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<document id="72D04C1549CE551E3559C1AADB9C388D" ID-ISSN="1175­5334" ID-ZooBank="505937B0-9F57-4068-82E6-8553826DD5AA" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629822157238" checkinUser="felipe" docAuthor="Judson, Sarah W. &amp; Nelson, C. Riley" docDate="2012" docId="03B387E715488119FF5AF942FDFC52B2" docLanguage="en" docName="zt03541p118.pdf" docOrigin="Zootaxa 3541" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Isoperla kozlovi Zhiltzova 1972" docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FF8AFF9F15668136FFCDFFA7FFAF5015" masterDocTitle="3541" masterLastPageNumber="118" masterPageNumber="1" pageNumber="47" updateTime="1699117090469" updateUser="plazi" zenodo-license-document="CLOSED">
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<subSubSection id="C300657A15488118FF5AF942FE935751" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA536F115488118FF5AF942FDB256EA" blockId="46.[151,541,1765,1791]" box="[151,541,1765,1791]" pageId="46" pageNumber="47">
<heading id="D0ED819D15488118FF5AF942FDB256EA" bold="true" box="[151,541,1765,1791]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<taxonomicName id="4C1A4D7215488118FF5AF942FDB256EA" ID-CoL="6MZY7" authority="Zhiltzova, 1972" authorityName="Zhiltzova" authorityYear="1972" box="[151,541,1765,1791]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="B96EEAE315488118FF5AF942FDB256EA" bold="true" box="[151,541,1765,1791]" pageId="46" pageNumber="47">
<emphasis id="B96EEAE315488118FF5AF942FEF956EA" bold="true" box="[151,342,1765,1791]" italics="true" pageId="46" pageNumber="47">Isoperla kozlovi</emphasis>
Zhiltzova, 1972
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA536F115488118FF5AF88CFE935751" blockId="46.[151,480,1835,1894]" box="[151,316,1835,1860]" pageId="46" pageNumber="47">
<heading id="D0ED819D15488118FF5AF88CFE935751" bold="true" box="[151,316,1835,1860]" fontSize="10" level="2" pageId="46" pageNumber="47" reason="2">
<emphasis id="B96EEAE315488118FF5AF88CFE935751" bold="true" box="[151,316,1835,1860]" pageId="46" pageNumber="47">SYNONYMY</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300657A15488118FF5AF8F7FE4F5773" box="[151,480,1872,1894]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA536F115488118FF5AF8F7FE4F5773" blockId="46.[151,480,1835,1894]" box="[151,480,1872,1894]" pageId="46" pageNumber="47">
<taxonomicName id="4C1A4D7215488118FF5AF8F7FE4F5773" authority="Zhiltzova, 1972" authorityName="Zhiltzova" authorityYear="1972" box="[151,480,1872,1894]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="B96EEAE315488118FF5AF8F7FE955773" box="[151,314,1872,1894]" italics="true" pageId="46" pageNumber="47">Isoperla kozlovi</emphasis>
Zhiltzova, 1972
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C300657A15488118FF5AF834FDCC57BF" box="[151,611,1938,1963]" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BA536F115488118FF5AF834FDCC57BF" blockId="46.[151,1436,1938,2034]" box="[151,611,1938,1963]" pageId="46" pageNumber="47">
<materialsCitation id="3B723CAC15488118FF5AF834FDF057BF" box="[151,607,1938,1963]" country="Mongolia" location="Khentii" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Hentiy" typeStatus="holotype">
<emphasis id="B96EEAE315488118FF5AF834FE2B57BE" bold="true" box="[151,388,1938,1963]" pageId="46" pageNumber="47">
<typeStatus id="54A1885315488118FF5AF834FF4D57BE" box="[151,226,1939,1963]" pageId="46" pageNumber="47">TYPE</typeStatus>
LOCALITY:
</emphasis>
<collectingCountry id="F30D766115488118FE41F835FE5157BF" box="[396,510,1938,1962]" name="Mongolia" pageId="46" pageNumber="47">Mongolia</collectingCountry>
:
<collectingRegion id="49DEF81315488118FDCAF835FDF057BF" box="[519,607,1938,1962]" country="Mongolia" name="Hentiy" pageId="46" pageNumber="47">Khentii</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15488119FF0AF811FD3451DA" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BA536F115488119FF0AF811FD3451DA" blockId="46.[151,1436,1938,2034]" lastBlockId="47.[151,1436,151,680]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B96EEAE315488118FF0AF811FECB57DA" bold="true" box="[199,356,1974,1999]" pageId="46" pageNumber="47">DIAGNOSIS</emphasis>
