364 lines
33 KiB
XML
364 lines
33 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6103236" ID-GBIF-Dataset="f0d443f2-b8b9-46fa-a1aa-2d0f0d3c8748" ID-GBIF-Taxon="114923350" ID-ISSN="1175-5326" ID-Zenodo-Dep="6103236" ID-ZooBank="CEAB4AA5-C4F8-437E-A2F4-D82F383579C1" checkinTime="1447288886152" checkinUser="donat" docAuthor="Taylor, Robert W." docDate="2015" docId="03B3878D9B1FFFE6FF534E319D1FFD57" docLanguage="en" docName="29627.pdf.imf" docOrigin="Zootaxa 3947 (2)" docTitle="Heteroponera relicta Wheeler" docType="treatment" docVersion="5" lastPageId="10" lastPageNumber="161" masterDocId="FF8AFFF59B18FFECFFC44D279F4AF867" masterDocTitle="Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus" masterLastPageNumber="180" masterPageNumber="151" pageId="7" pageNumber="158" updateTime="1645000973839" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Taylor, Robert W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>3947</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>151</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3947.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="ZooBank">CEAB4AA5-C4F8-437E-A2F4-D82F383579C1</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6103236" ID-GBIF-Taxon="114923350" ID-Zenodo-Dep="6103236" LSID="urn:lsid:plazi:treatment:03B3878D9B1FFFE6FF534E319D1FFD57" httpUri="http://treatment.plazi.org/id/03B3878D9B1FFFE6FF534E319D1FFD57" lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158">
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<subSubSection pageId="7" pageNumber="158" type="nomenclature">
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<paragraph box="[151,564,790,816]" pageId="7" pageNumber="158">
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<emphasis bold="true" box="[151,564,790,816]" pageId="7" pageNumber="158">
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<heading bold="true" box="[151,564,790,816]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
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1.
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<taxonomicName authority="Wheeler" authorityName="Wheeler" box="[180,564,790,816]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis bold="true" box="[180,431,790,816]" italics="true" pageId="7" pageNumber="158">Heteroponera relicta</emphasis>
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(Wheeler)
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</taxonomicName>
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</heading>
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</emphasis>
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</paragraph>
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<paragraph box="[151,323,825,850]" pageId="7" pageNumber="158">
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(
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<figureCitation box="[159,259,825,850]" captionStart="FIGURES 2 – 5" captionStartId="8.[151,263,1953,1975]" captionTargetBox="[189,1397,196,1931]" captionTargetId="figure@8.[189,1397,193,1931]" captionTargetPageId="8" captionText="FIGURES 2 – 5. Heteroponera relicta (Wheeler), worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68 mm); Figs 3 – 5 Black Mountain Rd, Kuranda (HW = 1.27 mm, PW = 0.91 mm, GW = 1.05 mm)." pageId="7" pageNumber="158">Figs 2–5</figureCitation>
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,
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<figureCitation box="[271,314,825,850]" captionStart="FIGURES 6 – 9" captionStartId="9.[151,263,1954,1976]" captionTargetBox="[187,1399,193,1931]" captionTargetId="figure@9.[187,1400,193,1932]" captionTargetPageId="9" captionText="FIGURES 6 – 9. Heteroponera relicta (Wheeler), gyne (Malanda Falls), standard views (HW = 1.28 mm, PW = 0.97 mm, WL = 1.59 mm, GW = 1.58 mm)." pageId="7" pageNumber="158">6–9</figureCitation>
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)
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</paragraph>
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<paragraph pageId="7" pageNumber="158">
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<taxonomicName authority="Wheeler, 1915: 118" authorityName="Wheeler" authorityPageNumber="118" authorityYear="1915" box="[151,600,897,919]" class="Insecta" family="Formicidae" genus="Paranomopone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis box="[151,382,897,919]" italics="true" pageId="7" pageNumber="158">Paranomopone relicta</emphasis>
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Wheeler, 1915: 118
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</taxonomicName>
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, pl. 8; Worker, Gyne; Type locality:
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<emphasis bold="true" box="[996,1145,897,919]" pageId="7" pageNumber="158">AUSTRALIA</emphasis>
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: QUEENSLAND:
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<taxonomicName box="[1349,1436,897,919]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
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(16°49'S, 145°38'E). Holotype examined (Museum of Comparative Zoology, Cambridge, Massachusetts, USA).
