628 lines
65 KiB
XML
628 lines
65 KiB
XML
<document id="E82E038718C8F00C6DA175C5D4819D43" ID-CLB-Dataset="26573" ID-DOI="10.11646/zootaxa.4613.3.9" ID-GBIF-Dataset="38fd24df-4471-45a4-ac5a-e85c57f94b07" ID-ISSN="1175-5326" ID-Zenodo-Dep="3239933" ID-ZooBank="D9CF5E43-22C7-4658-A747-286DAAE4CC9E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559796795157" checkinUser="plazi" docAuthor="Ziegler, Thomas, Pham, Cuong The, Nguyen, Tan Van, Nguyen, Truong Quang, Wang, Jian, Wang, Ying-Yong, Stuart, Bryan L. & Le, Minh Duc" docDate="2019" docId="03AAD15A7F6C0D63FF6CF920FE07F7D5" docLanguage="en" docName="zootaxa.4613.3.9.pdf" docOrigin="Zootaxa 4613 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opisthotropis haihaensis Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="586" masterDocId="FF93A9227F6E0D65FFFBFFA1FFCFF65E" masterDocTitle="A new species of Opisthotropis from northern Vietnam previously misidentified as the Yellow-spotted Mountain Stream Keelback O. maculosa Stuart & Chuaynkern, 2007 (Squamata: Natricidae)" masterLastPageNumber="586" masterPageNumber="579" pageNumber="579" updateTime="1698733350685" updateUser="ExternalLinkService">
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<mods:title id="928DC8ABCAE4B5A27981D2034501C3AE">A new species of Opisthotropis from northern Vietnam previously misidentified as the Yellow-spotted Mountain Stream Keelback O. maculosa Stuart & Chuaynkern, 2007 (Squamata: Natricidae)</mods:title>
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<mods:namePart id="A50F2D5C5717F0653C662944BB7804BB">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="5AB548914C7F2819B83F1B0C4F3400A4">Pham, Cuong The</mods:namePart>
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<mods:namePart id="006B681E665DB7A7B8EE9FCE11CFF7DC">Nguyen, Tan Van</mods:namePart>
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<mods:namePart id="1AE7662CA9E2F8F293DC65AA764E986B">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="433FB49CCEDB08F53086E581BC09765E">Wang, Jian</mods:namePart>
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<mods:namePart id="B0FFFC61D040219A940FF636AB83A934">Wang, Ying-Yong</mods:namePart>
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<mods:namePart id="E04E2CC16A1C88920364AFFA695682E9">Stuart, Bryan L.</mods:namePart>
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<mods:namePart id="FEB7D9A04AC4B6A0BB4B2D6E55BC64A8">Le, Minh Duc</mods:namePart>
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<mods:title id="9FF749D7ECD9AB1BAAB1B3D523D495E8">Zootaxa</mods:title>
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<mods:part id="66743E055E6A690CE4EB6006482A8C41">
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<mods:date id="4448C71BF4AE43E2A4762DC47A292102">2019</mods:date>
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<mods:number id="0A3CE8D2A8D935EF3C9DD736979EA2BD">2019-06-06</mods:number>
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<mods:number id="0802445F32F259B47F0CF177FA397FD8">4613</mods:number>
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<mods:start id="E555EB2CE7933B88E510E964EC4C7C5B">579</mods:start>
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<mods:identifier id="83DA3A6060D25F0B6AAA277E82C29EFE" type="DOI">10.11646/zootaxa.4613.3.9</mods:identifier>
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<treatment id="03AAD15A7F6C0D63FF6CF920FE07F7D5" ID-GBIF-Taxon="157422531" LSID="urn:lsid:plazi:treatment:03AAD15A7F6C0D63FF6CF920FE07F7D5" httpUri="http://treatment.plazi.org/id/03AAD15A7F6C0D63FF6CF920FE07F7D5" lastPageId="6" pageId="2">
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<subSubSection id="C31933C77F6C0D67FF6CF920FDE6F0C5" box="[151,553,1665,1692]" pageId="2" type="nomenclature">
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<paragraph id="8BBC604C7F6C0D67FF6CF920FDE6F0C5" blockId="2.[151,553,1665,1692]" box="[151,553,1665,1692]" pageId="2">
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<heading id="D0F4D7207F6C0D67FF6CF920FDE6F0C5" bold="true" box="[151,553,1665,1692]" fontSize="11" level="1" pageId="2" reason="1">
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<emphasis id="B977BC5E7F6C0D67FF6CF920FDE6F0C5" bold="true" box="[151,553,1665,1692]" pageId="2">
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<taxonomicName id="4C031BCF7F6C0D67FF6CF920FE0BF0C2" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[151,452,1665,1692]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
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<emphasis id="B977BC5E7F6C0D67FF6CF920FE0BF0C2" bold="true" box="[151,452,1665,1692]" italics="true" pageId="2">Opisthotropis haihaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24401257F6C0D67FE30F920FDE6F0C5" box="[459,553,1665,1691]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31933C77F6C0D67FF6CF964FC9CF083" box="[151,851,1733,1758]" pageId="2" type="reference_group">
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<paragraph id="8BBC604C7F6C0D67FF6CF964FC9CF083" blockId="2.[151,851,1733,1758]" box="[151,851,1733,1758]" pageId="2">
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<treatmentCitationGroup id="AB1347627F6C0D67FF6CF964FC9CF083" box="[151,851,1733,1758]" pageId="2">
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<taxonomicName id="4C031BCF7F6C0D67FF6CF964FC80F083" authority="Nguyen, Nguyen, Pham & Ziegler 2018: 74" authorityName="Nguyen, Nguyen, Pham & Ziegler" authorityPageNumber="74" authorityYear="2018" box="[151,847,1733,1758]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="maculosa">
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<emphasis id="B977BC5E7F6C0D67FF6CF964FE46F083" box="[151,393,1733,1757]" italics="true" pageId="2">Opisthotropis maculosa</emphasis>
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<treatmentCitation id="0AA2465D7F6C0D67FE74F967FC80F083" author="Nguyen, T. V. & Nguyen, T. Q. & Pham, C. T. & Ziegler, T." box="[399,847,1733,1758]" page="74" pageId="2" year="2018">
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<bibRefCitation id="EF921DBD7F6C0D67FE74F967FCE5F083" author="Nguyen, T. V. & Nguyen, T. Q. & Pham, C. T. & Ziegler, T." box="[399,810,1733,1758]" pageId="2" pagination="74 - 78" refId="ref5842" refString="Nguyen, T. V., Nguyen, T. Q., Pham, C. T. & Ziegler, T. (2018) First country record of Opisthotropis maculosa Stuart et Chuaynkern, 2007 from Vietnam. Russian Journal of Herpetology, 25, 74 - 78." type="journal article" year="2018">Nguyen, Nguyen, Pham & Ziegler 2018</bibRefCitation>
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: 74
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31933C77F6C0D67FF6CF8ADFE5CF1CC" pageId="2" type="materials_examined">
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<paragraph id="8BBC604C7F6C0D67FF6CF8ADFE5CF1CC" blockId="2.[151,1437,1803,2010]" pageId="2">