: Head of adult is infuscate on the anterior half, with a pale oval spot in the ocellar region bordered by a black, U-shaped line which connects all the ocelli (
<figureCitation id="13212A7415488118FC43F87DFC5C57E7" box="[910,1011,2010,2034]" captionStart="FIGURES 326330" captionStartId="92.[151,264,1880,1903]" captionTargetBox="[171,1436,1090,1859]" captionTargetId="figure-82@92.[151,1436,1090,1859]" captionTargetPageId="92" captionText="FIGURES 326330. Isoperla kozlovi. 326: Habitus (preserved); 327: Male terminalia, dorsal; 328: Male, ventral, showing penial sclerites; 329: Female terminalia, ventral; 330: Predicted ENM map with documented occurrences and range estimate." pageId="46" pageNumber="47">Fig. 326</figureCitation>
). The pronotum has a well-delimited pale stripe, otherwise the rest of the disc is dark with darker rugosities. The mesoscutum has anterior golden-yellow markings (
<figureCitation id="13212A7415498119FEDCFF1BFEDD50C1" box="[273,370,188,212]" captionStart="FIGURES 326330" captionStartId="92.[151,264,1880,1903]" captionTargetBox="[171,1436,1090,1859]" captionTargetId="figure-82@92.[151,1436,1090,1859]" captionTargetPageId="92" captionText="FIGURES 326330. Isoperla kozlovi. 326: Habitus (preserved); 327: Male terminalia, dorsal; 328: Male, ventral, showing penial sclerites; 329: Female terminalia, ventral; 330: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 326</figureCitation>
) similar to
<taxonomicName id="4C1A4D7215498119FE37FF1AFDC250C1" authorityName="Samal" authorityYear="1939" box="[506,621,189,212]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="B96EEAE315498119FE37FF1AFDC250C1" box="[506,621,189,212]" italics="true" pageId="47" pageNumber="48">I. asiatica</emphasis>
</taxonomicName>
(
<figureCitation id="13212A7415498119FDB3FF1BFD4D50C1" box="[638,738,188,212]" captionStart="FIGURES 316320" captionStartId="91.[151,264,1883,1906]" captionTargetBox="[156,1436,1093,1862]" captionTargetId="figure-79@91.[151,1436,1093,1862]" captionTargetPageId="91" captionText="FIGURES 316320. Isoperla asiatica. 316: Habitus (preserved); 317: Male terminalia, dorsal; 318: Male, ventral; 319: Female terminalia, ventral; 320: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 316</figureCitation>
),
<taxonomicName id="4C1A4D7215498119FD34FF1AFC2750C1" authorityName="Zhiltzova" authorityYear="1972" box="[761,904,188,212]" class="Insecta" family="Perlodidae" genus="Filchneria" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="B96EEAE315498119FD34FF1AFC2750C1" box="[761,904,188,212]" italics="true" pageId="47" pageNumber="48">I. mongolica</emphasis>
</taxonomicName>
(
<figureCitation id="13212A7415498119FC57FF1BFC5150C1" box="[922,1022,188,212]" captionStart="FIGURES 336340" captionStartId="93.[151,263,1880,1903]" captionTargetBox="[151,1436,1090,1859]" captionTargetId="figure-78@93.[151,1436,1090,1859]" captionTargetPageId="93" captionText="FIGURES 336340. Isoperla mongolica. 336: Habitus (preserved); 337: Male terminalia, dorsal; 338: Male, lateral; 339: Female terminalia, ventral; 340: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 336</figureCitation>
), and
<taxonomicName id="4C1A4D7215498119FB8BFF1AFB4B50C1" baseAuthorityName="Navas" baseAuthorityYear="1923" box="[1094,1252,188,212]" class="Insecta" family="Perlodidae" genus="Kaszabia" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="B96EEAE315498119FB8BFF1AFB4B50C1" box="[1094,1252,188,212]" italics="true" pageId="47" pageNumber="48">K. nigricauda</emphasis>
</taxonomicName>
(
<figureCitation id="13212A7415498119FB3BFF1BFAF450C1" box="[1270,1371,188,212]" captionStart="FIGURES 350354" captionStartId="95.[151,258,942,963]" captionTargetBox="[151,1436,159,928]" captionTargetId="figure-61@95.[151,1436,159,928]" captionTargetPageId="95" captionText="FIGURES 350354. Kaszabia nigricauda. 350: Habitus (preserved); 351: Male terminalia, dorsal; 352: Male abdomen, lateral; 353: Female terminalia, ventral; 354: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 350</figureCitation>
