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<document id="CDD9D6839C9E282C993F9D7B3042993D" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFDA1F49FF50FEC8FD368977" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax borhidiana Muniz" docType="treatment" docVersion="2" lastPageNumber="39" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="38" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="CCF030A9680FE302053065BB478D2CE2">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:date id="3409CBC63821AEB024A3ED224466B70A">2023</mods:date>
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<treatment id="03B387DAFFDA1F49FF50FEC8FD368977" ID-DOI="http://doi.org/10.5281/zenodo.8389283" ID-Zenodo-Dep="8389283" LSID="urn:lsid:plazi:treatment:03B387DAFFDA1F49FF50FEC8FD368977" httpUri="http://treatment.plazi.org/id/03B387DAFFDA1F49FF50FEC8FD368977" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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<paragraph id="8BA536CCFFDA1F4AFF50FEC8FD9089C9" blockId="37.[136,1452,369,466]" pageId="37" pageNumber="38">
<emphasis id="B96EEADEFFDA1F4AFF50FEC8FB7C8990" bold="true" box="[136,1157,369,396]" pageId="37" pageNumber="38">
1.7.
<taxonomicName id="4C1A4D4FFFDA1F4AFF62FEC8FDCF8990" ID-CoL="WLXY" ID-ENA="480896" authority="Muniz" authorityName="Muniz" box="[186,566,369,396]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="borhidiana">
<emphasis id="B96EEADEFFDA1F4AFF62FEC8FE198997" bold="true" box="[186,480,369,396]" italics="true" pageId="37" pageNumber="38">Coccothrinax borhidiana</emphasis>
Muñiz
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFDA1F4AFD86FEC8FB868990" author="Borhidi, A. &amp; Imchanitskaya, N. &amp; Muniz, O." box="[606,1151,369,396]" pageId="37" pageNumber="38" pagination="428 - 437" refId="ref61192" refString="Borhidi, A., Imchanitskaya, N. &amp; Muniz, O. (1978) Dendrological novelties in Cuba. Fam.: Arecaceae. Acta Agronomica Academiae Scientarum Hungaricae 27: 428 - 437." type="journal article" year="1978">Borhidi, Imchanitskaya &amp; Muñiz (1978: 437)</bibRefCitation>
.
</emphasis>
<materialsCitation id="3B723C91FFDA1F4AFB56FECBFD9089C9" collectingDate="1970-05-14" collectionCode="O, HAC, BP" country="Cuba" location="Jibacoa" municipality="Punta Seburoco" pageId="37" pageNumber="38" specimenCount="2" stateProvince="Matanzas" typeStatus="lectotype">
<typeStatus id="54A1886EFFDA1F4AFB56FECBFB078997" box="[1166,1278,370,396]" pageId="37" pageNumber="38">Lectotype</typeStatus>
(designated by
<bibRefCitation id="EF8B4B3DFFDA1F4AFF50FE2CFEF089B4" author="Moya, C." box="[136,265,405,431]" pageId="37" pageNumber="38" pagination="1 - 27" refId="ref63288" refString="Moya, C. (2021) An annotated checklist of Cuban palms 6. Coccothrinax, Pt. 2. 1972 - 1995. Nomenclature, typification, and distribution. PalmArbor 3: 1 - 27." type="journal article" year="2021">Moya 2021</bibRefCitation>
):—
<collectingCountry id="F30D765CFFDA1F4AFEECFE2CFE7E89B4" box="[308,391,405,431]" name="Cuba" pageId="37" pageNumber="38">CUBA</collectingCountry>
.