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<materialsCitation id="3B6B6A117F6C0D67FF6CF8ADFE5CF1CC" ID-GBIF-Occurrence="2248132647" collectingDate="2016-05-09" collectionCode="IEBR" collectorName="Field No. QN & Cuong The Pham & Tan Van Nguyen" country="Vietnam" county="Hai Ha District" elevation="950" location="Tai Chi Village" municipality="Quang Son Commune" pageId="2" specimenCount="1" stateProvince="Quang Ninh" typeStatus="holotype">
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<emphasis id="B977BC5E7F6C0D67FF6CF8ADFEC4F178" bold="true" box="[151,267,1804,1830]" pageId="2">
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<typeStatus id="54B8DEEE7F6C0D67FF6CF8ADFEC9F178" box="[151,262,1804,1830]" pageId="2" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED12F8897F6C0D67FEE9F8ADFE9DF178" box="[274,338,1804,1830]" pageId="2">IEBR</collectionCode>
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A.2016.34 [
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<collectorName id="26F6059A7F6C0D67FE21F8ADFDBAF178" box="[474,629,1804,1830]" pageId="2">Field No. QN</collectorName>
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2016.91], adult female, from the forest near
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<location id="8EDC36977F6C0D67FB92F8ADFADAF178" LSID="urn:lsid:plazi:treatment:03AAD15A7F6C0D63FF6CF920FE07F7D5:8EDC36977F6C0D67FB92F8ADFADAF178" box="[1129,1301,1804,1830]" country="Vietnam" county="Hai Ha District" municipality="Quang Son Commune" name="Tai Chi Village" pageId="2" stateProvince="Quang Ninh">Tai Chi Village</location>
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,
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<collectingMunicipality id="6BD8FA367F6C0D67FAE4F8ADFEC0F114" pageId="2">Quang Son Commune</collectingMunicipality>
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,
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<collectingCounty id="62DD18C07F6C0D67FEE2F891FE09F114" box="[281,454,1840,1866]" pageId="2">Hai Ha District</collectingCounty>
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,
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<collectingRegion id="49C7AEAE7F6C0D67FE2BF891FD0AF114" box="[464,709,1840,1866]" country="Vietnam" name="Quang Ninh" pageId="2">Quang Ninh Province</collectingRegion>
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,
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<elevation id="002E877F7F6C0D67FD2BF88EFC8FF114" box="[720,832,1839,1866]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" unit="m" value="950.0">
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<quantity id="4CFBCDA97F6C0D67FD2BF88EFCD9F114" box="[720,790,1839,1866]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" unit="m" value="950.0">950 m</quantity>
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asl
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</elevation>
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.,
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<collectingCountry id="F31420DC7F6C0D67FCB4F891FC61F114" box="[847,942,1840,1866]" name="Vietnam" pageId="2">Vietnam</collectingCountry>
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[exact locality and coordinates not provided owing to threat from collection for the pet trade (below)], collected by
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<collectorName id="26F6059A7F6C0D67FC4BF8F5FBBBF130" box="[944,1140,1876,1902]" pageId="2">Cuong The Pham</collectorName>
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and
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<collectorName id="26F6059A7F6C0D67FB52F8F5FAAAF130" box="[1193,1381,1876,1902]" pageId="2">Tan Van Nguyen</collectorName>
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on
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<collectingDate id="EFF9BF647F6C0D67FA75F8F5FEC5F1CF" pageId="2" value="2016-05-09">
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<date id="FFBD468C7F6C0D67FA75F8F5FEC5F1CF" pageId="2" value="2016-05-09">9 May 2016</date>
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</collectingDate>
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||
(
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<figureCitation id="13387CC97F6C0D67FEE2F8D9FE4AF1CC" box="[281,389,1912,1938]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1525,1550]" captionStartId-1="5.[151,250,2009,2034]" captionTargetBox-0="[151,1436,549,1513]" captionTargetBox-1="[151,1436,1089,1997]" captionTargetId-0="figure@4.[151,1436,549,1513]" captionTargetId-1="figure@5.[151,1436,1089,1997]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Holotype of Opisthotropis haihaensis sp. nov. (IEBR A.2016.34) in preserved state." captionText-1="FIGURE 3. Head views of the holotype of Opisthotropis haihaensis sp. nov. (IEBR A.2016.34) in preserved state." httpUri-0="https://zenodo.org/record/3239937/files/figure.png" httpUri-1="https://zenodo.org/record/3239939/files/figure.png" pageId="2">Figs. 2–3</figureCitation>
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||
).
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||
</materialsCitation>
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31933C77F6C0D66FF3CF83AFD78F71D" lastPageId="3" pageId="2" type="diagnosis">
|
||
<paragraph id="8BBC604C7F6C0D66FF3CF83AFD78F71D" blockId="2.[151,1437,1803,2010]" lastBlockId="3.[151,1437,152,323]" lastPageId="3" pageId="2">
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||
<emphasis id="B977BC5E7F6C0D67FF3CF83AFE8DF1E8" bold="true" box="[199,322,1947,1974]" pageId="2">Diagnosis.</emphasis>
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A species of the genus
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||
<taxonomicName id="4C031BCF7F6C0D67FDB2F83DFD2DF1E8" box="[585,738,1948,1974]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="genus">
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||
<emphasis id="B977BC5E7F6C0D67FDB2F83DFD2DF1E8" box="[585,738,1948,1974]" italics="true" pageId="2">Opisthotropis</emphasis>
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||
</taxonomicName>
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||
characterized by having the following combination of characters: (1) internasal not in contact with loreal; (2) prefrontal not touching supraocular; (3) frontal touching preocular; (4) one preocular; (5) one postocular; (6) one anterior temporal; (7) one posterior temporal; (8) supralabials eight, fourth and fifth in contact with eye; (9) 24 maxillary teeth; (10) anterior pair of chin shields longer than posterior pair; (11) ventrals 169 (+ 2 preventrals); (12) subcaudals 79; (13) 15 dorsal scale rows at neck, at midbody and before vent; (14) body and tail scales smooth; (15) chin shields yellow with brownish black mottling; (16) body and tail dorsum dark with each a light spot per scale.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF7C30C47F6D0D66FF6CFB3AFDC3F2AA" httpUri="https://zenodo.org/record/3239935/files/figure.png" pageId="3" startId="3.[151,250,1179,1204]" targetBox="[188,1399,395,1135]" targetPageId="3">