). The male paraprocts are lightly sclerotized, sharp and extend on to tergum 10. The aedeagus has large paired sclerites, which are apparent through the body wall of sternum 10 as if the segment itself is pigmented (
<figureCitation id="13212A7415498119FB62FEA3FABE5109" box="[1199,1297,260,284]" captionStart="FIGURES 326330" captionStartId="92.[151,264,1880,1903]" captionTargetBox="[171,1436,1090,1859]" captionTargetId="figure-82@92.[151,1436,1090,1859]" captionTargetPageId="92" captionText="FIGURES 326330. Isoperla kozlovi. 326: Habitus (preserved); 327: Male terminalia, dorsal; 328: Male, ventral, showing penial sclerites; 329: Female terminalia, ventral; 330: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 328</figureCitation>
). The apices of the sclerites are directed medially towards one another but do not touch. The female subgenital plate is triangular with a rounded apex that is sometimes shallowly notched (
<figureCitation id="13212A7415498119FCEBFEEBFC275171" box="[806,904,332,356]" captionStart="FIGURES 326330" captionStartId="92.[151,264,1880,1903]" captionTargetBox="[171,1436,1090,1859]" captionTargetId="figure-82@92.[151,1436,1090,1859]" captionTargetPageId="92" captionText="FIGURES 326330. Isoperla kozlovi. 326: Habitus (preserved); 327: Male terminalia, dorsal; 328: Male, ventral, showing penial sclerites; 329: Female terminalia, ventral; 330: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 329</figureCitation>
). Overall, the plate resembles that of
<taxonomicName id="4C1A4D7215498119FAE7FEEAFEC85192" authority="(Zetterstedt, 1840)" baseAuthorityName="Zetterstedt" baseAuthorityYear="1840" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B96EEAE315498119FAE7FEEAFA345171" box="[1322,1435,332,356]" italics="true" pageId="47" pageNumber="48">I. obscura</emphasis>
(Zetterstedt, 1840)
</taxonomicName>
(
<figureCitation id="13212A7415498119FEBBFEC8FE795192" box="[374,470,367,391]" captionStart="FIGURES 341344" captionStartId="94.[151,263,983,1006]" captionTargetBox="[157,1434,193,920]" captionTargetId="figure-74@94.[151,1436,193,962]" captionTargetPageId="94" captionText="FIGURES 341344. Isoperla obscura. 341: Habitus (preserved); 342: Male terminalia, dorsal; 343: Female terminalia, ventral; 344: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 343</figureCitation>
) and
<taxonomicName id="4C1A4D7215498119FDD9FED7FD50519D" baseAuthorityName="Navas" baseAuthorityYear="1923" box="[532,767,368,392]" class="Insecta" family="Perlodidae" genus="Kaszabia" kingdom="Animalia" order="Plecoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="B96EEAE315498119FDD9FED7FD50519D" box="[532,767,368,392]" italics="true" pageId="47" pageNumber="48">Kaszabia nigricauda</emphasis>
</taxonomicName>
(
<figureCitation id="13212A7415498119FCC3FEC8FCDF5192" box="[782,880,367,391]" captionStart="FIGURES 350354" captionStartId="95.[151,258,942,963]" captionTargetBox="[151,1436,159,928]" captionTargetId="figure-61@95.[151,1436,159,928]" captionTargetPageId="95" captionText="FIGURES 350354. Kaszabia nigricauda. 350: Habitus (preserved); 351: Male terminalia, dorsal; 352: Male abdomen, lateral; 353: Female terminalia, ventral; 354: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 353</figureCitation>
), but is not as long and only extends slightly onto segment 9. Nymph head coloration (
<figureCitation id="13212A7415498119FDE2FE33FD2151B9" box="[559,654,404,428]" captionStart="FIGURES 302310" captionStartId="90.[151,264,1845,1868]" captionTargetBox="[190,1420,193,1824]" captionTargetId="figure-61@90.[167,1421,193,1824]" captionTargetPageId="90" captionText="FIGURES 302310. Isoperla and Kaszabia nymphs. Illustrations after Teslenko and Zhiltzova (2006). 302: I. altaica; 303: I. asiatica; 304: I. eximia; 305: I. kozlovi; 306: I. lunigera; 307: I. mongolica; 308: I. obscura; 309: I. potanini; 310: K. nigricauda." pageId="47" pageNumber="48">Fig. 305</figureCitation>
) is dark from the epicranial arm to the M-line with no dark markings posterior to the lateral ocelli or on the labrum.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15498119FF0AFE7CFE2D5202" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BA536F115498119FF0AFE7CFE2D5202" blockId="47.[151,1436,151,680]" pageId="47" pageNumber="48">
<emphasis id="B96EEAE315498119FF0AFE7CFDA351E1" bold="true" box="[199,524,475,500]" pageId="47" pageNumber="48">DISTRIBUTION—Global:</emphasis>
East Palearctic—
<emphasis id="B96EEAE315498119FD1AFE7CFCE551E1" bold="true" box="[727,842,475,500]" pageId="47" pageNumber="48">Regional:</emphasis>
AOB, IDB*, POB*—
<emphasis id="B96EEAE315498119FB81FE7CFB0B51E1" bold="true" box="[1100,1188,475,500]" pageId="47" pageNumber="48">Aimag:</emphasis>
AR, BO*, BU, DO*, KhE*, KhG, SE, TO.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15498119FF0AFD84FDFC52B2" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8BA536F115498119FF0AFD84FDFC52B2" blockId="47.[151,1436,151,680]" pageId="47" pageNumber="48">
<emphasis id="B96EEAE315498119FF0AFD84FED45229" bold="true" box="[199,379,547,572]" pageId="47" pageNumber="48">DISCUSSION:</emphasis>
<materialsCitation id="3B723CAC15498119FE47FD83FDE052B2" country="Mongolia" county="This" location="When" municipality="Zhiltzova" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Selenge">
<collectingCounty id="62C44E7D15498119FE47FD83FE145229" box="[394,443,548,572]" pageId="47" pageNumber="48">This</collectingCounty>
species was originally described from
<collectingCountry id="F30D766115498119FC52FD83FBA35229" box="[927,1036,548,572]" name="Mongolia" pageId="47" pageNumber="48">Mongolia</collectingCountry>
by
<collectingMunicipality id="6BC1AC8B15498119FB88FD83FB1F5229" box="[1093,1200,548,572]" pageId="47" pageNumber="48">Zhiltzova</collectingMunicipality>
(1972) in her first contribution to the
<taxonomicName id="4C1A4D7215498119FEB3FDEFFE63524A" box="[382,460,584,607]" class="Insects" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="47" pageNumber="48" phylum="Arthropoda" rank="class">Insects</taxonomicName>
of
<collectingCountry id="F30D766115498119FE30FDE0FDC5524A" box="[509,618,583,607]" name="Mongolia" pageId="47" pageNumber="48">Mongolia</collectingCountry>
series.
<location id="8EC5602A15498119FD06FDE0FCA1524A" LSID="urn:lsid:plazi:treatment:03B387E715488119FF5AF942FDFC52B2:8EC5602A15498119FD06FDE0FCA1524A" box="[715,782,583,607]" country="Mongolia" county="This" municipality="Zhiltzova" name="When" pageId="47" pageNumber="48" stateProvince="Selenge">When</location>
considering only our study, this species appears to be restricted to the
<collectingRegion id="49DEF81315498119FE99FDCBFE025291" box="[340,429,620,644]" country="Mongolia" name="Selenge" pageId="47" pageNumber="48">Selenge</collectingRegion>
<location id="8EC5602A15498119FE7BFDCBFE5B5291" LSID="urn:lsid:plazi:treatment:03B387E715488119FF5AF942FDFC52B2:8EC5602A15498119FE7BFDCBFE5B5291" box="[438,500,620,644]" country="Mongolia" county="This" municipality="Zhiltzova" name="River" pageId="47" pageNumber="48" stateProvince="Selenge">River</location>
basin (
<figureCitation id="13212A7415498119FD86FDCBFD1F5291" box="[587,688,620,644]" captionStart="FIGURES 326330" captionStartId="92.[151,264,1880,1903]" captionTargetBox="[171,1436,1090,1859]" captionTargetId="figure-82@92.[151,1436,1090,1859]" captionTargetPageId="92" captionText="FIGURES 326330. Isoperla kozlovi. 326: Habitus (preserved); 327: Male terminalia, dorsal; 328: Male, ventral, showing penial sclerites; 329: Female terminalia, ventral; 330: Predicted ENM map with documented occurrences and range estimate." pageId="47" pageNumber="48">Fig. 330</figureCitation>
), but is predicted to occur in the eastern
<collectingRegion id="49DEF81315498119FB40FDCBFA9D5291" box="[1165,1330,620,644]" country="Mongolia" name="Hentiy" pageId="47" pageNumber="48">Khentii region</collectingRegion>
which is where the
<typeStatus id="54A1885315498119FEC1FD37FE9352BD" box="[268,316,656,680]" pageId="47" pageNumber="48">type</typeStatus>
specimen was collected
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>