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<taxonomicName authority="W.L. Brown, 1958: 196" authorityName="W.L. Brown" authorityPageNumber="196" authorityYear="1958" box="[151,612,962,984]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis box="[151,364,962,984]" italics="true" pageId="7" pageNumber="158">Heteroponera relicta</emphasis>
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W.L. Brown, 1958: 196
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="158" type="distribution">
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<paragraph pageId="7" pageNumber="158">
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<emphasis bold="true" box="[151,541,1029,1053]" pageId="7" pageNumber="158">Distribution, material examined.</emphasis>
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Abundantly known from many localities centered on the Atherton Tableland of NE Qld, north to near
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<taxonomicName box="[407,516,1066,1089]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="165" phylum="Arthropoda" rank="genus">Mossman</taxonomicName>
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and south almost to the latitude of Townsville (Grid cells -16/145, -17/145 and - 18/146), as follows:
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<emphasis bold="true" box="[384,741,1101,1125]" pageId="7" pageNumber="158">EA4 (Mt Carbine Tableland):</emphasis>
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<taxonomicName authority="Bluff Track" authorityName="Bluff Track" box="[749,990,1101,1125]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis box="[749,990,1101,1125]" italics="true" pageId="7" pageNumber="158">Mossman Bluff Track</emphasis>
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</taxonomicName>
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;
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<taxonomicName authority="Gorge" authorityName="Gorge" box="[1006,1192,1102,1125]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis box="[1006,1192,1102,1125]" italics="true" pageId="7" pageNumber="158">Mossman Gorge</emphasis>
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</taxonomicName>
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;
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<emphasis box="[1208,1308,1102,1125]" italics="true" pageId="7" pageNumber="158">Mt Demi</emphasis>
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;
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<emphasis box="[1324,1428,1102,1125]" italics="true" pageId="7" pageNumber="158">Mt Lewis</emphasis>
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;
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<emphasis box="[151,391,1137,1162]" italics="true" pageId="7" pageNumber="158">2.5 km N of Mt Lewis</emphasis>
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;
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<emphasis box="[406,679,1137,1162]" italics="true" pageId="7" pageNumber="158">2 km SE of Mt Spurgeon</emphasis>
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;
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<emphasis box="[694,797,1137,1162]" italics="true" pageId="7" pageNumber="158">The Bluff</emphasis>
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(11 km W
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<taxonomicName box="[924,1038,1137,1162]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="165" phylum="Arthropoda" rank="genus">Mossman</taxonomicName>
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);
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<emphasis box="[1058,1183,1137,1162]" italics="true" pageId="7" pageNumber="158">Pauls Luck</emphasis>
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;
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<taxonomicName authority="Creek." authorityName="Creek." box="[1198,1375,1137,1162]" class="Insecta" family="Curculionidae" genus="Platypus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis box="[1198,1369,1137,1162]" italics="true" pageId="7" pageNumber="158">Platypus Creek</emphasis>
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.
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</taxonomicName>
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<emphasis bold="true" pageId="7" pageNumber="158">EA6 (Black Mountain):</emphasis>
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<emphasis box="[382,563,1173,1197]" italics="true" pageId="7" pageNumber="158">Black Mountain</emphasis>
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(17 km ESE Julatten);
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<emphasis box="[832,1052,1173,1197]" italics="true" pageId="7" pageNumber="158">Black Mountain Rd</emphasis>
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(via
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<taxonomicName box="[1112,1214,1173,1197]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
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);
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<emphasis box="[1234,1384,1173,1197]" italics="true" pageId="7" pageNumber="158">Clacherty Rd</emphasis>
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(via Julatten);
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<emphasis box="[267,366,1209,1234]" italics="true" pageId="7" pageNumber="158">
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<taxonomicName box="[267,366,1209,1234]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
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</emphasis>
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;
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<emphasis box="[386,555,1209,1234]" italics="true" pageId="7" pageNumber="158">Lyons Lookout</emphasis>
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(near
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<taxonomicName box="[636,750,1209,1234]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="165" phylum="Arthropoda" rank="genus">Mossman</taxonomicName>
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).
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<emphasis bold="true" box="[773,1025,1209,1234]" pageId="7" pageNumber="158">EA7 (Lamb Range):</emphasis>
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<emphasis box="[1036,1235,1209,1234]" italics="true" pageId="7" pageNumber="158">Davies Creek Rd</emphasis>
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;
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<emphasis box="[1254,1428,1209,1234]" italics="true" pageId="7" pageNumber="158">Emerald Creek</emphasis>
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;
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<emphasis box="[151,293,1245,1269]" italics="true" pageId="7" pageNumber="158">Lamb Range</emphasis>
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.
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<emphasis bold="true" box="[308,724,1245,1269]" pageId="7" pageNumber="158">EA8 (Walsh/ Hugh Nelson Range):</emphasis>
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<taxonomicName authority="Fire Track" authorityName="Fire Track" box="[731,926,1245,1269]" class="Insecta" family="Staphylinidae" genus="Bartle" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis box="[731,926,1245,1269]" italics="true" pageId="7" pageNumber="158">Bartle Fire Track</emphasis>
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</taxonomicName>
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(17 km W Millaa Millaa);
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<emphasis box="[1232,1428,1245,1269]" italics="true" pageId="7" pageNumber="158">Herberton Range</emphasis>
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;
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<emphasis box="[151,376,1281,1306]" italics="true" pageId="7" pageNumber="158">Hugh Nelson Range</emphasis>
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(2.5 km S of Crater NP);
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<emphasis box="[668,1026,1281,1306]" italics="true" pageId="7" pageNumber="158">Mt Fisher; 21 km S of Atherton.</emphasis>
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<emphasis bold="true" box="[1032,1354,1281,1306]" pageId="7" pageNumber="158">EA9 (Atherton Tableland):</emphasis>
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<emphasis italics="true" pageId="7" pageNumber="158">3.2 km E of Lake Barrine</emphasis>
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;
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<emphasis box="[372,565,1317,1341]" italics="true" pageId="7" pageNumber="158">Lake Barrine NP</emphasis>
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;
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<emphasis box="[581,779,1317,1341]" italics="true" pageId="7" pageNumber="158">Lake Eacham NP</emphasis>
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;
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<emphasis box="[796,962,1317,1341]" italics="true" pageId="7" pageNumber="158">Malanda Falls</emphasis>
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;
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<emphasis box="[978,1199,1317,1341]" italics="true" pageId="7" pageNumber="158">12 km SE of Millaa</emphasis>
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;
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<emphasis box="[1215,1428,1317,1341]" italics="true" pageId="7" pageNumber="158">7 km SW of Millaa</emphasis>
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;
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<emphasis box="[151,337,1353,1378]" italics="true" pageId="7" pageNumber="158">Danbulla Forest</emphasis>
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;
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<taxonomicName box="[353,465,1353,1378]" class="Insecta" family="Staphylinidae" genus="Gadgarra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis box="[353,465,1353,1378]" italics="true" pageId="7" pageNumber="158">Gadgarra</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" box="[481,757,1353,1378]" pageId="7" pageNumber="158">
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EA10 (
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<taxonomicName authority="Ker" authorityName="Ker" box="[564,743,1353,1378]" class="Insecta" family="Staphylinidae" genus="Bellenden" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Bellenden Ker</taxonomicName>
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):
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</emphasis>
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<emphasis box="[765,1008,1353,1378]" italics="true" pageId="7" pageNumber="158">
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<taxonomicName authority="Ker Range" authorityName="Ker Range" box="[765,1008,1353,1378]" class="Insecta" family="Staphylinidae" genus="Bellenden" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Bellenden Ker Range</taxonomicName>
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</emphasis>
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(E slope, sited near various specified cable-car towers);
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<emphasis box="[365,452,1390,1413]" italics="true" pageId="7" pageNumber="158">Boonjie</emphasis>
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(13 km ESE Malanda);
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<emphasis box="[742,808,1390,1413]" italics="true" pageId="7" pageNumber="158">Topaz</emphasis>
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area.
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<emphasis bold="true" box="[882,1303,1389,1413]" pageId="7" pageNumber="158">EA11 (Malbon Thompson Range):</emphasis>
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<emphasis box="[1314,1436,1390,1413]" italics="true" pageId="7" pageNumber="158">Mt Gorton</emphasis>
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(near Yarrabah);
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<emphasis box="[346,531,1425,1450]" italics="true" pageId="7" pageNumber="158">Bell Peak North</emphasis>
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(near Gordonvale).
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<emphasis bold="true" box="[765,1092,1425,1450]" pageId="7" pageNumber="158">EA12 (Walter Hill Range):</emphasis>
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<emphasis box="[1102,1328,1425,1450]" italics="true" pageId="7" pageNumber="158">25 km SE of Millaa</emphasis>
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;
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<emphasis italics="true" pageId="7" pageNumber="158">Boulder Creek</emphasis>
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(via Tully);
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<emphasis box="[361,523,1461,1485]" italics="true" pageId="7" pageNumber="158">Downie Creek</emphasis>
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;
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<emphasis box="[539,731,1461,1485]" italics="true" pageId="7" pageNumber="158">Koombooloomba</emphasis>
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;
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<emphasis box="[747,980,1461,1485]" italics="true" pageId="7" pageNumber="158">Tully River Crossing</emphasis>
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;
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<emphasis box="[996,1162,1461,1485]" italics="true" pageId="7" pageNumber="158">Tully Falls NP</emphasis>
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;
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<emphasis box="[1178,1396,1461,1485]" italics="true" pageId="7" pageNumber="158">Crawfords Lookout</emphasis>
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;
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<emphasis italics="true" pageId="7" pageNumber="158">W of McNamee Creek</emphasis>
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.
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<emphasis box="[383,534,1497,1522]" italics="true" pageId="7" pageNumber="158">Laceys Creek</emphasis>
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;
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<emphasis box="[550,738,1497,1522]" italics="true" pageId="7" pageNumber="158">McNamee Creek</emphasis>
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(15 km SW of Innisfail);
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<emphasis box="[1035,1237,1497,1522]" italics="true" pageId="7" pageNumber="158">Mt Father Clancy</emphasis>
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(9 km S of Milla Milla);
|
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<emphasis box="[242,417,1533,1557]" italics="true" pageId="7" pageNumber="158">Palmerston NP</emphasis>
|
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.
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<emphasis bold="true" box="[437,873,1533,1557]" pageId="7" pageNumber="158">EA13 (Kirrama/ Cardwell Range):</emphasis>
|
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<emphasis box="[886,1095,1533,1557]" italics="true" pageId="7" pageNumber="158">MacAlister Range</emphasis>
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(W slope).
|
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<emphasis bold="true" pageId="7" pageNumber="158">EA16 (Paluma/ Bluewater Ranges):</emphasis>
|
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<emphasis box="[393,484,1570,1593]" italics="true" pageId="7" pageNumber="158">Mt Spec</emphasis>
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.
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</paragraph>
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<paragraph pageId="7" pageNumber="158">
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All samples with ecological data are labelled “Rainforest” or “RF”. Relevant collectors include: E.H. Bourne, W.L. Brown, A.N. Burns, H. Janetzki, DWC, JDF, BBL, GBM, RWT, GIT, DKY.
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<emphasis box="[1092,1329,1641,1666]" italics="true" pageId="7" pageNumber="158">
|
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<taxonomicName box="[1092,1329,1641,1666]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="relicta">Heteroponera relicta</taxonomicName>
|
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</emphasis>
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has been identified here from QMBA berlesates numbered: 238, 249, 298, 300, 312, 365, 408, 413, 466, 549, 598, 609, 611, 612 and 618.
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</paragraph>
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<paragraph pageId="7" pageNumber="158">
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<emphasis bold="true" box="[199,497,1749,1773]" pageId="7" pageNumber="158">Elevational distribution.</emphasis>
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Specified elevations range from 60m (
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<taxonomicName authority="Gorge" authorityName="Gorge" box="[977,1173,1749,1773]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">Mossman Gorge</taxonomicName>
|
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) to 1000–1200m (Mt Spurgeon, Lamb Range and Herberton Range).
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</paragraph>
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</subSubSection>
|
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<subSubSection lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158" type="description">
|
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<paragraph lastPageId="10" lastPageNumber="161" pageId="7" pageNumber="158">
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<emphasis bold="true" box="[199,417,1821,1845]" pageId="7" pageNumber="158">Worker diagnosis.</emphasis>
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General features, conformation, color and major sculpturation as in
|
||
<figureCitation box="[1191,1289,1821,1845]" captionStart="FIGURES 2 – 5" captionStartId="8.[151,263,1953,1975]" captionTargetBox="[189,1397,196,1931]" captionTargetId="figure@8.[189,1397,193,1931]" captionTargetPageId="8" captionText="FIGURES 2 – 5. Heteroponera relicta (Wheeler), worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68 mm); Figs 3 – 5 Black Mountain Rd, Kuranda (HW = 1.27 mm, PW = 0.91 mm, GW = 1.05 mm)." pageId="7" pageNumber="158">Figs 2–5</figureCitation>
|
||
and relevant key couplets above. Body color frequently more nearly black than illustrated. Described in adequate detail by Wheeler (1915). Readily identified by key characters cited above and the accompanying illustrations. Humeral angles and pronotal epaulets well-developed. Scrobes generally smooth and shining, usually with very light rippling of the surfaces of the medial scrobal areas. Propodeal declivity smooth and shining, sometimes with slight traces of irregular longitudinal sculpturing. Lateral margins of declivity simple, essentially the unelaborated edges of the lateral and dorsal propodeal sculpturation. A small smooth, shining area medially on the posterior surface of the node.
|
||
<emphasis box="[267,399,189,212]" italics="true" pageId="10" pageNumber="161">Dimensions</emphasis>
|
||
: (smallest available specimen (Black Mountain Rd,
|
||
<taxonomicName box="[1011,1112,188,212]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
|
||
), largest available specimen (Downey Creek)): TL (ca): 5.1, 6.0; HW: 1.19, 1.47; HL: 1.14, 1.34; CI: 104, 109; EL: 0. 19, 0.24; SL: 0.70, 0.82; SI: 58, 56; PW: 0.83, 1.07; WL: 1.42, 1.72; petH: –, 0.96; petW: 0.43, 0.62; GW 0.98, 1.21.
|
||
</paragraph>
|
||
<caption pageId="8" pageNumber="159" targetBox="[189,1397,196,1931]" targetPageId="8">
|
||
<paragraph pageId="8" pageNumber="159">
|
||
<emphasis bold="true" box="[151,315,1953,1975]" pageId="8" pageNumber="159">FIGURES 2–5.</emphasis>
|
||
<taxonomicName authority="Wheeler" authorityName="Wheeler" box="[322,648,1953,1975]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[322,536,1953,1975]" italics="true" pageId="8" pageNumber="159">Heteroponera relicta</emphasis>
|
||
(Wheeler)
|
||
</taxonomicName>
|
||
, worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68mm); Figs 3–5 Black Mountain Rd,
|
||
<taxonomicName box="[363,450,1983,2005]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
|
||
(HW = 1.27mm, PW = 0.91mm, GW = 1.05mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="9" pageNumber="160" targetBox="[187,1399,193,1931]" targetPageId="9">
|
||
<paragraph pageId="9" pageNumber="160">
|
||
<emphasis bold="true" box="[151,314,1954,1976]" pageId="9" pageNumber="160">FIGURES 6–9.</emphasis>
|
||
<taxonomicName authority="Wheeler" authorityName="Wheeler" box="[320,644,1954,1976]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[320,533,1954,1976]" italics="true" pageId="9" pageNumber="160">Heteroponera relicta</emphasis>
|
||
(Wheeler)
|
||
</taxonomicName>
|
||
, gyne (Malanda Falls), standard views (HW = 1.28mm, PW = 0.97mm, WL = 1.59mm, GW = 1.58mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="10" pageNumber="161">
|
||
<emphasis box="[199,258,296,320]" italics="true" pageId="10" pageNumber="161">Gyne</emphasis>
|
||
: General features and major sculpturation as in
|
||
<figureCitation box="[863,971,295,320]" captionStart="FIGURES 6 – 9" captionStartId="9.[151,263,1954,1976]" captionTargetBox="[187,1399,193,1931]" captionTargetId="figure@9.[187,1400,193,1932]" captionTargetPageId="9" captionText="FIGURES 6 – 9. Heteroponera relicta (Wheeler), gyne (Malanda Falls), standard views (HW = 1.28 mm, PW = 0.97 mm, WL = 1.59 mm, GW = 1.58 mm)." pageId="10" pageNumber="161">Figs 6–9</figureCitation>
|
||
. Head size, proportions, structure and sculpturation close to the worker range. One specimen (illustrated here) has an apparently unreduced median ocellus (as did Wheeler’s (1915)
|
||
<taxonomicName box="[535,632,367,392]" class="Insecta" family="Derbidae" genus="Kuranda" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="158" phylum="Arthropoda" rank="genus">Kuranda</taxonomicName>
|
||
gyne). Mesosoma remarkably worker-like (considering that structural reduction from alate ancestry must have occurred separately from that in the evolution of the worker caste), with vestigial traces of ancestral alate-related structure as illustrated; sculpturation less strongly expressed than in worker; PW and WL generally within the worker range, WL ranging slightly higher.
|
||
<emphasis box="[1111,1243,477,500]" italics="true" pageId="10" pageNumber="161">Dimensions</emphasis>
|
||
: Four specimens from colonies collected at
|
||
<taxonomicName authority="Gorge, Malanda Falls, Lake Eacham" authorityName="Gorge, Malanda Falls, Lake Eacham" box="[451,984,511,536]" class="Insecta" family="Staphylinidae" genus="Mossman" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">Mossman Gorge, Malanda Falls, Lake Eacham</taxonomicName>
|
||
, and Palmerston National Park have the following dimensions: TL (ca): 5.7, 6.6, 6.5, 6.5; HW: 1.28, 1.38, 1.38, 1.40; HL: 1.23, 1.31, 1.33, 1.30; CI: 104, 105, 103, 107; EL: 0.23, 0.25, 0.25, 0.26; SL: 0.72, 0.76, 0.75, 0.74; SI: 56, 55, 54, 53; PW: 0.97, 0.97, 1.02, 1.06; WL: 1.59, 1.75, 1.73, 1.79; petH: -, 1.15, 1.00, 1.11; petW: 0.70, 0.81, 0.75, 0.72; GW (across gastral segment III): 1.58, 1.68, 1.69, 1.66. Two specimens from
|
||
<taxonomicName authority="Creek" authorityName="Creek" box="[644,815,655,680]" class="Insecta" family="Curculionidae" genus="Platypus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">Platypus Creek</taxonomicName>
|
||
have HW 1.36mm.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="161">
|
||
<emphasis bold="true" box="[199,356,693,716]" pageId="10" pageNumber="161">Related taxa.</emphasis>
|
||
<emphasis box="[363,531,692,716]" italics="true" pageId="10" pageNumber="161">
|
||
<taxonomicName box="[363,531,692,716]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="rhodopygea">H. rhodopygea</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis box="[585,692,692,716]" italics="true" pageId="10" pageNumber="161">
|
||
<taxonomicName box="[585,692,692,716]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">H. relicta</taxonomicName>
|
||
</emphasis>
|
||
are closely similar and apparently phylogenetically cognate. They are not known to occur sympatrically, though both are recorded from (well separated) localities in
|
||
<emphasis bold="true" box="[1276,1329,727,752]" pageId="10" pageNumber="161">EA4</emphasis>
|
||
(Carbine Tableland) and otherwise have extensive, well-defined allopatric distributional ranges.
|
||
<taxonomicName box="[1193,1436,764,788]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[1193,1436,764,788]" italics="true" pageId="10" pageNumber="161">Heteroponera relicta</emphasis>
|
||
</taxonomicName>
|
||
workers are larger on average. HW of 40 randomly selected.
|
||
<taxonomicName box="[861,994,799,824]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="rhodopygea">
|
||
<emphasis box="[861,994,799,824]" italics="true" pageId="10" pageNumber="161">rhodopygea</emphasis>
|
||
</taxonomicName>
|
||
workers ranged from 1.18 to 1.39mm (mean 1.32mm, distribution roughly bimodal with modes at 1.28mm and 1.34mm. The main range was 1.26mm to 1.39mm, with a divergent small individual at I.18mm (see full measurements below). HW of 40 randomly selected
|
||
<emphasis box="[151,260,908,932]" italics="true" pageId="10" pageNumber="161">
|
||
<taxonomicName box="[151,260,908,932]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">H. relicta</taxonomicName>
|
||
</emphasis>
|
||
workers ranged from 1.19mm to 1.47mm (mean 1.36mm) with frequency distribution essentially flat. The main range was 1.32–1.47mm, with 4 small (?nanitic) individuals in the range 1.20–1.28mm. Thus 42% of measured
|
||
<taxonomicName box="[269,380,980,1004]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[269,380,980,1004]" italics="true" pageId="10" pageNumber="161">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
workers were larger than any measured
|
||
<taxonomicName box="[864,998,980,1004]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="rhodopygea">
|
||
<emphasis box="[864,998,980,1004]" italics="true" pageId="10" pageNumber="161">rhodopygea</emphasis>
|
||
</taxonomicName>
|
||
worker. These comparisons could be affected by generational differences in stature among workers in growing colonies, and by geographical variation in maximum worker size.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="161">
|
||
<taxonomicName box="[199,543,1087,1112]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="darlingtonorum">
|
||
<emphasis box="[199,543,1087,1112]" italics="true" pageId="10" pageNumber="161">Heteroponera darlingtonorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[605,724,1087,1112]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="wilsoni">
|
||
<emphasis box="[605,724,1087,1112]" italics="true" pageId="10" pageNumber="161">H. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
are both similar to
|
||
<taxonomicName box="[962,1072,1087,1112]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[962,1072,1087,1112]" italics="true" pageId="10" pageNumber="161">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
, but with distinctive sculptural differences, and somewhat simplified morphological structural detail, especially in
|
||
<taxonomicName box="[1093,1209,1124,1148]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="wilsoni">
|
||
<emphasis box="[1093,1209,1124,1148]" italics="true" pageId="10" pageNumber="161">H. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
. In both species the propodeum is dorsally slightly inflated and less strongly sloping posterodorsally in profile view than in
|
||
<taxonomicName box="[1312,1418,1159,1184]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[1312,1418,1159,1184]" italics="true" pageId="10" pageNumber="161">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="161">
|
||
<emphasis bold="true" box="[199,588,1196,1220]" pageId="10" pageNumber="161">In-group sympatric associations.</emphasis>
|
||
Sympatric variously with
|
||
<taxonomicName box="[884,1061,1196,1220]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="pendergrasti">
|
||
<emphasis box="[884,1061,1196,1220]" italics="true" pageId="10" pageNumber="161">H. pendergrasti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[1115,1230,1196,1220]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="wilsoni">
|
||
<emphasis box="[1115,1230,1196,1220]" italics="true" pageId="10" pageNumber="161">H. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
(see those species entries for details), with
|
||
<taxonomicName box="[425,567,1233,1256]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="ecarinata">
|
||
<emphasis box="[425,567,1233,1256]" italics="true" pageId="10" pageNumber="161">H. ecarinata</emphasis>
|
||
</taxonomicName>
|
||
at Crawfords Lookout, and with
|
||
<taxonomicName box="[939,1151,1231,1256]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="darlingtonorum">
|
||
<emphasis box="[939,1151,1231,1256]" italics="true" pageId="10" pageNumber="161">H. darlingtonorum</emphasis>
|
||
</taxonomicName>
|
||
in the Paluma area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="161" type="discussion">
|
||
<paragraph pageId="10" pageNumber="161">
|
||
<emphasis bold="true" box="[199,315,1269,1292]" pageId="10" pageNumber="161">Remarks.</emphasis>
|
||
<taxonomicName box="[330,444,1268,1292]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis box="[330,444,1268,1292]" italics="true" pageId="10" pageNumber="161">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
was included in Feitosa’s (2011) phylogenetic analysis as a sister species to
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="rhodopygea">
|
||
<emphasis italics="true" pageId="10" pageNumber="161">H. rhodopygea</emphasis>
|
||
</taxonomicName>
|
||
(his “
|
||
<taxonomicName box="[352,509,1305,1328]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[352,509,1305,1328]" italics="true" pageId="10" pageNumber="161">Heteroponera</emphasis>
|
||
</taxonomicName>
|
||
sp A”).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |