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<document id="11AF761C68CBF8AAACA45AEEB90A737C" ID="10.11646/zootaxa.4139.3.2" ID-DOI="10.11646/zootaxa.4139.3.2" ID-GBIF-Dataset="51358935-98b0-4d89-a47f-e4341171e5b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="262765" ID-ZooBank="DCF4220B-4A73-407C-837C-54DEAE29F435" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469091585085" checkinUser="plazi" docAuthor="Last, Peter R., Naylor, Gavin J. P. &amp; Manjaji-Matsumoto, B. Mabel" docDate="2016" docId="03A7CA1D857A1440E7C6FCC7FBFE0179" docLanguage="en" docName="zootaxa.4139.3.2.pdf" docOrigin="Zootaxa 4139 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Himantura Muller &amp; Henle 1837" docType="treatment" docVersion="9" lastPageNumber="361" masterDocId="FF9EB265856A1450E751FF83FFDE0766" masterDocTitle="A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights" masterLastPageNumber="368" masterPageNumber="345" pageNumber="361" updateTime="1698365466156" updateUser="plazi">
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<mods:title id="033BC4BBA011CD27EE70212802FA0F23">A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights</mods:title>
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<mods:namePart id="1E5249D6CA3B6D4C62B59506FA84CAC6">Manjaji-Matsumoto, B. Mabel</mods:namePart>
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<mods:date id="DC4A57D05F3FDF2B2CBFC92FA83E1987">2016</mods:date>
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<treatment id="03A7CA1D857A1440E7C6FCC7FBFE0179" ID-DOI="http://doi.org/10.5281/zenodo.6085235" ID-GBIF-Taxon="123198561" ID-Zenodo-Dep="6085235" LSID="urn:lsid:plazi:treatment:03A7CA1D857A1440E7C6FCC7FBFE0179" httpUri="http://treatment.plazi.org/id/03A7CA1D857A1440E7C6FCC7FBFE0179" lastPageNumber="361" pageId="16" pageNumber="361">
<subSubSection id="C3142880857A1440E7C6FCC7FD530438" box="[151,653,836,862]" pageId="16" pageNumber="361" type="nomenclature">
<paragraph id="8BB17B0B857A1440E7C6FCC7FD530438" blockId="16.[151,653,836,862]" box="[151,653,836,862]" pageId="16" pageNumber="361">
<heading id="D0F9CC67857A1440E7C6FCC7FD530438" bold="true" box="[151,653,836,862]" fontSize="11" level="1" pageId="16" pageNumber="361" reason="1">
<emphasis id="B97AA719857A1440E7C6FCC7FD530438" bold="true" box="[151,653,836,862]" pageId="16" pageNumber="361">
Genus
<taxonomicName id="4C0E0088857A1440E7BFFCC7FD530438" ID-CoL="4XB4" ID-ENA="86362" authority="Muller &amp; Henle, 1837" authorityName="Muller &amp; Henle" authorityYear="1837" box="[238,653,836,862]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="genus">
<emphasis id="B97AA719857A1440E7BFFCC7FEAB0438" bold="true" box="[238,373,836,862]" italics="true" pageId="16" pageNumber="361">Himantura</emphasis>
Müller &amp; Henle, 1837
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</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3142880857A1440E7C6FC08FEB2020B" pageId="16" pageNumber="361" type="reference_group">
<paragraph id="8BB17B0B857A1440E7C6FC08FE4604D9" blockId="16.[151,1436,906,959]" pageId="16" pageNumber="361">
<taxonomicName id="4C0E0088857A1440E7C6FC08FDC304C6" ID-CoL="4XB4" authority="Muller &amp; Henle, 1837:400" authorityName="Muller &amp; Henle" authorityPageNumber="400" authorityYear="1837" box="[151,541,906,928]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="genus">
<emphasis id="B97AA719857A1440E7C6FC08FED804C6" box="[151,262,907,928]" italics="true" pageId="16" pageNumber="361">Himantura</emphasis>
Müller &amp; Henle, 1837:400
</taxonomicName>
.
<typeStatus id="54B5C5A9857A1440E579FC08FD8404C6" box="[552,602,907,928]" pageId="16" pageNumber="361">Type</typeStatus>
species
<taxonomicName id="4C0E0088857A1440E5E1FC08FBC404C6" authority="Forsskal, 1775" authorityName="Forsskal" authorityYear="1775" box="[688,1050,906,928]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="subSpecies" species="sephen" subSpecies="uarnak">
<emphasis id="B97AA719857A1440E5E1FC08FCA504C6" box="[688,891,906,928]" italics="true" pageId="16" pageNumber="361">Raja sephen uarnak</emphasis>
Forsskål, 1775
</taxonomicName>
; by subsequent designation (
<collectingCountry id="F3193B9B857A1440E26CFC09FA5C04C6" box="[1341,1410,906,928]" name="Jordan" pageId="16" pageNumber="361">Jordan</collectingCountry>
&amp; Evermann 1896:82).
</paragraph>
<paragraph id="8BB17B0B857A1440E7C6FC68FEB2020B" blockId="16.[151,1437,1003,1568]" pageId="16" pageNumber="361">
<emphasis id="B97AA719857A1440E7C6FC68FED10362" bold="true" box="[151,271,1003,1028]" pageId="16" pageNumber="361">Definition</emphasis>
. Large dasyatids (adults to
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DW) characterised by the following: robust, suboval to rhombic disc with pectoral-fin apex broadly rounded to narrowly angular; snout broadly angular, moderately elongate (1.7 2.8 times combined orbit and spiracle length); eye small and protruding; nasal curtain broadly skirt shaped; mouth narrow with 45 oral papillae (lateral papillae always present); tail very long, whip-like (length 2.53.7 times DW), its base narrow and oval to almost circular in cross section; pelvic fins small, almost entirely concealed by disc; dorsal fold and ventral folds absent; caudal sting close to tail base (distance from pectoral-fin insertion to caudalsting base 1.72.3 times interspiracular width); 13 mid-scapular thorns or in row on nape, no other scapular thorns; denticle band well developed with diffuse edge, skin on rest of disc with patchy denticles in adults; no row of enlarged median thorns on tail, small thorns and denticles posteriorly in adults; dorsal surface with strong colour pattern (spots, ocelli and/or reticulations); ventral surface white; posterior tail typically banded in young; marine, Indo
<collectingRegion id="49CAB5E9857A1440E787FAD7FED1020B" box="[214,271,1364,1389]" country="Tanzania" name="Mjini Magharibi" pageId="16" pageNumber="361">West</collectingRegion>
Pacific.
</paragraph>
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<subSubSection id="C3142880857A1440E796FAF4FDC002D3" pageId="16" pageNumber="361" type="description">
<paragraph id="8BB17B0B857A1440E796FAF4FDC002D3" blockId="16.[151,1437,1003,1568]" pageId="16" pageNumber="361">
<emphasis id="B97AA719857A1440E796FAF4FEC002F6" bold="true" box="[199,286,1399,1424]" pageId="16" pageNumber="361">Species</emphasis>
.
<taxonomicName id="4C0E0088857A1440E661FAFAFC9002F6" authority="Manjaji-Matsumoto &amp; Last, 2008" authorityName="Manjaji-Matsumoto &amp; Last" authorityYear="2008" box="[304,846,1399,1424]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B97AA719857A1440E661FAFAFE6402F6" box="[304,442,1399,1424]" italics="true" pageId="16" pageNumber="361">H. leoparda</emphasis>
Manjaji-Matsumoto &amp;
<bibRefCitation id="EF9F06FA857A1440E580FAFBFC9002F6" author="Last" box="[721,846,1400,1424]" pageId="16" pageNumber="361" refString="Last, P. R. &amp; White, W. T. (2008) Resurrection of the genus Neotrygon Castelnau (Myliobatoidei: Dasyatidae) with the description of Neotrygon picta sp. nov., a new species from northern Australia. In: Last, P. R., White, W. T. &amp; Pogonoski, J. J. (Eds.), Descriptions of New Australian Chondrichthyans. CSIRO Marine and Atmospheric Research Paper, 0 22, pp. 315 - 325." type="book chapter" year="2008">Last, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0E0088857A1440E40CFAFAFB4F02F6" authority="Gmelin, 1789" authorityName="Gmelin" authorityYear="1789" box="[861,1169,1399,1424]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="species" species="uarnak">
<emphasis id="B97AA719857A1440E40CFAFAFC0A02F6" box="[861,980,1399,1424]" italics="true" pageId="16" pageNumber="361">H. uarnak</emphasis>
(Gmelin, 1789)
</taxonomicName>
,
<taxonomicName id="4C0E0088857A1440E3F0FAFAFF0402D3" authority="Bleeker, 1852" authorityName="Bleeker" authorityYear="1852" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="species" species="undulata">
<emphasis id="B97AA719857A1440E3F0FAFAFAF502F6" box="[1185,1323,1399,1424]" italics="true" pageId="16" pageNumber="361">H. undulata</emphasis>
(Bleeker, 1852)
</taxonomicName>
, and an undescribed species.
</paragraph>
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<subSubSection id="C3142880857A1440E796FA43FBFE0179" pageId="16" pageNumber="361" type="discussion">
<paragraph id="8BB17B0B857A1440E796FA43FBFE0179" blockId="16.[151,1437,1003,1568]" pageId="16" pageNumber="361">
<emphasis id="B97AA719857A1440E796FA43FEEA02BE" bold="true" box="[199,308,1472,1496]" pageId="16" pageNumber="361">Remarks</emphasis>
. The genus
<taxonomicName id="4C0E0088857A1440E6EAFA42FDE802BE" box="[443,566,1473,1496]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="361" phylum="Chordata" rank="genus">
<emphasis id="B97AA719857A1440E6EAFA42FDE802BE" box="[443,566,1473,1496]" italics="true" pageId="16" pageNumber="361">Himantura</emphasis>
</taxonomicName>
, which was once represented widely in the Indo
<collectingRegion id="49CAB5E9857A1440E30DFA3CFB4B02BE" box="[1116,1173,1471,1496]" country="Tanzania" name="Mjini Magharibi" pageId="16" pageNumber="361">West</collectingRegion>
Pacific by ~20 species, is now restricted to a group of four large IndoPacific species with strong colour patterns. The taxonomy of this group is under review (PL &amp; MM) and a new species exists in Australasian seas.
</paragraph>
</subSubSection>
</treatment>
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