="4C1CEADD1A50FFA99672704D5849830E" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[357,611,1860,1886]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="partitus" status="sp. nov.">
<emphasis id="B9684D4C1A50FFA99672704D5849830E" box="[357,611,1860,1886]" italics="true" pageId="10" pageNumber="11">Cladocarpus partitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A50FFA99578704C58FA830F" box="[623,720,1861,1887]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, fertile cormoid (A) and phylactocarp (B) from sample MNHN-IK-2015-529 (holotype).
<emphasis id="B9684D4C1A50FFA9954F7061589083D1" bold="true" box="[600,698,1895,1922]" pageId="10" pageNumber="11">— C–E</emphasis>
.
<taxonomicName id="4C1CEADD1A50FFA995C7706159CA83D1" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[720,992,1896,1922]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="pennatus" status="sp. nov.">
<emphasis id="B9684D4C1A50FFA995C7706159CA83D1" box="[720,992,1896,1922]" italics="true" pageId="10" pageNumber="11">Cladocarpus pennatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A50FFA994F970615E7883D2" box="[1006,1106,1896,1922]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, fertile colony (C) and phylactocarp with male gonothecae (D) from sample MNHN-IK-2015-534 (holotype); phylactocarp with female gonothecae from sample MNHN-IK-2015-537 (one paratype). Scale bars: A, C–E = 1 cm; B = 1 mm.
</paragraph>
</caption>
<caption id="DF63C1D61A51FFA897AA702A5B3583BE" ID-DOI="http://doi.org/10.5281/zenodo.3711012" ID-Zenodo-Dep="3711012" httpUri="https://zenodo.org/record/3711012/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1827,1853]" targetBox="[202,1398,265,1768]" targetPageId="11">
<paragraph id="8BA3915E1A51FFA897AA702A5B3583BE" blockId="11.[189,1399,1827,2030]" pageId="11" pageNumber="12">
<emphasis id="B9684D4C1A51FFA897AA702A5B26836E" bold="true" box="[189,268,1827,1854]" pageId="11" pageNumber="12">Fig. 7.</emphasis>
<taxonomicName id="4C1CEADD1A51FFA89602702D583A836D" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[277,528,1828,1854]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="11" pageNumber="12" phylum="Cnidaria" rank="species" species="partitus" status="sp. nov.">
<emphasis id="B9684D4C1A51FFA89602702D583A836D" box="[277,528,1828,1854]" italics="true" pageId="11" pageNumber="12">Cladocarpus partitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A51FFA8950D702D5852836E" box="[538,632,1828,1854]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, from sample MNHN-IK-2015-529. Portions from proximal (A, seen frontally; B, seen laterally) and distal (C, with insertion of phylactocarp) parts of stem. Hydrotheca seen laterally (D) and frontally (E), and details of its basal (F) and distal (G) parts; mesial nematotheca (H) seen frontally (left) and apically (right); internal septa in an apical view of the hydrotheca (I). Distal end of phylactocarp with two gonothecae (J). Scale bars: A–C = 500 µm; D–G, J = 200 µm; H, I = 100 µm.
</paragraph>
</caption>
<subSubSection id="C306C2D51A56FFAF97AA72135F2F8329" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BA3915E1A56FFAF97AA72135B058164" blockId="12.[189,303,1306,1332]" box="[189,303,1306,1332]" pageId="12" pageNumber="13">
<heading id="D0EB26321A56FFAF97AA72135B058164" bold="true" box="[189,303,1306,1332]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B9684D4C1A56FFAF97AA72135B058164" bold="true" box="[189,303,1306,1332]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A56FFAF97AA72405F5D829C" blockId="12.[189,1399,1353,1740]" pageId="12" pageNumber="13">
Although not comprehensively described and/or illustrated, especially upon an apical examination of the hydrothecae, the internal septa of
<taxonomicName id="4C1CEADD1A56FFAF955172645E3281D7" authority="(G. O. Sars, 1874)" baseAuthorityName="G. O. Sars" baseAuthorityYear="1874" box="[582,1048,1389,1415]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="bicuspis">
<emphasis id="B9684D4C1A56FFAF95517264596E81D7" box="[582,836,1389,1415]" italics="true" pageId="12" pageNumber="13">Cladocarpus bicuspis</emphasis>
(G.O.
<bibRefCitation id="EF8DECAF1A56FFAF948372645E3A81D7" author="Sars G. O." box="[916,1040,1389,1415]" firstAuthor="Sars" journalOrPublisher="Forhandlinger i Videnskabs-Selskabet i Kristiania" pageId="12" pageNumber="13" pagination="91 - 150" part="1873" refId="ref23605" refString="Sars G. O. 1874. Bidrag til Kundskaben om Norges Hydroider. Forhandlinger i Videnskabs-Selskabet i Kristiania 1873: 91 - 150." title="Bidrag til Kundskaben om Norges Hydroider" type="journal article" year="1874">Sars, 1874</bibRefCitation>
)
</taxonomicName>
possibly share morphological affinities with those of the present species. Indeed, a transverse septum, originating from the lateroabaxial hydrothecal walls, projects nearly horizontally midway into the lumen, then plunges down so as to meet the distal, swollen border of the adaxial hydrothecal ridge (G.O.
<bibRefCitation id="EF8DECAF1A56FFAF930072D05EBA81A3" author="Sars G. O." box="[1047,1168,1497,1523]" firstAuthor="Sars" journalOrPublisher="Forhandlinger i Videnskabs-Selskabet i Kristiania" pageId="12" pageNumber="13" pagination="91 - 150" part="1873" refId="ref23605" refString="Sars G. O. 1874. Bidrag til Kundskaben om Norges Hydroider. Forhandlinger i Videnskabs-Selskabet i Kristiania 1873: 91 - 150." title="Bidrag til Kundskaben om Norges Hydroider" type="journal article" year="1874">Sars 1874</bibRefCitation>
: pl. 2 fig. 9;
<bibRefCitation id="EF8DECAF1A56FFAF923272D05AD68248" author="Kramp P. L." firstAuthor="Kramp" journalOrPublisher="Danmarks Fauna" pageId="12" pageNumber="13" pagination="1 - 207" part="41" refId="ref22799" refString="Kramp P. L. 1935. Polypdyr (Coelenterata). I. Ferskvardspolypper og Goplepolypper. Danmarks Fauna 41: 1 - 207." title="Polypdyr (Coelenterata). I. Ferskvardspolypper og Goplepolypper" type="journal article" year="1935">Kramp 1935</bibRefCitation>
: fig. 71b;
<bibRefCitation id="EF8DECAF1A56FFAF966F72F458198248" author="Cornelius P. F. S." box="[376,563,1533,1560]" firstAuthor="Cornelius" journalOrPublisher="Synopses of the British Fauna" pageId="12" pageNumber="13" pagination="1 - 386" part="50" refId="ref22008" refString="Cornelius P. F. S. 1995. North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae. Synopses of the British Fauna 50: 1 - 386." title="North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae" type="journal article" year="1995">Cornelius 1995</bibRefCitation>
: fig. 47c). Frontal views of the hydrotheca illustrated in the literature (
<bibRefCitation id="EF8DECAF1A56FFAF97D271285B7D826B" author="Broch H." box="[197,343,1569,1595]" firstAuthor="Broch" journalOrPublisher="Danish Ingolf-Expedition" pageId="12" pageNumber="13" pagination="1 - 206" part="5" refId="ref21849" refString="Broch H. 1918. Hydroida (Part II). Danish Ingolf-Expedition 5 (7): 1 - 206." title="Hydroida (Part II)" type="journal article" year="1918">Broch 1918</bibRefCitation>
: fig. 48c;
<bibRefCitation id="EF8DECAF1A56FFAF96D8712858A7826B" author="Schuchert P." box="[463,653,1569,1595]" firstAuthor="Schuchert" journalOrPublisher="Meddelelser om GrOnland, Bioscience" pageId="12" pageNumber="13" pagination="1 - 184" part="53" refId="ref23630" refString="Schuchert P. 2001. Hydroids of Greenland and Iceland. Meddelelser om GrOnland, Bioscience 53: 1 - 184." title="Hydroids of Greenland and Iceland" type="journal article" year="2001">Schuchert 2001</bibRefCitation>
: fig. 114b) do not provide sufficient information on the global aspect of the abaxial projection. Besides this possible common feature, the hydrothecae of
<taxonomicName id="4C1CEADD1A56FFAF93DA714C5F67820F" baseAuthorityName="G. O. Sars" baseAuthorityYear="1874" box="[1229,1357,1605,1631]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="bicuspis">
<emphasis id="B9684D4C1A56FFAF93DA714C5F67820F" box="[1229,1357,1605,1631]" italics="true" pageId="12" pageNumber="13">C. bicuspis</emphasis>
</taxonomicName>
are distinctive through the presence of a median, abaxial pair of marginal cusps, separated from a squared to V-shaped gap prolonged downwards, till the level of abaxial ridge, as a perisarcal thickening; additional features related to its gonosome allow a rapid separation from the present new species (
<bibRefCitation id="EF8DECAF1A56FFAF93A771B85F40829C" author="Cornelius P. F. S." box="[1200,1386,1713,1740]" firstAuthor="Cornelius" journalOrPublisher="Synopses of the British Fauna" pageId="12" pageNumber="13" pagination="1 - 386" part="50" refId="ref22008" refString="Cornelius P. F. S. 1995. North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae. Synopses of the British Fauna 50: 1 - 386." title="North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae" type="journal article" year="1995">Cornelius 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BA3915E1A56FFAF97AA71FA5F2F8329" blockId="12.[189,1398,1779,1914]" pageId="12" pageNumber="13">
Also similar seems to be the abaxial septum of
<taxonomicName id="4C1CEADD1A56FFAF95F671FA5F78835D" authority="Vervoort &amp; Watson, 2003" authorityName="Vervoort &amp; Watson" authorityYear="2003" box="[737,1362,1779,1806]" class="Hydrozoa" family="Aglaopheniidae" genus="Wanglaophenia" kingdom="Animalia" order="Leptothecata" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="longicarpa">
<emphasis id="B9684D4C1A56FFAF95F671FA5E0A835D" box="[737,1056,1779,1805]" italics="true" pageId="12" pageNumber="13">Wanglaophenia longicarpa</emphasis>
<bibRefCitation id="EF8DECAF1A56FFAF933171FD5F78835D" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[1062,1362,1779,1806]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="12" pageNumber="13" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson, 2003</bibRefCitation>
</taxonomicName>
, as observed from the illustration in
<bibRefCitation id="EF8DECAF1A56FFAF95577011595F8361" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[576,885,1815,1842]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="12" pageNumber="13" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson (2003</bibRefCitation>
: fig.
<quantity id="4CE43CBB1A56FFAF94B8701E59F48361" box="[943,990,1815,1841]" metricMagnitude="-2" metricUnit="kg" metricValue="8.1" pageId="12" pageNumber="13" unit="g" value="81.0">81g</quantity>
). This species, however, possesses hydrothecae with a distinct, abaxial, thorn-shaped crest prolonged downwards as a longitudinal ridge, and its phylactocarp has forked costae provided with hydrothecae (
<bibRefCitation id="EF8DECAF1A56FFAF94DA70565ED38329" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[973,1273,1887,1914]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="12" pageNumber="13" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A56FFAF97AA70A859DF83BC" box="[189,1013,1953,2028]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BA3915E1A56FFAF97AA70A85B7C83EB" blockId="12.[189,1013,1953,2028]" box="[189,342,1953,1979]" pageId="12" pageNumber="13">
<heading id="D0EB26321A56FFAF97AA70A85B7C83EB" bold="true" box="[189,342,1953,1979]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B9684D4C1A56FFAF97AA70A85B7C83EB" bold="true" box="[189,342,1953,1979]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A56FFAF97AA70D859DF83BC" blockId="12.[189,1013,1953,2028]" box="[189,1013,2001,2028]" pageId="12" pageNumber="13">
Known only from its
<typeStatus id="54A72FFC1A56FFAF96AD70D85BC783BB" box="[442,493,2001,2027]" pageId="12" pageNumber="13">type</typeStatus>
locality, off
<collectingCountry id="F30BD1CE1A56FFAF959670D8591D83BB" box="[641,823,2001,2027]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>
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