<collectingRegion id="49DEF82EFFDA1F4AFE49FE2CFE0689B4" box="[401,511,405,431]" country="Cuba" name="Matanzas" pageId="37" pageNumber="38">Matanzas</collectingRegion>
,
<collectingMunicipality id="6BC1ACB6FFDA1F4AFDD2FE2CFD4289B4" box="[522,699,405,431]" pageId="37" pageNumber="38">Punta Seburoco</collectingMunicipality>
inter
<location id="8EC56017FFDA1F4AFD23FE2CFCB689B4" LSID="urn:lsid:plazi:treatment:03B387DAFFDA1F49FF50FEC8FD368977:8EC56017FFDA1F4AFD23FE2CFCB689B4" box="[763,847,405,431]" country="Cuba" municipality="Punta Seburoco" name="Jibacoa" pageId="37" pageNumber="38" stateProvince="Matanzas">Jibacoa</location>
y
<collectingRegion id="49DEF82EFFDA1F4AFCB3FE2CFC2089B4" box="[875,985,405,431]" country="Cuba" name="Matanzas" pageId="37" pageNumber="38">Matanzas</collectingRegion>
,
<date id="FFA4100CFFDA1F4AFC3CFE2CFB8589B4" box="[996,1148,405,431]" pageId="37" pageNumber="38" value="1970-05-14">
<collectingDate id="EFE0E9E4FFDA1F4AFC3CFE2CFB8589B4" box="[996,1148,405,431]" pageId="37" pageNumber="38" value="1970-05-14">14 May 1970</collectingDate>
</date>
,
<emphasis id="B96EEADEFFDA1F4AFB5FFE2CFAC089B4" box="[1159,1337,405,431]" italics="true" pageId="37" pageNumber="38">
<collectionCode id="ED0BAE09FFDA1F4AFB5FFE2CFB6789B4" box="[1159,1182,405,431]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="37" pageNumber="38" type="Herbarium">O</collectionCode>
. Muñiz 27118
</emphasis>
(
<typeStatus id="54A1886EFFDA1F4AFA90FE2CFA5589B4" box="[1352,1452,405,431]" pageId="37" pageNumber="38">holotype</typeStatus>
<collectionCode id="ED0BAE09FFDA1F4AFF50FE01FF3289C9" box="[136,203,440,466]" country="Cuba" httpUri="http://grbio.org/cool/cenz-te88" name="Instituto de Ecología y Sistemática" pageId="37" pageNumber="38" type="Herbarium">HAC</collectionCode>
!, isotype
<collectionCode id="ED0BAE09FFDA1F4AFEEAFE01FEAA89C9" box="[306,339,440,466]" country="Hungary" lsid="urn:lsid:biocol.org:col:13261" name="Hungarian Natural History Museum" pageId="37" pageNumber="38" type="Herbarium">BP</collectionCode>
<emphasis id="B96EEADEFFDA1F4AFE83FE00FE7889C9" box="[347,385,441,466]" italics="true" pageId="37" pageNumber="38">n.v.</emphasis>
,
<collectionCode id="ED0BAE09FFDA1F4AFE57FE01FE4989C9" box="[399,432,440,466]" country="Hungary" lsid="urn:lsid:biocol.org:col:13261" name="Hungarian Natural History Museum" pageId="37" pageNumber="38" type="Herbarium">BP</collectionCode>
image!).
<figureCitation id="13212A49FFDA1F4AFDC4FE01FD9089C9" box="[540,617,440,466]" captionStart="PLATE 7" captionStartId="37.[136,214,2022,2043]" captionTargetBox="[151,1433,1059,1996]" captionTargetId="figure-366@37.[150,1435,1058,1998]" captionTargetPageId="37" captionText="PLATE 7. Coccothrinax borhidiana, Cuba, with stems covered with persistent dead leaves, giving a skirt of dead leaves, and short petioles giving a compact crown of leaves. (Image by Héctor M. Díaz)." figureDoi="http://doi.org/10.5281/zenodo.8389365" httpUri="https://zenodo.org/record/8389365/files/figure.png" pageId="37" pageNumber="38">Plate 7</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFDA1F4AFF50FE47FBC58C1D" pageId="37" pageNumber="38" type="description">
<paragraph id="8BA536CCFFDA1F4AFF50FE47FBC58C1D" blockId="37.[136,1452,510,1030]" pageId="37" pageNumber="38">
<emphasis id="B96EEADEFFDA1F4AFF50FE47FF298A03" bold="true" box="[136,208,510,536]" pageId="37" pageNumber="38">Stems</emphasis>
length not recorded, 14.0 cm diameter, solitary.
<emphasis id="B96EEADEFFDA1F4AFD2CFE47FCBF8A03" bold="true" box="[756,838,510,536]" pageId="37" pageNumber="38">Leaves</emphasis>
persistent, giving a skirt of dead leaves covering the stem; leaf sheath fibers 0.5(0.40.6) mm diameter, thin, flimsy, loosely woven, free and greatly elongate at the apices; petioles 16.4(15.218.2) mm diameter just below the apex; palmans 19.8(18.021.5) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 3435 per leaf, the middle ones 58.6(52.062.5) cm long and 3.7(3.04.5) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially; leaf segments densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown center, without transverse veinlets.
<emphasis id="B96EEADEFFDA1F4AFF50FCE7FEC88B63" bold="true" box="[136,305,862,888]" pageId="37" pageNumber="38">Inflorescences</emphasis>
erect, at least initially, amongst or above the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 5; proximalmost rachillae straight, 9.2(7.015.0) cm long and 1.1(1.01.2) mm diameter in fruit; rachillae at or near anthesis not recorded; stamens 8; fruit pedicels 2.8(2.23.2) mm long;
<emphasis id="B96EEADEFFDA1F4AFCC8FC71FCA88BF9" bold="true" box="[784,849,968,994]" pageId="37" pageNumber="38">fruits</emphasis>
6.2(5.86.5) mm long and 7.1(6.38.0) mm diameter, black; fruit surfaces smooth or sometimes with projecting fibers; seeds not recorded.
</paragraph>
</subSubSection>
<caption id="DF656644FFDA1F4AFF50F85FFCCC8004" ID-DOI="http://doi.org/10.5281/zenodo.8389365" ID-Zenodo-Dep="8389365" httpUri="https://zenodo.org/record/8389365/files/figure.png" pageId="37" pageNumber="38" startId="37.[136,214,2022,2043]" targetBox="[151,1433,1059,1996]" targetPageId="37" targetType="plate">
<paragraph id="8BA536CCFFDA1F4AFF50F85FFCCC8004" blockId="37.[136,1451,2022,2080]" pageId="37" pageNumber="38">
<emphasis id="B96EEADEFFDA1F4AFF50F85FFF128FE0" bold="true" box="[136,235,2022,2043]" pageId="37" pageNumber="38">PLATE 7</emphasis>
.
<taxonomicName id="4C1A4D4FFFDA1F4AFF21F85FFE178FE0" box="[249,494,2022,2043]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="borhidiana">
<emphasis id="B96EEADEFFDA1F4AFF21F85FFE178FE0" box="[249,494,2022,2043]" italics="true" pageId="37" pageNumber="38">Coccothrinax borhidiana</emphasis>
</taxonomicName>
, Cuba, with stems covered with persistent dead leaves, giving a skirt of dead leaves, and short petioles giving a compact crown of leaves. (Image by Héctor M. Díaz).
</paragraph>
</caption>
<subSubSection id="C3006547FFD91F49FF65FF27FD368977" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BA536CCFFD91F49FF65FF27FB6D88C7" blockId="38.[136,1452,158,365]" pageId="38" pageNumber="39">
<emphasis id="B96EEADEFFD91F49FF65FF27FDF188A3" bold="true" box="[189,520,158,184]" pageId="38" pageNumber="39">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFD91F49FDD0FF27FDBA88A3" box="[520,579,158,184]" name="Cuba" pageId="38" pageNumber="39">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFD91F49FD8BFF27FD3B88A3" box="[595,706,158,184]" country="Cuba" name="Matanzas" pageId="38" pageNumber="39">Matanzas</collectingRegion>
) (
<figureCitation id="13212A49FFD91F49FD02FF27FCD488A3" box="[730,813,158,184]" captionStart="FIGURE 10" captionStartId="32.[136,229,1508,1530]" captionTargetBox="[155,1429,198,1479]" captionTargetId="figure-27@32.[150,1434,193,1484]" captionTargetPageId="32" captionText="FIGURE 10. Distribution of Coccothrinax baracoensis, C. bonettiana, C. borhidiana, and C. boschiana." figureDoi="http://doi.org/10.5281/zenodo.8389355" httpUri="https://zenodo.org/record/8389355/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
), open areas on sandy or rocky soils near the sea at
<quantity id="4CE29B29FFD91F49FAA7FF26FA5288A2" box="[1407,1451,159,185]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="7.0">7 m</quantity>
elevation.
<bibRefCitation id="EF8B4B3DFFD91F49FF25FF7BFE7188C7" author="Craft, P." box="[253,392,194,220]" pageId="38" pageNumber="39" refId="ref61560" refString="Craft, P. (2017) The Palms of Cuba. Palm Nut Pages, Florida, 232 pp." type="book" year="2017">Craft (2017)</bibRefCitation>
described the habitat as littoral scrub forest on dogs tooth limestone.
</paragraph>
<paragraph id="8BA536CCFFD91F49FF65FF5FFD368977" blockId="38.[136,1452,158,365]" pageId="38" pageNumber="39">
<emphasis id="B96EEADEFFD91F49FF65FF5FFE55891B" bold="true" box="[189,428,230,256]" pageId="38" pageNumber="39">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFD91F49FD16FF5FFC10891B" box="[718,1001,230,256]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="borhidiana">
<emphasis id="B96EEADEFFD91F49FD16FF5FFC10891B" box="[718,1001,230,256]" italics="true" pageId="38" pageNumber="39">Coccothrinax borhidiana</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. It appears similar to
<taxonomicName id="4C1A4D4FFFD91F49FBE5FEB2FB55893F" authorityName="Beccari" authorityYear="1907" baseAuthorityName="Hoyt" box="[1085,1196,267,293]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="crinita">
<emphasis id="B96EEADEFFD91F49FBE5FEB2FB55893F" box="[1085,1196,267,293]" italics="true" pageId="38" pageNumber="39">C. crinita</emphasis>
</taxonomicName>
in its elongate sheath fibers, differing in its persistent leaves, giving a skirt of dead leaves covering the stem and smaller fruits. Petioles are relatively short, hence the compact crown of leaves.
</paragraph>
</subSubSection>
</treatment>
</document>