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<paragraph id="8BBC604C7F6D0D66FF6CFB3AFDC3F2AA" blockId="3.[151,1437,1179,1268]" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF6CFB3AFEDBF2EA" bold="true" box="[151,276,1179,1204]" pageId="3">FIGURE 1.</emphasis>
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||
Fifty percent majority-rule consensus phylogram based on the combined Bayesian analysis. Numbers above and below branches are MP/ML bootstrap values and combined/partitioned Bayesian posterior probabilities (>50%), respectively. Asterisks (*) represent 100% values.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31933C77F6D0D61FF3CFABCFE67F111" lastPageId="4" pageId="3" type="description">
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<paragraph id="8BBC604C7F6D0D66FF3CFABCFB82F3DE" blockId="3.[151,1437,1309,2021]" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF3CFABCFDDFF366" bold="true" box="[199,528,1309,1336]" pageId="3">
|
||
Description of the
|
||
<typeStatus id="54B8DEEE7F6D0D66FE52FABFFDDFF366" box="[425,528,1310,1336]" pageId="3" type="holotype">holotype</typeStatus>
|
||
</emphasis>
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||
. Body slender, round to oval in cross section; tail thin, pointed; head small, indistinct from neck, depressed, dorsally covered with large shields; nostrils in dorsal position and oblique, narrow, piercing the upper middle of nasal, which is divided beneath; eye small, pupil round.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6D0D66FF3CFA2AFBBEF3FA" blockId="3.[151,1437,1309,2021]" box="[199,1137,1417,1445]" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF3CFA2AFF3BF3FB" box="[199,244,1419,1445]" italics="true" pageId="3">Size</emphasis>
|
||
. HL:
|
||
<quantity id="4CFBCDA97F6D0D66FECDFA2BFE52F3FB" box="[310,413,1418,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="3" unit="mm" value="13.7">13.7 mm</quantity>
|
||
; SVL:
|
||
<quantity id="4CFBCDA97F6D0D66FE17FA2BFD83F3FB" box="[492,588,1418,1445]" metricMagnitude="-1" metricUnit="m" metricValue="3.96" pageId="3" unit="mm" value="396.0">396 mm</quantity>
|
||
; TaL:
|
||
<quantity id="4CFBCDA97F6D0D66FD68FA2BFD3DF3FB" box="[659,754,1418,1445]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="3" unit="mm" value="113.0">113 mm</quantity>
|
||
; TL:
|
||
<quantity id="4CFBCDA97F6D0D66FCD6FA28FC42F3FB" box="[813,909,1417,1445]" metricMagnitude="-1" metricUnit="m" metricValue="5.09" pageId="3" unit="mm" value="509.0">509 mm</quantity>
|
||
; ratio TaL/TL: 0.22.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6D0D66FF3CFA0EFECDF3B2" blockId="3.[151,1437,1309,2021]" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF3CFA0EFEFEF396" box="[199,305,1455,1480]" italics="true" pageId="3">Dentition</emphasis>
|
||
. Maxillary teeth: right upper maxilla with 24 subequal teeth or sockets, teeth small, curved, without diastema.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6D0D66FF3CFA57FA5FF04E" blockId="3.[151,1437,1309,2021]" box="[199,1424,1525,1553]" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF3CFA57FEA0F04E" box="[199,367,1526,1552]" italics="true" pageId="3">Body scalation</emphasis>
|
||
. Dorsal scale rows 15–15–15, entirely smooth. Outer dorsal scales enlarged. All scales glossy.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6D0D66FF3CF9BBFC38F06A" blockId="3.[151,1437,1309,2021]" box="[199,1015,1562,1589]" pageId="3">169 ventrals (+ 2 preventrals); 79 subcaudals, all paired; cloacal divided.</paragraph>
|
||
<paragraph id="8BBC604C7F6D0D61FF3CF99FFB38F689" blockId="3.[151,1437,1309,2021]" lastBlockId="4.[151,1437,152,539]" lastPageId="4" pageId="3">
|
||
<emphasis id="B977BC5E7F6D0D66FF3CF99FFEBDF006" box="[199,370,1598,1624]" italics="true" pageId="3">Head scalation</emphasis>
|
||
. Rostral heptagonal, wider than high, visible from above; nasals large, much longer than high, weakly divided below nostril by a distinct furrow; nasals surrounded by the first two supralabials, rostral, internasal, prefrontal and loreal; internasals two, strongly curved, longer than wide, in contact with rostral anteriorly, nasal, and prefrontal; prefrontal single, heptagonal, anteriorly pointed, about twice as wide as long, in contact with internasals, nasals, loreals, preoculars, and frontal; frontal pentagonal, 1.26 times wider than long, tapering posteriorly, equal to its distance from tip of snout; parietals longer than wide, in contact approximately equal to the length of frontal, 1.88 times longer than frontal; 1 / 1 supraocular, distinctly wider than high, not in contact with prefrontal; 1 / 1 loreal, pentagonal, wider than high, neither in contact with internasal nor eye, surrounded by second and third supralabials, nasal, prefrontal and preocular; 1 / 1 preocular, large, hexagonal, higher than wide, reaching frontal, in broad contact with prefrontal; subocular absent; 1 / 1 postocular, curved, higher than wide; 8 / 8 supralabials, anterior ones higher than wide, fourth and fifth entering orbit, seventh largest; 1+1 / 1+1 temporals, anterior one very long and narrow, in broad contact with SL 5–7 and parietals, posterior one more strongly developed; infralabials 8 / 8, first pair in contact behind small mental, 1–4 /
|
||
<quantity id="4CFBCDA97F6A0D61FDDBFF39FDA8F6ED" box="[544,615,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="4" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
|
||
contact with anterior chin shields, IL 5 largest, apex directed posteriorly; posterior chin shields shorter than anterior ones, separated from each other by a pair of small scales.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6A0D61FF3CFF40FDAEF445" blockId="4.[151,1437,152,539]" pageId="4">
|
||
<emphasis id="B977BC5E7F6A0D61FF3CFF40FDC9F6A5" box="[199,518,225,251]" italics="true" pageId="4">Coloration (in preservative)</emphasis>
|
||
. Ground colour of upper head and body surface dark brown, that of venter yellowish-beige. Dorsal scales with light blotch in the centre each. From behind the neck, the dorsal blotches become more elongated. The blotches become wider towards the body sides; they are largest at the outermost dorsal scale row, where the light blotches stretch towards the posterior scale end. Dorsal tail scales likewise with light central blotches. Dorsal head surface in part with indistinct light mottling that becomes more obvious on temporals. Anterior supralabials with light mottling, supralabials 4–7 with distinct light blotch each. Infralabials, chin shields and smaller throat scales anterior to ventrals brown with light mottling / blotches per scale. The belly with few, scattered dark flecks. Outermost edges of light ventrals brown. Ground colour of subcaudals brown with transversally enlarged light blotches at each scale end.
|
||
</paragraph>
|
||
<caption id="DF7C30C47F6A0D61FF6CFA54FBA2F053" box="[151,1133,1525,1550]" httpUri="https://zenodo.org/record/3239937/files/figure.png" pageId="4" startId="4.[151,250,1525,1550]" targetBox="[151,1436,549,1513]" targetPageId="4">
|
||
<paragraph id="8BBC604C7F6A0D61FF6CFA54FBA2F053" blockId="4.[151,1133,1525,1550]" box="[151,1133,1525,1550]" pageId="4">
|
||
<emphasis id="B977BC5E7F6A0D61FF6CFA54FEDCF050" bold="true" box="[151,275,1525,1550]" pageId="4">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C031BCF7F6A0D61FE62FA57FD59F050" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[409,662,1526,1550]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6A0D61FE62FA57FD59F050" box="[409,662,1526,1550]" italics="true" pageId="4">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6A0D61FD66FA57FD22F050" box="[669,749,1526,1550]" pageId="4" rank="species">
|
||
<emphasis id="B977BC5E7F6A0D61FD66FA57FD22F050" bold="true" box="[669,749,1526,1550]" pageId="4">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(IEBR A.2016.34) in preserved state.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC604C7F6A0D61FF3CF999FAB4F0E0" blockId="4.[151,1437,1592,2015]" pageId="4">
|
||
<emphasis id="B977BC5E7F6A0D61FF3CF999FE6FF00C" box="[199,416,1592,1618]" italics="true" pageId="4">Coloration (in life)</emphasis>
|
||
. Eye black; scales on dorsal surface of head glossy black with scattered yellow flecking; chin shields yellow with brownish black mottling; body and tail glossy black with iridescence above, with single yellow spot on each scale, yellow spots becoming larger on sides of body; ventrals yellow with brownish black lateral margins and scattered brown flecks; subcaudals yellow with brownish black anterior and lateral margins.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6A0D61FF3CF966FE67F111" blockId="4.[151,1437,1592,2015]" pageId="4">
|
||
<emphasis id="B977BC5E7F6A0D61FF3CF966FEA2F0BC" bold="true" box="[199,365,1735,1762]" pageId="4">Comparisons.</emphasis>
|
||
Only six other species of
|
||
<taxonomicName id="4C031BCF7F6A0D61FD62F969FCFDF0BC" box="[665,818,1736,1762]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="B977BC5E7F6A0D61FD62F969FCFDF0BC" box="[665,818,1736,1762]" italics="true" pageId="4">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
and the related genus
|
||
<taxonomicName id="4C031BCF7F6A0D61FBC8F969FB4DF0BC" authorityName="Thomson" authorityYear="1913" box="[1075,1154,1736,1762]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="B977BC5E7F6A0D61FBC8F969FB4DF0BC" box="[1075,1154,1736,1762]" italics="true" pageId="4">Hebius</emphasis>
|
||
</taxonomicName>
|
||
have 15 midbody dorsal scale rows (see review in
|
||
<bibRefCitation id="EF921DBD7F6A0D61FE4EF94DFD47F158" author="Ziegler, T. & David, P. & Ziegler, T. N. & Pham, C. T. & Nguyen, T. Q. & Le, M. D." box="[437,648,1772,1798]" pageId="4" pagination="476 - 496" refId="ref6889" refString="Ziegler, T., David, P., Ziegler, T. N., Pham, C. T., Nguyen, T. Q. & Le, M. D. (2018) Morphological and molecular review of Jacob's Mountain Stream Keelback Opisthotropis jacobi Angel & Bourret, 1933 (Squamata: Natricidae) with description of a sibling species from northern Vietnam. Zootaxa, 4374 (4), 476 - 496. https: // doi. org / 10.11646 / zootaxa. 4374.4.2" type="journal article" year="2018">
|
||
Ziegler
|
||
<emphasis id="B977BC5E7F6A0D61FDF6F94CFD8AF158" box="[525,581,1772,1798]" italics="true" pageId="4">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C031BCF7F6A0D61FD60F94CFCE7F158" authorityName="Nguyen, Nguyen, Pham & Ziegler" authorityYear="2018" box="[667,808,1772,1799]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6A0D61FD60F94CFCE7F158" box="[667,808,1772,1799]" italics="true" pageId="4">O. maculosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C031BCF7F6A0D61FCCEF94CFBACF159" authority="Angel & Bourret" authorityName="Angel & Bourret" authorityYear="1933" box="[821,1123,1772,1799]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="jacobi">
|
||
<emphasis id="B977BC5E7F6A0D61FCCEF94CFC52F158" box="[821,925,1772,1799]" italics="true" pageId="4">O. jacobi</emphasis>
|
||
Angel & Bourret
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C031BCF7F6A0D61FB95F94CFB14F158" authorityName="Ziegler, David, Ziegler, Pham, Nguyen & Le" authorityYear="2018" box="[1134,1243,1773,1799]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="voquyi">
|
||
<emphasis id="B977BC5E7F6A0D61FB95F94CFB14F158" box="[1134,1243,1773,1799]" italics="true" pageId="4">O. voquyi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C031BCF7F6A0D61FB13F94CFE44F175" authority="Zhao, Jiang & Huang" authorityName="Zhao, Jiang & Huang" authorityYear="1978" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6A0D61FB13F94CFA53F158" box="[1256,1436,1773,1799]" italics="true" pageId="4">O. guangxiensis</emphasis>
|
||
Zhao, Jiang & Huang
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C031BCF7F6A0D61FE6CF8B0FD3BF175" authority="(Okada & Takara)" authorityName="Okada & Takara" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" box="[407,756,1808,1835]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="kikuzatoi">
|
||
<emphasis id="B977BC5E7F6A0D61FE6CF8B0FDD0F174" box="[407,543,1808,1835]" italics="true" pageId="4">O. kikuzatoi</emphasis>
|
||
(Okada & Takara)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C031BCF7F6A0D61FCD4F8B0FBFCF175" authority="Bourret" authorityName="Bourret" authorityYear="1934" box="[815,1075,1809,1835]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="B977BC5E7F6A0D61FCD4F8B0FC19F174" box="[815,982,1809,1834]" italics="true" pageId="4">H. annamensis</emphasis>
|
||
Bourret
|
||
</taxonomicName>
|
||
(which may have 15 or 17 midbody dorsal scale rows).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31933C77F6A0D60FF3CF8F9FB86F7F1" lastPageId="5" pageId="4" type="reference_group">
|
||
<paragraph id="8BBC604C7F6A0D60FF3CF8F9FEDCF71D" blockId="4.[151,1437,1592,2015]" lastBlockId="5.[151,1437,153,1080]" lastPageId="5" pageId="4">
|
||
<taxonomicName id="4C031BCF7F6A0D61FF3CF8F9FE10F12C" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,479,1880,1906]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6A0D61FF3CF8F9FE10F12C" box="[199,479,1880,1906]" italics="true" pageId="4">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6A0D61FE1CF8F9FD8FF12C" box="[487,576,1880,1906]" pageId="4" rank="species">
|
||
<emphasis id="B977BC5E7F6A0D61FE1CF8F9FD8FF12C" bold="true" box="[487,576,1880,1906]" pageId="4">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6A0D61FD2EF8F8FCACF12C" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[725,867,1880,1907]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" sensu="stricto" species="maculosa">
|
||
<emphasis id="B977BC5E7F6A0D61FD2EF8F8FD26F12D" box="[725,745,1881,1907]" italics="true" pageId="4">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6A0D61FD03F8F9FCACF12C" box="[760,867,1880,1906]" italics="true" pageId="4">maculosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6A0D61FC91F8F8FC37F12C" box="[874,1016,1881,1906]" pageId="4" sensu="stricto">
|
||
<emphasis id="B977BC5E7F6A0D61FC91F8F8FC37F12C" box="[874,1016,1881,1906]" italics="true" pageId="4">sensu stricto</emphasis>
|
||
</taxonomicNameLabel>
|
||
(adult male
|
||
<typeStatus id="54B8DEEE7F6A0D61FB7EF8F9FB28F12C" box="[1157,1255,1880,1906]" pageId="4" type="holotype">holotype</typeStatus>
|
||
from northeastern
|
||
<collectingCountry id="F31420DC7F6A0D61FF39F8DDFEE7F1C8" box="[194,296,1916,1942]" name="Thailand" pageId="4">Thailand</collectingCountry>
|
||
, see
|
||
<bibRefCitation id="EF921DBD7F6A0D61FEA5F8DDFD5EF1C8" author="Stuart, B. L. & Chuaynkern, Y." box="[350,657,1916,1942]" pageId="4" pagination="35 - 40" refId="ref6136" refString="Stuart, B. L. & Chuaynkern, Y. (2007) A new Opisthotropis (Serpentes: Colubridae: Natricinae) from Northeastern Thailand. Current Herpetology, 26, 35 - 40. https: // doi. org / 10.3105 / 1345 - 5834 (2007) 26 [35: ANOSCN] 2.0. CO; 2" type="journal article" year="2007">Stuart & Chuaynkern 2007</bibRefCitation>
|
||
), with which the species was confused in the previous study (
|
||
<bibRefCitation id="EF921DBD7F6A0D61FAB9F8DDFEC0F1E4" author="Nguyen, T. V. & Nguyen, T. Q. & Pham, C. T. & Ziegler, T." pageId="4" pagination="74 - 78" refId="ref5842" refString="Nguyen, T. V., Nguyen, T. Q., Pham, C. T. & Ziegler, T. (2018) First country record of Opisthotropis maculosa Stuart et Chuaynkern, 2007 from Vietnam. Russian Journal of Herpetology, 25, 74 - 78." type="journal article" year="2018">Nguyen et al. 2018</bibRefCitation>
|
||
), by having prefrontal not touching supraocular (
|
||
<emphasis id="B977BC5E7F6A0D61FCD4F800FCB8F1E4" box="[815,887,1953,1978]" italics="true" pageId="4">versus</emphasis>
|
||
prefrontal touching supraocular in
|
||
<taxonomicName id="4C031BCF7F6A0D61FB05F800FA44F1E4" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[1278,1419,1952,1979]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6A0D61FB05F800FADDF1E5" box="[1278,1298,1953,1979]" italics="true" pageId="4">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6A0D61FADBF801FA44F1E4" box="[1312,1419,1952,1978]" italics="true" pageId="4">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); frontal touching preocular (
|
||
<emphasis id="B977BC5E7F6A0D61FE30F864FDDCF180" box="[459,531,1989,2014]" italics="true" pageId="4">versus</emphasis>
|
||
frontal not touching preocular in
|
||
<taxonomicName id="4C031BCF7F6A0D61FC77F864FBD6F180" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[908,1049,1988,2015]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6A0D61FC77F864FC6FF181" box="[908,928,1989,2015]" italics="true" pageId="4">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6A0D61FC55F865FBD6F180" box="[942,1049,1988,2014]" italics="true" pageId="4">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); one postocular (
|
||
<emphasis id="B977BC5E7F6A0D61FB1EF864FAE2F180" box="[1253,1325,1989,2014]" italics="true" pageId="4">versus</emphasis>
|
||
two post- oculars in
|
||
<taxonomicName id="4C031BCF7F6B0D60FEF7FF38FE55F6ED" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[268,410,153,179]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FEF7FF38FEEFF6ED" box="[268,288,153,179]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FED4FF38FE55F6ED" box="[303,410,153,179]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); fourth and fifth supralabials in contact with eye (
|
||
<emphasis id="B977BC5E7F6B0D60FC23FF3BFBEFF6ED" box="[984,1056,154,179]" italics="true" pageId="5">versus</emphasis>
|
||
fourth supralabial in contact with eye in
|
||
<taxonomicName id="4C031BCF7F6B0D60FF1CFF1CFEB9F689" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[231,374,189,215]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FF1CFF1CFF34F689" box="[231,251,189,215]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FEF0FF1CFEB9F689" box="[267,374,189,215]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); anterior pair of chin shields longer than posterior pair (
|
||
<emphasis id="B977BC5E7F6B0D60FBF5FF1FFB99F689" box="[1038,1110,190,215]" italics="true" pageId="5">versus</emphasis>
|
||
vice versa in
|
||
<taxonomicName id="4C031BCF7F6B0D60FB00FF1CFA44F689" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[1275,1419,189,215]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FB00FF1CFAC0F689" box="[1275,1295,189,215]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FADBFF1CFA44F689" box="[1312,1419,189,215]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); lower number of ventrals (169 + 2 preventrals versus 182 + 2 preventrals in
|
||
<taxonomicName id="4C031BCF7F6B0D60FC02FF40FB48F6A5" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[1017,1159,225,251]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FC02FF40FBC2F6A5" box="[1017,1037,225,251]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FBE7FF40FB48F6A5" box="[1052,1159,225,251]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); higher number of subcaudals (79
|
||
<emphasis id="B977BC5E7F6B0D60FEE4FEA7FEA8F741" box="[287,359,262,287]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FE95FEA4FE68F741" box="[366,423,261,287]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" pageId="5" unit="in" value="67.0">67 in</quantity>
|
||
<taxonomicName id="4C031BCF7F6B0D60FE54FEA4FDF3F741" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[431,572,261,287]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FE54FEA4FE0CF741" box="[431,451,261,287]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FE2AFEA4FDF3F741" box="[465,572,261,287]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
); and chin shields yellow with brownish black mottling (
|
||
<emphasis id="B977BC5E7F6B0D60FB3AFEA7FAC6F741" box="[1217,1289,262,287]" italics="true" pageId="5">versus</emphasis>
|
||
yellow in
|
||
<taxonomicName id="4C031BCF7F6B0D60FA7AFEA4FECDF71D" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B977BC5E7F6B0D60FA7AFEA4FA5AF741" box="[1409,1429,261,287]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FF6CFE88FECDF71D" box="[151,258,297,323]" italics="true" pageId="5">maculosa</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFEECFB86F7F1" blockId="5.[151,1437,153,1080]" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FF3CFEECFE2FF739" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,480,333,359]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFEECFE2FF739" box="[199,480,333,359]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FE13FEECFD8DF739" box="[488,578,333,359]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FE13FEECFD8DF739" bold="true" box="[488,578,333,359]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6B0D60FD23FEECFC8EF739" authorityName="Angel & Bourret" authorityYear="1933" box="[728,833,333,359]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="jacobi">
|
||
<emphasis id="B977BC5E7F6B0D60FD23FEECFD23F739" box="[728,748,333,359]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FD00FEECFC8EF739" box="[763,833,333,359]" italics="true" pageId="5">jacobi</emphasis>
|
||
</taxonomicName>
|
||
by having 24 maxillary teeth (
|
||
<emphasis id="B977BC5E7F6B0D60FB64FEEFFB28F739" box="[1183,1255,334,359]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FB14FEECFA9CF739" box="[1263,1363,333,359]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.842" metricValueMin="4.826" pageId="5" unit="in" value="21.0" valueMax="23.0" valueMin="19.0">19–23 in</quantity>
|
||
<taxonomicName id="4C031BCF7F6B0D60FAA1FEECFF08F7D5" authorityName="Angel & Bourret" authorityYear="1933" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="jacobi">
|
||
<emphasis id="B977BC5E7F6B0D60FAA1FEECFF08F7D5" italics="true" pageId="5">O. jacobi</emphasis>
|
||
</taxonomicName>
|
||
); smooth dorsal scales on tail, without keels (
|
||
<emphasis id="B977BC5E7F6B0D60FD3CFED3FCC0F7D5" box="[711,783,370,395]" italics="true" pageId="5">versus</emphasis>
|
||
few faint keels usually present in
|
||
<taxonomicName id="4C031BCF7F6B0D60FB70FED0FB3CF7D5" authorityName="Angel & Bourret" authorityYear="1933" box="[1163,1267,369,395]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="jacobi">
|
||
<emphasis id="B977BC5E7F6B0D60FB70FED0FB3CF7D5" box="[1163,1267,369,395]" italics="true" pageId="5">O. jacobi</emphasis>
|
||
</taxonomicName>
|
||
); body and tail dorsum dark with a light spot per scale (
|
||
<emphasis id="B977BC5E7F6B0D60FDA1FE37FD6DF7F1" box="[602,674,406,431]" italics="true" pageId="5">versus</emphasis>
|
||
dorsum uniformly dark in
|
||
<taxonomicName id="4C031BCF7F6B0D60FC2BFE34FBF7F7F1" authorityName="Angel & Bourret" authorityYear="1933" box="[976,1080,405,431]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="jacobi">
|
||
<emphasis id="B977BC5E7F6B0D60FC2BFE34FBF7F7F1" box="[976,1080,405,431]" italics="true" pageId="5">O. jacobi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31933C77F6B0D60FF3CFE18FBE8F5B1" pageId="5" type="description">
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFE18FD07F445" blockId="5.[151,1437,153,1080]" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FF3CFE18FE2FF78D" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,480,441,467]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFE18FE2FF78D" box="[199,480,441,467]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FE13FE18FD8CF78D" box="[488,579,441,467]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FE13FE18FD8CF78D" bold="true" box="[488,579,441,467]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6B0D60FD22FE18FC87F78D" authorityName="Ziegler, David, Ziegler, Pham, Nguyen & Le" authorityYear="2018" box="[729,840,441,467]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="voquyi">
|
||
<emphasis id="B977BC5E7F6B0D60FD22FE18FC87F78D" box="[729,840,441,467]" italics="true" pageId="5">O. voquyi</emphasis>
|
||
</taxonomicName>
|
||
by having one postocular (
|
||
<emphasis id="B977BC5E7F6B0D60FB85FE1BFB09F78D" box="[1150,1222,442,467]" italics="true" pageId="5">versus</emphasis>
|
||
two in
|
||
<taxonomicName id="4C031BCF7F6B0D60FAE6FE18FA44F78D" authorityName="Ziegler, David, Ziegler, Pham, Nguyen & Le" authorityYear="2018" box="[1309,1419,441,467]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="voquyi">
|
||
<emphasis id="B977BC5E7F6B0D60FAE6FE18FA44F78D" box="[1309,1419,441,467]" italics="true" pageId="5">O. voquyi</emphasis>
|
||
</taxonomicName>
|
||
); body and tail dorsum dark with each a light spot per scale (
|
||
<emphasis id="B977BC5E7F6B0D60FCD2FE7FFCBEF7A9" box="[809,881,478,503]" italics="true" pageId="5">versus</emphasis>
|
||
dorsal pattern dark, with dorsal scales posteriorly more or less edged with pale brown in
|
||
<taxonomicName id="4C031BCF7F6B0D60FDB0FDA0FD77F445" authorityName="Ziegler, David, Ziegler, Pham, Nguyen & Le" authorityYear="2018" box="[587,696,513,539]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="voquyi">
|
||
<emphasis id="B977BC5E7F6B0D60FDB0FDA0FD77F445" box="[587,696,513,539]" italics="true" pageId="5">O. voquyi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFD84FD2BF491" blockId="5.[151,1437,153,1080]" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FF3CFD84FE10F461" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,479,549,575]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFD84FE10F461" box="[199,479,549,575]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FE1EFD84FDF0F461" box="[485,575,549,575]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FE1EFD84FDF0F461" bold="true" box="[485,575,549,575]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6B0D60FD29FD84FC49F461" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[722,902,549,575]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FD29FD84FC49F461" box="[722,902,549,575]" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
by having eight supralabials (
|
||
<emphasis id="B977BC5E7F6B0D60FB2CFD87FAD0F461" box="[1239,1311,550,575]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FADDFD84FAB5F461" box="[1318,1402,549,575]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="5" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
<taxonomicName id="4C031BCF7F6B0D60FA7AFD84FEE6F43D" authorityName="Zhao, Jiang & Huang" authorityYear="1978" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FA7AFD84FEE6F43D" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
); one postocular (
|
||
<emphasis id="B977BC5E7F6B0D60FE01FDEBFD8DF43D" box="[506,578,586,611]" italics="true" pageId="5">versus</emphasis>
|
||
two in
|
||
<taxonomicName id="4C031BCF7F6B0D60FD61FDE8FC9FF43D" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[666,848,585,611]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FD61FDE8FC9FF43D" box="[666,848,585,611]" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
); one posterior temporal in the new species (
|
||
<emphasis id="B977BC5E7F6B0D60FAAEFDEBFA52F43D" box="[1365,1437,586,611]" italics="true" pageId="5">versus</emphasis>
|
||
two in
|
||
<taxonomicName id="4C031BCF7F6B0D60FF1FFDCCFE57F4D9" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[228,408,621,647]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FF1FFDCCFE57F4D9" box="[228,408,621,647]" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
); 15 dorsal scale rows at neck in the new species (
|
||
<emphasis id="B977BC5E7F6B0D60FC31FDCFFBDDF4D9" box="[970,1042,622,647]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FBE3FDCCFB9EF4D9" box="[1048,1105,621,647]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="5" unit="in" value="17.0">17 in</quantity>
|
||
<taxonomicName id="4C031BCF7F6B0D60FBACFDCCFAC4F4D9" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[1111,1291,621,647]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FBACFDCCFAC4F4D9" box="[1111,1291,621,647]" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
); 79 subcaudals (
|
||
<emphasis id="B977BC5E7F6B0D60FF2EFD33FED2F4F5" box="[213,285,658,683]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FEDFFD31FE48F4F5" box="[292,391,656,683]" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.524" metricValueMin="1.2953999999999999" pageId="5" unit="in" value="55.5" valueMax="60.0" valueMin="51.0">51–60 in</quantity>
|
||
<taxonomicName id="4C031BCF7F6B0D60FE75FD30FD8DF4F5" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[398,578,657,683]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">
|
||
<emphasis id="B977BC5E7F6B0D60FE75FD30FD8DF4F5" box="[398,578,657,683]" italics="true" pageId="5">O. guangxiensis</emphasis>
|
||
</taxonomicName>
|
||
); body and tail dorsum dark with a light spot per scale (
|
||
<emphasis id="B977BC5E7F6B0D60FB41FD33FACDF4F5" box="[1210,1282,658,683]" italics="true" pageId="5">versus</emphasis>
|
||
dorsal colour pattern dark with pale crossbars
|
||
<emphasis id="B977BC5E7F6B0D60FDF9FD17FD1CF491" box="[514,723,693,719]" italics="true" pageId="5">
|
||
in
|
||
<taxonomicName id="4C031BCF7F6B0D60FDE4FD14FD1CF491" authorityName="Zhao, Jiang & Huang" authorityYear="1978" box="[543,723,693,719]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="guangxiensis">O. guangxiensis</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFD78FD13F501" blockId="5.[151,1437,153,1080]" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FF3CFD78FE2EF4AD" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,481,729,755]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFD78FE2EF4AD" box="[199,481,729,755]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FE11FD78FD8AF4AD" box="[490,581,729,755]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FE11FD78FD8AF4AD" bold="true" box="[490,581,729,755]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6B0D60FD25FD78FCA8F4AD" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" box="[734,871,729,755]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="kikuzatoi">
|
||
<emphasis id="B977BC5E7F6B0D60FD25FD78FD3DF4AD" box="[734,754,729,755]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FCF9FD78FCA8F4AD" box="[770,871,729,755]" italics="true" pageId="5">kikuzatoi</emphasis>
|
||
</taxonomicName>
|
||
by having eight supralabials (
|
||
<emphasis id="B977BC5E7F6B0D60FB39FD7BFAC5F4AD" box="[1218,1290,730,755]" italics="true" pageId="5">versus</emphasis>
|
||
six in
|
||
<taxonomicName id="4C031BCF7F6B0D60FAA0FD78FF27F549" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="kikuzatoi">
|
||
<emphasis id="B977BC5E7F6B0D60FAA0FD78FAA0F4AD" box="[1371,1391,729,755]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FA84FD78FF27F549" italics="true" pageId="5">kikuzatoi</emphasis>
|
||
</taxonomicName>
|
||
); one preocular (
|
||
<emphasis id="B977BC5E7F6B0D60FE50FD5FFE3CF549" box="[427,499,766,791]" italics="true" pageId="5">versus</emphasis>
|
||
two in
|
||
<taxonomicName id="4C031BCF7F6B0D60FDB2FD5CFD1EF549" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" box="[585,721,765,791]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="kikuzatoi">
|
||
<emphasis id="B977BC5E7F6B0D60FDB2FD5CFD92F549" box="[585,605,765,791]" italics="true" pageId="5">O</emphasis>
|
||
.
|
||
<emphasis id="B977BC5E7F6B0D60FD97FD5CFD1EF549" box="[620,721,765,791]" italics="true" pageId="5">kikuzatoi</emphasis>
|
||
</taxonomicName>
|
||
); 169 ventrals and 79 subcaudals (
|
||
<emphasis id="B977BC5E7F6B0D60FBA0FD5FFB6CF549" box="[1115,1187,766,791]" italics="true" pageId="5">versus</emphasis>
|
||
180–198 ventrals and 58–73 subcaudals in
|
||
<taxonomicName id="4C031BCF7F6B0D60FE84FC80FDCAF565" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" box="[383,517,801,827]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="kikuzatoi">
|
||
<emphasis id="B977BC5E7F6B0D60FE84FC80FDCAF565" box="[383,517,801,827]" italics="true" pageId="5">O. kikuzatoi</emphasis>
|
||
</taxonomicName>
|
||
); body and tail dorsum dark with a light spot per scale (
|
||
<emphasis id="B977BC5E7F6B0D60FB91FC83FB7DF565" box="[1130,1202,802,827]" italics="true" pageId="5">versus</emphasis>
|
||
dorsal colour pattern dark with dorsolateral orange spots
|
||
<emphasis id="B977BC5E7F6B0D60FDD3FCE7FD03F501" box="[552,716,837,863]" italics="true" pageId="5">
|
||
in
|
||
<taxonomicName id="4C031BCF7F6B0D60FDBFFCE4FD03F501" baseAuthorityName="Okada & Takara" baseAuthorityYear="1958" box="[580,716,837,863]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="kikuzatoi">O. kikuzatoi</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFCC8FBE8F5B1" blockId="5.[151,1437,153,1080]" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FF3CFCC8FE10F5DD" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[199,479,873,899]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFCC8FE10F5DD" box="[199,479,873,899]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FE1EFCC8FDF0F5DD" box="[485,575,873,899]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FE1EFCC8FDF0F5DD" bold="true" box="[485,575,873,899]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C031BCF7F6B0D60FD29FCCBFCB7F5DD" authorityName="Bourret" authorityYear="1934" box="[722,888,874,899]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="B977BC5E7F6B0D60FD29FCCBFCB7F5DD" box="[722,888,874,899]" italics="true" pageId="5">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
by having one preocular (
|
||
<emphasis id="B977BC5E7F6B0D60FB66FCCBFB2AF5DD" box="[1181,1253,874,899]" italics="true" pageId="5">versus</emphasis>
|
||
two in
|
||
<taxonomicName id="4C031BCF7F6B0D60FAC2FCCBFF2CF5F9" authorityName="Bourret" authorityYear="1934" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="B977BC5E7F6B0D60FAC2FCCBFF2CF5F9" italics="true" pageId="5">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
); 15 dorsal scale rows at neck (
|
||
<emphasis id="B977BC5E7F6B0D60FDB9FC2FFD45F5F9" box="[578,650,910,935]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FD6BFC2CFD3DF5F9" box="[656,754,909,935]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="4.826" metricValueMin="4.318" pageId="5" unit="in" value="18.0" valueMax="19.0" valueMin="17.0">17–19 in</quantity>
|
||
<emphasis id="B977BC5E7F6B0D60FD03FC2FFC68F5F9" box="[760,935,909,935]" italics="true" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FD03FC2FFC51F5F9" authorityName="Bourret" authorityYear="1934" box="[760,926,909,935]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">H. annamensis</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
; body scales smooth (
|
||
<emphasis id="B977BC5E7F6B0D60FB64FC2FFB28F5F9" box="[1183,1255,910,935]" italics="true" pageId="5">versus</emphasis>
|
||
keeled throughout in
|
||
<taxonomicName id="4C031BCF7F6B0D60FF1AFC13FE47F595" authorityName="Bourret" authorityYear="1934" box="[225,392,946,971]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="B977BC5E7F6B0D60FF1AFC13FE47F595" box="[225,392,946,971]" italics="true" pageId="5">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
); 79 subcaudals (
|
||
<emphasis id="B977BC5E7F6B0D60FDA9FC13FD55F595" box="[594,666,946,971]" italics="true" pageId="5">versus</emphasis>
|
||
<quantity id="4CFBCDA97F6B0D60FD5AFC10FCEFF595" box="[673,800,945,971]" metricMagnitude="0" metricUnit="m" metricValue="3.3274" metricValueMax="3.7084" metricValueMin="2.9463999999999997" pageId="5" unit="in" value="131.0" valueMax="146.0" valueMin="116.0">116–146 in</quantity>
|
||
<emphasis id="B977BC5E7F6B0D60FCD3FC13FC16F595" box="[808,985,945,971]" italics="true" pageId="5">
|
||
<taxonomicName id="4C031BCF7F6B0D60FCD3FC13FC1FF595" authorityName="Bourret" authorityYear="1934" box="[808,976,945,971]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">H. annamensis</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
; body and tail dorsum dark with a light spot per scale (
|
||
<emphasis id="B977BC5E7F6B0D60FEBBFC77FE47F5B1" box="[320,392,982,1007]" italics="true" pageId="5">versus</emphasis>
|
||
dorsum dark with orange bars and spots in
|
||
<taxonomicName id="4C031BCF7F6B0D60FC8AFC77FBD8F5B1" authorityName="Bourret" authorityYear="1934" box="[881,1047,982,1007]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="B977BC5E7F6B0D60FC8AFC77FBD8F5B1" box="[881,1047,982,1007]" italics="true" pageId="5">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31933C77F6B0D60FF3CFC58FA42F266" pageId="5" type="etymology">
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFC58FAB1F24D" blockId="5.[151,1437,153,1080]" box="[199,1406,1017,1043]" pageId="5">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFC58FE87F24D" bold="true" box="[199,328,1017,1043]" pageId="5">Etymology</emphasis>
|
||
. This species is named after its
|
||
<typeStatus id="54B8DEEE7F6B0D60FD55FC58FD11F24D" box="[686,734,1017,1043]" pageId="5">type</typeStatus>
|
||
locality in Hai Ha District,
|
||
<collectingRegion id="49C7AEAE7F6B0D60FBEDFC58FAC3F24D" box="[1046,1292,1017,1043]" country="Vietnam" name="Quang Ninh" pageId="5">Quang Ninh Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31420DC7F6B0D60FAECFC58FAB5F24D" box="[1303,1402,1017,1043]" name="Vietnam" pageId="5">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F6B0D60FF3CFBBDFA42F266" blockId="5.[151,1437,153,1080]" box="[199,1421,1052,1080]" pageId="5">
|
||
<emphasis id="B977BC5E7F6B0D60FF3CFBBDFDC8F269" bold="true" box="[199,519,1052,1079]" pageId="5">Suggested common names.</emphasis>
|
||
Hai Ha Mountain Stream Keelback (English) and Rắn trán hải hà (Vietnamese).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7C30C47F6B0D60FF6CF878FAE9F1AC" box="[151,1318,2009,2034]" httpUri="https://zenodo.org/record/3239939/files/figure.png" pageId="5" startId="5.[151,250,2009,2034]" targetBox="[151,1436,1089,1997]" targetPageId="5">
|
||
<paragraph id="8BBC604C7F6B0D60FF6CF878FAE9F1AC" blockId="5.[151,1318,2009,2034]" box="[151,1318,2009,2034]" pageId="5">
|
||
<emphasis id="B977BC5E7F6B0D60FF6CF878FEDCF1AC" bold="true" box="[151,275,2009,2034]" pageId="5">FIGURE 3.</emphasis>
|
||
Head views of the holotype of
|
||
<taxonomicName id="4C031BCF7F6B0D60FDA9F87BFC81F1AC" authority="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le, 2019" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[594,846,2010,2034]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="haihaensis" status="sp. nov.">
|
||
<emphasis id="B977BC5E7F6B0D60FDA9F87BFC81F1AC" box="[594,846,2010,2034]" italics="true" pageId="5">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24401257F6B0D60FCAEF87BFC69F1AC" box="[853,934,2010,2034]" pageId="5" rank="species">
|
||
<emphasis id="B977BC5E7F6B0D60FCAEF87BFC69F1AC" bold="true" box="[853,934,2010,2034]" pageId="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(IEBR A.2016.34) in preserved state.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31933C77F680D63FF3CFF39FE07F7D5" pageId="6" type="distribution">
|
||
<paragraph id="8BBC604C7F680D63FF3CFF39FDD0F689" blockId="6.[151,1437,152,395]" pageId="6">
|
||
<emphasis id="B977BC5E7F680D63FF3CFF39FE96F6ED" bold="true" box="[199,345,152,179]" pageId="6">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C031BCF7F680D63FE9CFF38FDB1F6ED" authorityName="Ziegler & Pham & Nguyen & Nguyen & Wang & Wang & Stuart & Le" authorityYear="2019" box="[359,638,153,179]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="6" phylum="Chordata" rank="species" species="haihaensis">
|
||
<emphasis id="B977BC5E7F680D63FE9CFF38FDB1F6ED" box="[359,638,153,179]" italics="true" pageId="6">Opisthotropis haihaensis</emphasis>
|
||
</taxonomicName>
|
||
is currently known only from the
|
||
<typeStatus id="54B8DEEE7F680D63FC0CFF38FBE8F6ED" box="[1015,1063,153,179]" pageId="6">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49C7AEAE7F680D63FB58FF38FA57F6ED" box="[1187,1432,153,179]" country="Vietnam" name="Quang Ninh" pageId="6">Quang Ninh Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31420DC7F680D63FF6CFF1CFF39F689" box="[151,246,189,215]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
|
||
(see
|
||
<bibRefCitation id="EF921DBD7F680D63FECAFF1CFDDBF689" author="Nguyen, T. V. & Nguyen, T. Q. & Pham, C. T. & Ziegler, T." box="[305,532,189,215]" pageId="6" pagination="74 - 78" refId="ref5842" refString="Nguyen, T. V., Nguyen, T. Q., Pham, C. T. & Ziegler, T. (2018) First country record of Opisthotropis maculosa Stuart et Chuaynkern, 2007 from Vietnam. Russian Journal of Herpetology, 25, 74 - 78." type="journal article" year="2018">
|
||
Nguyen
|
||
<emphasis id="B977BC5E7F680D63FE6AFF1FFE0DF689" box="[401,450,189,215]" italics="true" pageId="6">et al</emphasis>
|
||
., 2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBC604C7F680D63FF3CFF40FE07F7D5" blockId="6.[151,1437,152,395]" pageId="6">
|
||
<emphasis id="B977BC5E7F680D63FF3CFF40FEB0F6A5" bold="true" box="[199,383,224,251]" pageId="6">Natural history</emphasis>
|
||
. The adult female
|
||
<typeStatus id="54B8DEEE7F680D63FDB5FF40FD7FF6A5" box="[590,688,225,251]" pageId="6" type="holotype">holotype</typeStatus>
|
||
was found at night in a small rocky stream at 2130h. The surrounding habitat was secondary evergreen forest consisting of small hardwood trees, bamboo, and shrubs (see
|
||
<bibRefCitation id="EF921DBD7F680D63FABFFEA4FED5F71D" author="Nguyen, T. V. & Nguyen, T. Q. & Pham, C. T. & Ziegler, T." pageId="6" pagination="74 - 78" refId="ref5842" refString="Nguyen, T. V., Nguyen, T. Q., Pham, C. T. & Ziegler, T. (2018) First country record of Opisthotropis maculosa Stuart et Chuaynkern, 2007 from Vietnam. Russian Journal of Herpetology, 25, 74 - 78." type="journal article" year="2018">
|
||
Nguyen
|
||
<emphasis id="B977BC5E7F680D63FF6CFE8BFF00F71D" box="[151,207,297,323]" italics="true" pageId="6">et al.</emphasis>
|
||
, 2018
|
||
</bibRefCitation>
|
||
). The air temperature was 24–29
|
||
<superScript id="7C76CD047F680D63FD73FE89FD40F768" attach="right" box="[648,655,296,310]" fontSize="6" pageId="6">°</superScript>
|
||
C and the relative humidity was 65–88%. Dissection revealed that the
|
||
<typeStatus id="54B8DEEE7F680D63FF6CFEECFF36F739" box="[151,249,333,359]" pageId="6" type="holotype">holotype</typeStatus>
|
||
contained three large eggs having maximum length of
|
||
<quantity id="4CFBCDA97F680D63FCA5FEEDFC0BF739" box="[862,964,332,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="6" unit="mm" value="16.5">16.5 mm</quantity>
|
||
, with folded oviducts indicating that some eggs had already been laid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |