treatments-xml/data/03/98/A2/0398A275FFBFFF98FF73238D49FE5404.xml

<document id="C827FFC614E850B9DB2D01D7EF52D8E2" ID-DOI="10.5281/zenodo.183370" ID-GBIF-Dataset="5e888339-e311-4074-a736-a54ec0d2de31" ID-ISSN="1175-5326" ID-Zenodo-Dep="183370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459904775507" checkinUser="plazi" docAuthor="Fontoura, Nelson Ferreira &amp; Conter, Maria-Rotraut" docDate="2008" docId="0398A275FFBFFF98FF73238D49FE5404" docLanguage="en" docName="zt01849p034.pdf" docOrigin="Zootaxa 1849" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Parastacus brasiliensis subsp. promatensis Fontoura &amp; Conter, 2008, subsp. n." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFA1DA0DFFBEFF9EFFE4240849325736" masterDocTitle="Description of a new subspecies of the crayfish Parastacus brasiliensis (Von Martens, 1869) from São Francisco de Paula, RS, Brazil (Decapoda, Parastacidae)" masterLastPageNumber="34" masterPageNumber="28" pageNumber="29" updateTime="1698228937222" updateUser="plazi">
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<mods:title id="68EDA43BBC0B0D606342CD0AEE239B59">Description of a new subspecies of the crayfish Parastacus brasiliensis (Von Martens, 1869) from São Francisco de Paula, RS, Brazil (Decapoda, Parastacidae)</mods:title>
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<mods:namePart id="5D05AE7CBDE26994B197E6E902B3A95F">Fontoura, Nelson Ferreira</mods:namePart>
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<mods:namePart id="2F7BA4025AEFAC72711A772537E31438">Conter, Maria-Rotraut</mods:namePart>
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<mods:date id="FA99889F4FDEC89B97FF9EDC4CF4A472">2008</mods:date>
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<mods:number id="FC385CFDE308D9173A21504340871E26">1849</mods:number>
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<subSubSection id="C32B40E8FFBFFF9FFF73238D4A2E50A9" box="[151,796,1925,1951]" pageId="1" pageNumber="29" type="nomenclature">
<paragraph id="8B8E1363FFBFFF9FFF73238D4A2E50A9" blockId="1.[151,796,1925,1951]" box="[151,796,1925,1951]" pageId="1" pageNumber="29">
<heading id="D0C6A40FFFBFFF9FFF73238D4A2E50A9" bold="true" box="[151,796,1925,1951]" fontSize="11" level="1" pageId="1" pageNumber="29" reason="1">
<emphasis id="B945CF71FFBFFF9FFF73238D4A2E50A9" bold="true" box="[151,796,1925,1951]" pageId="1" pageNumber="29">
<taxonomicName id="4C3168E0FFBFFF9FFF73238D4B7750A9" ID-CoL="7W94S" box="[151,581,1925,1951]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBFFF9FFF73238D4B7750A9" bold="true" box="[151,581,1925,1951]" italics="true" pageId="1" pageNumber="29">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276720AFFBFFF9FFDA9238D4B8950A9" box="[589,699,1925,1951]" pageId="1" pageNumber="29" rank="subSpecies">subsp. n.</taxonomicNameLabel>
(
<figureCitation id="130A0FE6FFBFFF9FFD2F238D4A2350A9" box="[715,785,1925,1951]" captionStart="FIGURE 1" captionStartId="2.[151,255,1068,1092]" captionText="FIGURE 1. Parastacus brasiliensis promatensis subsp. n." pageId="1" pageNumber="29">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32B40E8FFBFFF9CFF7323D84B6557EA" lastPageId="2" lastPageNumber="30" pageId="1" pageNumber="29" type="materials_examined">
<paragraph id="8B8E1363FFBFFF9CFF7323D84B6557EA" blockId="1.[151,1436,2000,2026]" lastBlockId="2.[151,1436,152,220]" lastPageId="2" lastPageNumber="30" pageId="1" pageNumber="29">
<emphasis id="B945CF71FFBFFF9FFF7323D8487E50DC" bold="true" box="[151,332,2000,2026]" pageId="1" pageNumber="29">
<typeStatus id="548AADC1FFBFFF9FFF7323D849E050DC" box="[151,210,2000,2026]" pageId="1" pageNumber="29">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548AADC1FFBFFF9FFEB323D848FB50DC" box="[343,457,2000,2026]" pageId="1" pageNumber="29" type="holotype">Holotype</typeStatus>
: MCT 2085. 
<specimenCount id="9D37D8EAFFBFFF9FFD8323D84B4450DC" box="[615,630,2000,2026]" pageId="1" pageNumber="29" type="generic">1</specimenCount>
specimen, 
<collectingCountry id="F32653F3FFBFFF9FFD1A23D84A7A50DC" box="[766,840,2000,2026]" name="Brazil" pageId="1" pageNumber="29">Brazil</collectingCountry>
, Rio Grande do Sul, São Francisco de Paula, Pró- 
<collectingCountry id="F32653F3FFBCFF9CFF73249249E45782" box="[151,214,154,180]" name="Malta" pageId="2" pageNumber="30">Mata</collectingCountry>
, Garapiá stream (
<geoCoordinate id="EE0575A4FFBCFF9CFE4824924B785782" box="[428,586,152,180]" direction="south" orientation="latitude" pageId="2" pageNumber="30" precision="1" value="-29.489517">29o29.371’ S</geoCoordinate>
; 
<geoCoordinate id="EE0575A4FFBCFF9CFDBE24924A305782" box="[602,770,152,180]" direction="west" orientation="longitude" pageId="2" pageNumber="30" precision="1" value="-50.23">50o13.800’ W</geoCoordinate>
), 
<date id="FF8F35A3FFBCFF9CFCFF24924AA85782" box="[795,922,154,180]" pageId="2" pageNumber="30" value="1997-11-22">22.xi.1997</date>
, Leg. M. Conter. 
<typeStatus id="548AADC1FFBCFF9CFB9724924DDB5782" box="[1139,1257,154,180]" pageId="2" pageNumber="30" type="paratype">Paratypes</typeStatus>
: MCT 2086. 
<specimenCount id="9D37D8EAFFBCFF9CFA6924924CAE5782" box="[1421,1436,154,180]" pageId="2" pageNumber="30" type="generic">8</specimenCount>
specimens, same location as 
<typeStatus id="548AADC1FFBCFF9CFE0C24CA4B6157EA" box="[488,595,194,220]" pageId="2" pageNumber="30" type="holotype">holotype</typeStatus>
.
</paragraph>
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<caption id="DF4E43EBFFBCFF9CFF7320244A3E5372" box="[151,780,1068,1092]" pageId="2" pageNumber="30">
<paragraph id="8B8E1363FFBCFF9CFF7320244A3E5372" blockId="2.[151,780,1068,1092]" box="[151,780,1068,1092]" pageId="2" pageNumber="30">
<emphasis id="B945CF71FFBCFF9CFF732024482B5372" bold="true" box="[151,281,1068,1092]" pageId="2" pageNumber="30">FIGURE 1.</emphasis>
<taxonomicName id="4C3168E0FFBCFF9CFEC420254B935372" box="[288,673,1069,1092]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBCFF9CFEC420254B935372" box="[288,673,1069,1092]" italics="true" pageId="2" pageNumber="30">Parastacus brasiliensis promatensis</emphasis>
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<taxonomicNameLabel id="A276720AFFBCFF9CFD4C20244A3E5372" box="[680,780,1068,1092]" pageId="2" pageNumber="30" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C32B40E8FFBCFF9CFF22207F4817523F" pageId="2" pageNumber="30" type="etymology">
<paragraph id="8B8E1363FFBCFF9CFF22207F4817523F" blockId="2.[151,1437,1143,1289]" pageId="2" pageNumber="30">
<emphasis id="B945CF71FFBCFF9CFF22207F486153A7" bold="true" box="[198,339,1143,1169]" pageId="2" pageNumber="30">Etymology.</emphasis>
The subspecies was named after the only location from where it is known. 
<emphasis id="B945CF71FFBCFF9CFF22209748EE538F" bold="true" box="[198,476,1183,1209]" pageId="2" pageNumber="30">Diagnostic characters.</emphasis>
Identification should be based on a set of morphometrical data (
<tableCitation id="C6B326D8FFBCFF9CFB2D20974C1F538F" box="[1225,1325,1183,1209]" captionStart="TABLE 2" captionStartId="2.[151,239,1343,1367]" captionTargetBox="[159,1324,1399,1977]" captionTargetPageId="2" captionText="TABLE 2. Morphometrical measurements of Parastacus brasiliensis promatensis subsp. n.." httpUri="http://table.plazi.org/id/DF4E43EBFFBCFF9CFF7321374D415260" pageId="2" pageNumber="30" tableUuid="DF4E43EBFFBCFF9CFF7321374D415260">Tables 2</tableCitation>
and 
<tableCitation id="C6B326D8FFBCFF9CFA8020974C41538F" box="[1380,1395,1183,1209]" captionStart="TABLE 3" captionStartId="3.[151,239,151,175]" captionTargetBox="[159,1323,208,657]" captionTargetPageId="3" captionText="TABLE 3. Ratios for morphometrical identification of Parastacus brasiliensis promatensis subsp. n.." httpUri="http://table.plazi.org/id/DF4E43EBFFBDFF9DFF73249F4DE55799" pageId="2" pageNumber="30" tableUuid="DF4E43EBFFBDFF9DFF73249F4DE55799">3</tableCitation>
) as there is no morphological character that clearly distinguishes 
<taxonomicName id="4C3168E0FFBCFF9CFC9520C04D9E53D7" box="[881,1196,1224,1249]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBCFF9CFC9520C04D9E53D7" box="[881,1196,1224,1249]" italics="true" pageId="2" pageNumber="30">P. brasiliensis promatensis</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C3168E0FFBCFF9CFB1320C0482C523F" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBCFF9CFB1320C0482C523F" italics="true" pageId="2" pageNumber="30">P. brasiliensis brasiliensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF4E43EBFFBCFF9CFF7321374D415260" ID-Table-UUID="DF4E43EBFFBCFF9CFF7321374D415260" box="[151,1139,1343,1367]" httpUri="http://table.plazi.org/id/DF4E43EBFFBCFF9CFF7321374D415260" pageId="2" pageNumber="30" targetBox="[159,1324,1399,1977]" targetIsTable="true" targetPageId="2">
<paragraph id="8B8E1363FFBCFF9CFF7321374D415260" blockId="2.[151,1139,1343,1367]" box="[151,1139,1343,1367]" pageId="2" pageNumber="30">
<emphasis id="B945CF71FFBCFF9CFF732137483A5261" bold="true" box="[151,264,1343,1367]" pageId="2" pageNumber="30">TABLE 2.</emphasis>
Morphometrical measurements of 
<taxonomicName id="4C3168E0FFBCFF9CFD6421374D305260" box="[640,1026,1343,1366]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBCFF9CFD6421374D305260" box="[640,1026,1343,1366]" italics="true" pageId="2" pageNumber="30">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
<emphasis id="B945CF71FFBCFF9CFBED21374D5E5261" bold="true" box="[1033,1132,1343,1367]" pageId="2" pageNumber="30">
<taxonomicNameLabel id="A276720AFFBCFF9CFBED21374D5E5261" box="[1033,1132,1343,1367]" pageId="2" pageNumber="30" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<paragraph id="8B8E1363FFBCFF9CFF7B217F4DDD508F" pageId="2" pageNumber="30">
<table id="F931E1C3FFBC0061FF7B217F4C1E508F" box="[159,1324,1399,1977]" gridcols="5" gridrows="14" pageId="2" pageNumber="30">
<tr id="35011121FFBC0061FF7B217F4C1E52B8" box="[159,1324,1399,1422]" gridrow="0" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B217F480252B8" box="[159,304,1399,1422]" gridcol="0" gridrow="0" pageId="2" pageNumber="30">Measurement</th>
<th id="76D0785DFFBC0061FDEA217F4B5452B8" box="[526,614,1399,1422]" gridcol="1" gridrow="0" pageId="2" pageNumber="30">Average</th>
<th id="76D0785DFFBC0061FD10217F4A1152B8" box="[756,803,1399,1422]" gridcol="2" gridrow="0" pageId="2" pageNumber="30">SD</th>
<th id="76D0785DFFBC0061FC3E217F4D7B52B8" box="[986,1097,1399,1422]" gridcol="3" gridrow="0" pageId="2" pageNumber="30">Maximum</th>
<th id="76D0785DFFBC0061FB24217F4C1E52B8" box="[1216,1324,1399,1422]" gridcol="4" gridrow="0" pageId="2" pageNumber="30">Minimum</th>
</tr>
<tr id="35011121FFBC0061FF7B21AA4C1E528F" box="[159,1324,1442,1465]" gridrow="1" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B21AA4802528F" box="[159,304,1442,1465]" gridcol="0" gridrow="1" pageId="2" pageNumber="30">HCHE</th>
<td id="76D0785DFFBC0061FDEA21AA4B54528F" box="[526,614,1442,1465]" gridcol="1" gridrow="1" pageId="2" pageNumber="30">10.45</td>
<td id="76D0785DFFBC0061FD1021AA4A11528F" box="[756,803,1442,1465]" gridcol="2" gridrow="1" pageId="2" pageNumber="30">3.17</td>
<td id="76D0785DFFBC0061FC3E21AA4D7B528F" box="[986,1097,1442,1465]" gridcol="3" gridrow="1" pageId="2" pageNumber="30">18.00</td>
<td id="76D0785DFFBC0061FB2421AA4C1E528F" box="[1216,1324,1442,1465]" gridcol="4" gridrow="1" pageId="2" pageNumber="30">5.00</td>
</tr>
<tr id="35011121FFBC0061FF7B21C44C1E52D5" box="[159,1324,1484,1507]" gridrow="2" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B21C4480252D5" box="[159,304,1484,1507]" gridcol="0" gridrow="2" pageId="2" pageNumber="30">TCHE</th>
<td id="76D0785DFFBC0061FDEA21C44B5452D5" box="[526,614,1484,1507]" gridcol="1" gridrow="2" pageId="2" pageNumber="30">6.05</td>
<td id="76D0785DFFBC0061FD1021C44A1152D5" box="[756,803,1484,1507]" gridcol="2" gridrow="2" pageId="2" pageNumber="30">1.85</td>
<td id="76D0785DFFBC0061FC3E21C44D7B52D5" box="[986,1097,1484,1507]" gridcol="3" gridrow="2" pageId="2" pageNumber="30">10.80</td>
<td id="76D0785DFFBC0061FB2421C44C1E52D5" box="[1216,1324,1484,1507]" gridcol="4" gridrow="2" pageId="2" pageNumber="30">2.80</td>
</tr>
<tr id="35011121FFBC0061FF7B21FF4C1E5138" box="[159,1324,1527,1550]" gridrow="3" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B21FF48025138" box="[159,304,1527,1550]" gridcol="0" gridrow="3" pageId="2" pageNumber="30">LCHE</th>
<td id="76D0785DFFBC0061FDEA21FF4B545138" box="[526,614,1527,1550]" gridcol="1" gridrow="3" pageId="2" pageNumber="30">26.27</td>
<td id="76D0785DFFBC0061FD1021FF4A115138" box="[756,803,1527,1550]" gridcol="2" gridrow="3" pageId="2" pageNumber="30">9.75</td>
<td id="76D0785DFFBC0061FC3E21FF4D7B5138" box="[986,1097,1527,1550]" gridcol="3" gridrow="3" pageId="2" pageNumber="30">52.20</td>
<td id="76D0785DFFBC0061FB2421FF4C1E5138" box="[1216,1324,1527,1550]" gridcol="4" gridrow="3" pageId="2" pageNumber="30">11.70</td>
</tr>
<tr id="35011121FFBC0061FF7B222A4C1E510F" box="[159,1324,1570,1593]" gridrow="4" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B222A4802510F" box="[159,304,1570,1593]" gridcol="0" gridrow="4" pageId="2" pageNumber="30">LCAR5</th>
<td id="76D0785DFFBC0061FDEA222A4B54510F" box="[526,614,1570,1593]" gridcol="1" gridrow="4" pageId="2" pageNumber="30">4.86</td>
<td id="76D0785DFFBC0061FD10222A4A11510F" box="[756,803,1570,1593]" gridcol="2" gridrow="4" pageId="2" pageNumber="30">1.39</td>
<td id="76D0785DFFBC0061FC3E222A4D7B510F" box="[986,1097,1570,1593]" gridcol="3" gridrow="4" pageId="2" pageNumber="30">7.90</td>
<td id="76D0785DFFBC0061FB24222A4C1E510F" box="[1216,1324,1570,1593]" gridcol="4" gridrow="4" pageId="2" pageNumber="30">2.60</td>
</tr>
<tr id="35011121FFBC0061FF7B22444C1E5155" box="[159,1324,1612,1635]" gridrow="5" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B224448025155" box="[159,304,1612,1635]" gridcol="0" gridrow="5" pageId="2" pageNumber="30">LDAC5</th>
<td id="76D0785DFFBC0061FDEA22444B545155" box="[526,614,1612,1635]" gridcol="1" gridrow="5" pageId="2" pageNumber="30">3.63</td>
<td id="76D0785DFFBC0061FD1022444A115155" box="[756,803,1612,1635]" gridcol="2" gridrow="5" pageId="2" pageNumber="30">0.86</td>
<td id="76D0785DFFBC0061FC3E22444D7B5155" box="[986,1097,1612,1635]" gridcol="3" gridrow="5" pageId="2" pageNumber="30">5.20</td>
<td id="76D0785DFFBC0061FB2422444C1E5155" box="[1216,1324,1612,1635]" gridcol="4" gridrow="5" pageId="2" pageNumber="30">2.30</td>
</tr>
<tr id="35011121FFBC0061FF7B227F4C1E51B8" box="[159,1324,1655,1678]" gridrow="6" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B227F480251B8" box="[159,304,1655,1678]" gridcol="0" gridrow="6" pageId="2" pageNumber="30">LMER5</th>
<td id="76D0785DFFBC0061FDEA227F4B5451B8" box="[526,614,1655,1678]" gridcol="1" gridrow="6" pageId="2" pageNumber="30">8.27</td>
<td id="76D0785DFFBC0061FD10227F4A1151B8" box="[756,803,1655,1678]" gridcol="2" gridrow="6" pageId="2" pageNumber="30">2.18</td>
<td id="76D0785DFFBC0061FC3E227F4D7B51B8" box="[986,1097,1655,1678]" gridcol="3" gridrow="6" pageId="2" pageNumber="30">14.10</td>
<td id="76D0785DFFBC0061FB24227F4C1E51B8" box="[1216,1324,1655,1678]" gridcol="4" gridrow="6" pageId="2" pageNumber="30">4.70</td>
</tr>
<tr id="35011121FFBC0061FF7B22AA4C1E518F" box="[159,1324,1698,1721]" gridrow="7" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B22AA4802518F" box="[159,304,1698,1721]" gridcol="0" gridrow="7" pageId="2" pageNumber="30">LPRO5</th>
<td id="76D0785DFFBC0061FDEA22AA4B54518F" box="[526,614,1698,1721]" gridcol="1" gridrow="7" pageId="2" pageNumber="30">8.35</td>
<td id="76D0785DFFBC0061FD1022AA4A11518F" box="[756,803,1698,1721]" gridcol="2" gridrow="7" pageId="2" pageNumber="30">2.17</td>
<td id="76D0785DFFBC0061FC3E22AA4D7B518F" box="[986,1097,1698,1721]" gridcol="3" gridrow="7" pageId="2" pageNumber="30">12.20</td>
<td id="76D0785DFFBC0061FB2422AA4C1E518F" box="[1216,1324,1698,1721]" gridcol="4" gridrow="7" pageId="2" pageNumber="30">4.60</td>
</tr>
<tr id="35011121FFBC0061FF7B22C44C1E51D5" box="[159,1324,1740,1763]" gridrow="8" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B22C4480251D5" box="[159,304,1740,1763]" gridcol="0" gridrow="8" pageId="2" pageNumber="30">LCARAP</th>
<td id="76D0785DFFBC0061FDEA22C44B5451D5" box="[526,614,1740,1763]" gridcol="1" gridrow="8" pageId="2" pageNumber="30">33.33</td>
<td id="76D0785DFFBC0061FD1022C44A1151D5" box="[756,803,1740,1763]" gridcol="2" gridrow="8" pageId="2" pageNumber="30">7.65</td>
<td id="76D0785DFFBC0061FC3E22C44D7B51D5" box="[986,1097,1740,1763]" gridcol="3" gridrow="8" pageId="2" pageNumber="30">46.20</td>
<td id="76D0785DFFBC0061FB2422C44C1E51D5" box="[1216,1324,1740,1763]" gridcol="4" gridrow="8" pageId="2" pageNumber="30">19.60</td>
</tr>
<tr id="35011121FFBC0061FF7B22FF4C1E5038" box="[159,1324,1783,1806]" gridrow="9" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B22FF48025038" box="[159,304,1783,1806]" gridcol="0" gridrow="9" pageId="2" pageNumber="30">LABD</th>
<td id="76D0785DFFBC0061FDEA22FF4B545038" box="[526,614,1783,1806]" gridcol="1" gridrow="9" pageId="2" pageNumber="30">36.15</td>
<td id="76D0785DFFBC0061FD1022FF4A115038" box="[756,803,1783,1806]" gridcol="2" gridrow="9" pageId="2" pageNumber="30">7.87</td>
<td id="76D0785DFFBC0061FC3E22FF4D7B5038" box="[986,1097,1783,1806]" gridcol="3" gridrow="9" pageId="2" pageNumber="30">49.00</td>
<td id="76D0785DFFBC0061FB2422FF4C1E5038" box="[1216,1324,1783,1806]" gridcol="4" gridrow="9" pageId="2" pageNumber="30">20.50</td>
</tr>
<tr id="35011121FFBC0061FF7B232A4C1E500F" box="[159,1324,1826,1849]" gridrow="10" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B232A4802500F" box="[159,304,1826,1849]" gridcol="0" gridrow="10" pageId="2" pageNumber="30">LROS</th>
<td id="76D0785DFFBC0061FDEA232A4B54500F" box="[526,614,1826,1849]" gridcol="1" gridrow="10" pageId="2" pageNumber="30">5.22</td>
<td id="76D0785DFFBC0061FD10232A4A11500F" box="[756,803,1826,1849]" gridcol="2" gridrow="10" pageId="2" pageNumber="30">1.01</td>
<td id="76D0785DFFBC0061FC3E232A4D7B500F" box="[986,1097,1826,1849]" gridcol="3" gridrow="10" pageId="2" pageNumber="30">6.80</td>
<td id="76D0785DFFBC0061FB24232A4C1E500F" box="[1216,1324,1826,1849]" gridcol="4" gridrow="10" pageId="2" pageNumber="30">3.10</td>
</tr>
<tr id="35011121FFBC0061FF7B23444C1E5055" box="[159,1324,1868,1891]" gridrow="11" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B234448025055" box="[159,304,1868,1891]" gridcol="0" gridrow="11" pageId="2" pageNumber="30">LTEL</th>
<td id="76D0785DFFBC0061FDEA23444B545055" box="[526,614,1868,1891]" gridcol="1" gridrow="11" pageId="2" pageNumber="30">11.15</td>
<td id="76D0785DFFBC0061FD1023444A115055" box="[756,803,1868,1891]" gridcol="2" gridrow="11" pageId="2" pageNumber="30">2.65</td>
<td id="76D0785DFFBC0061FC3E23444D7B5055" box="[986,1097,1868,1891]" gridcol="3" gridrow="11" pageId="2" pageNumber="30">15.60</td>
<td id="76D0785DFFBC0061FB2423444C1E5055" box="[1216,1324,1868,1891]" gridcol="4" gridrow="11" pageId="2" pageNumber="30">6.10</td>
</tr>
<tr id="35011121FFBC0061FF7B237F4C1E50B8" box="[159,1324,1911,1934]" gridrow="12" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B237F480250B8" box="[159,304,1911,1934]" gridcol="0" gridrow="12" pageId="2" pageNumber="30">LURO1</th>
<td id="76D0785DFFBC0061FDEA237F4B5450B8" box="[526,614,1911,1934]" gridcol="1" gridrow="12" pageId="2" pageNumber="30">8.95</td>
<td id="76D0785DFFBC0061FD10237F4A1150B8" box="[756,803,1911,1934]" gridcol="2" gridrow="12" pageId="2" pageNumber="30">2.48</td>
<td id="76D0785DFFBC0061FC3E237F4D7B50B8" box="[986,1097,1911,1934]" gridcol="3" gridrow="12" pageId="2" pageNumber="30">14.50</td>
<td id="76D0785DFFBC0061FB24237F4C1E50B8" box="[1216,1324,1911,1934]" gridcol="4" gridrow="12" pageId="2" pageNumber="30">4.80</td>
</tr>
<tr id="35011121FFBC0061FF7B23AA4C1E508F" box="[159,1324,1954,1977]" gridrow="13" pageId="2" pageNumber="30">
<th id="76D0785DFFBC0061FF7B23AA4802508F" box="[159,304,1954,1977]" gridcol="0" gridrow="13" pageId="2" pageNumber="30">LURO2</th>
<td id="76D0785DFFBC0061FDEA23AA4B54508F" box="[526,614,1954,1977]" gridcol="1" gridrow="13" pageId="2" pageNumber="30">10.64</td>
<td id="76D0785DFFBC0061FD1023AA4A11508F" box="[756,803,1954,1977]" gridcol="2" gridrow="13" pageId="2" pageNumber="30">2.74</td>
<td id="76D0785DFFBC0061FC3E23AA4D7B508F" box="[986,1097,1954,1977]" gridcol="3" gridrow="13" pageId="2" pageNumber="30">16.00</td>
<td id="76D0785DFFBC0061FB2423AA4C1E508F" box="[1216,1324,1954,1977]" gridcol="4" gridrow="13" pageId="2" pageNumber="30">5.70</td>
</tr>
</table>
</paragraph>
<caption id="DF4E43EBFFBDFF9DFF73249F4DE55799" ID-Table-UUID="DF4E43EBFFBDFF9DFF73249F4DE55799" box="[151,1239,151,175]" httpUri="http://table.plazi.org/id/DF4E43EBFFBDFF9DFF73249F4DE55799" pageId="3" pageNumber="31" targetBox="[159,1323,208,657]" targetIsTable="true" targetPageId="3">
<paragraph id="8B8E1363FFBDFF9DFF73249F4DE55799" blockId="3.[151,1239,151,175]" box="[151,1239,151,175]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF73249F483A5799" bold="true" box="[151,264,151,175]" pageId="3" pageNumber="31">TABLE 3.</emphasis>
Ratios for morphometrical identification of 
<taxonomicName id="4C3168E0FFBDFF9DFD0024904D575799" box="[740,1125,152,175]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBDFF9DFD0024904D575799" box="[740,1125,152,175]" italics="true" pageId="3" pageNumber="31">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
<emphasis id="B945CF71FFBDFF9DFB88249F4DFD5799" bold="true" box="[1132,1231,151,175]" pageId="3" pageNumber="31">
<taxonomicNameLabel id="A276720AFFBDFF9DFB88249F4DFD5799" box="[1132,1231,151,175]" pageId="3" pageNumber="31" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<paragraph id="8B8E1363FFBDFF9DFF7B24D84C3855A7" pageId="3" pageNumber="31">
<table id="F931E1C3FFBD0061FF7B24D84C1955A7" box="[159,1323,208,657]" gridcols="5" gridrows="11" pageId="3" pageNumber="31">
<tr id="35011121FFBD0061FF7B24D84C1957D1" box="[159,1323,208,231]" gridrow="0" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B24D8481C57D1" box="[159,302,208,231]" gridcol="0" gridrow="0" pageId="3" pageNumber="31">Ratio</th>
<th id="76D0785DFFBD0061FDEA24D84B5457D1" box="[526,614,208,231]" gridcol="1" gridrow="0" pageId="3" pageNumber="31">Average</th>
<th id="76D0785DFFBD0061FD1024D84A0F57D1" box="[756,829,208,231]" gridcol="2" gridrow="0" pageId="3" pageNumber="31">SD</th>
<th id="76D0785DFFBD0061FC3E24D84D7B57D1" box="[986,1097,208,231]" gridcol="3" gridrow="0" pageId="3" pageNumber="31">Maximum</th>
<th id="76D0785DFFBD0061FB2424D84C1957D1" box="[1216,1323,208,231]" gridcol="4" gridrow="0" pageId="3" pageNumber="31">Minimum</th>
</tr>
<tr id="35011121FFBD0061FF7B24F24C195627" box="[159,1323,250,273]" gridrow="1" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B24F2481C5627" box="[159,302,250,273]" gridcol="0" gridrow="1" pageId="3" pageNumber="31">Labd/Lcarap</th>
<td id="76D0785DFFBD0061FDEA24F24B545627" box="[526,614,250,273]" gridcol="1" gridrow="1" pageId="3" pageNumber="31">1.0880</td>
<td id="76D0785DFFBD0061FD1024F24A0F5627" box="[756,829,250,273]" gridcol="2" gridrow="1" pageId="3" pageNumber="31">0.0430</td>
<td id="76D0785DFFBD0061FC3E24F24D7B5627" box="[986,1097,250,273]" gridcol="3" gridrow="1" pageId="3" pageNumber="31">1.2188</td>
<td id="76D0785DFFBD0061FB2424F24C195627" box="[1216,1323,250,273]" gridcol="4" gridrow="1" pageId="3" pageNumber="31">1.0310</td>
</tr>
<tr id="35011121FFBD0061FF7B252D4C19560A" box="[159,1323,293,316]" gridrow="2" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B252D481C560A" box="[159,302,293,316]" gridcol="0" gridrow="2" pageId="3" pageNumber="31">Lros/Lcarap</th>
<td id="76D0785DFFBD0061FDEA252D4B54560A" box="[526,614,293,316]" gridcol="1" gridrow="2" pageId="3" pageNumber="31">0.1583</td>
<td id="76D0785DFFBD0061FD10252D4A0F560A" box="[756,829,293,316]" gridcol="2" gridrow="2" pageId="3" pageNumber="31">0.0159</td>
<td id="76D0785DFFBD0061FC3E252D4D7B560A" box="[986,1097,293,316]" gridcol="3" gridrow="2" pageId="3" pageNumber="31">0.2227</td>
<td id="76D0785DFFBD0061FB24252D4C19560A" box="[1216,1323,293,316]" gridcol="4" gridrow="2" pageId="3" pageNumber="31">0.1432</td>
</tr>
<tr id="35011121FFBD0061FF7B25584C195651" box="[159,1323,336,359]" gridrow="3" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B2558481C5651" box="[159,302,336,359]" gridcol="0" gridrow="3" pageId="3" pageNumber="31">Ltel/Lcarap</th>
<td id="76D0785DFFBD0061FDEA25584B545651" box="[526,614,336,359]" gridcol="1" gridrow="3" pageId="3" pageNumber="31">0.3340</td>
<td id="76D0785DFFBD0061FD1025584A0F5651" box="[756,829,336,359]" gridcol="2" gridrow="3" pageId="3" pageNumber="31">0.0121</td>
<td id="76D0785DFFBD0061FC3E25584D7B5651" box="[986,1097,336,359]" gridcol="3" gridrow="3" pageId="3" pageNumber="31">0.3596</td>
<td id="76D0785DFFBD0061FB2425584C195651" box="[1216,1323,336,359]" gridcol="4" gridrow="3" pageId="3" pageNumber="31">0.3112</td>
</tr>
<tr id="35011121FFBD0061FF7B25724C1956A7" box="[159,1323,378,401]" gridrow="4" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B2572481C56A7" box="[159,302,378,401]" gridcol="0" gridrow="4" pageId="3" pageNumber="31">Luro1/Ltel</th>
<td id="76D0785DFFBD0061FDEA25724B5456A7" box="[526,614,378,401]" gridcol="1" gridrow="4" pageId="3" pageNumber="31">0.7984</td>
<td id="76D0785DFFBD0061FD1025724A0F56A7" box="[756,829,378,401]" gridcol="2" gridrow="4" pageId="3" pageNumber="31">0.0561</td>
<td id="76D0785DFFBD0061FC3E25724D7B56A7" box="[986,1097,378,401]" gridcol="3" gridrow="4" pageId="3" pageNumber="31">0.9932</td>
<td id="76D0785DFFBD0061FB2425724C1956A7" box="[1216,1323,378,401]" gridcol="4" gridrow="4" pageId="3" pageNumber="31">0.7121</td>
</tr>
<tr id="35011121FFBD0061FF7B25AD4C19568A" box="[159,1323,421,444]" gridrow="5" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B25AD481C568A" box="[159,302,421,444]" gridcol="0" gridrow="5" pageId="3" pageNumber="31">Luro2/Ltel</th>
<td id="76D0785DFFBD0061FDEA25AD4B54568A" box="[526,614,421,444]" gridcol="1" gridrow="5" pageId="3" pageNumber="31">0.9510</td>
<td id="76D0785DFFBD0061FD1025AD4A0F568A" box="[756,829,421,444]" gridcol="2" gridrow="5" pageId="3" pageNumber="31">0.0427</td>
<td id="76D0785DFFBD0061FC3E25AD4D7B568A" box="[986,1097,421,444]" gridcol="3" gridrow="5" pageId="3" pageNumber="31">1.0959</td>
<td id="76D0785DFFBD0061FB2425AD4C19568A" box="[1216,1323,421,444]" gridcol="4" gridrow="5" pageId="3" pageNumber="31">0.9000</td>
</tr>
<tr id="35011121FFBD0061FF7B25D84C1956D1" box="[159,1323,464,487]" gridrow="6" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B25D8481C56D1" box="[159,302,464,487]" gridcol="0" gridrow="6" pageId="3" pageNumber="31">Hche/Lche</th>
<td id="76D0785DFFBD0061FDEA25D84B5456D1" box="[526,614,464,487]" gridcol="1" gridrow="6" pageId="3" pageNumber="31">0.4097</td>
<td id="76D0785DFFBD0061FD1025D84A0F56D1" box="[756,829,464,487]" gridcol="2" gridrow="6" pageId="3" pageNumber="31">0.0478</td>
<td id="76D0785DFFBD0061FC3E25D84D7B56D1" box="[986,1097,464,487]" gridcol="3" gridrow="6" pageId="3" pageNumber="31">0.4710</td>
<td id="76D0785DFFBD0061FB2425D84C1956D1" box="[1216,1323,464,487]" gridcol="4" gridrow="6" pageId="3" pageNumber="31">0.2907</td>
</tr>
<tr id="35011121FFBD0061FF7B25F24C195527" box="[159,1323,506,529]" gridrow="7" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B25F2481C5527" box="[159,302,506,529]" gridcol="0" gridrow="7" pageId="3" pageNumber="31">Tche/Lche</th>
<td id="76D0785DFFBD0061FDEA25F24B545527" box="[526,614,506,529]" gridcol="1" gridrow="7" pageId="3" pageNumber="31">0.2382</td>
<td id="76D0785DFFBD0061FD1025F24A0F5527" box="[756,829,506,529]" gridcol="2" gridrow="7" pageId="3" pageNumber="31">0.0330</td>
<td id="76D0785DFFBD0061FC3E25F24D7B5527" box="[986,1097,506,529]" gridcol="3" gridrow="7" pageId="3" pageNumber="31">0.2774</td>
<td id="76D0785DFFBD0061FB2425F24C195527" box="[1216,1323,506,529]" gridcol="4" gridrow="7" pageId="3" pageNumber="31">0.1547</td>
</tr>
<tr id="35011121FFBD0061FF7B262D4C19550A" box="[159,1323,549,572]" gridrow="8" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B262D481C550A" box="[159,302,549,572]" gridcol="0" gridrow="8" pageId="3" pageNumber="31">Lpro5/Lcar5</th>
<td id="76D0785DFFBD0061FDEA262D4B54550A" box="[526,614,549,572]" gridcol="1" gridrow="8" pageId="3" pageNumber="31">0.7492</td>
<td id="76D0785DFFBD0061FD10262D4A0F550A" box="[756,829,549,572]" gridcol="2" gridrow="8" pageId="3" pageNumber="31">0.0360</td>
<td id="76D0785DFFBD0061FC3E262D4D7B550A" box="[986,1097,549,572]" gridcol="3" gridrow="8" pageId="3" pageNumber="31">0.8248</td>
<td id="76D0785DFFBD0061FB24262D4C19550A" box="[1216,1323,549,572]" gridcol="4" gridrow="8" pageId="3" pageNumber="31">0.6667</td>
</tr>
<tr id="35011121FFBD0061FF7B26584C195551" box="[159,1323,592,615]" gridrow="9" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B2658481C5551" box="[159,302,592,615]" gridcol="0" gridrow="9" pageId="3" pageNumber="31">Ldac5/Lcar5</th>
<td id="76D0785DFFBD0061FDEA26584B545551" box="[526,614,592,615]" gridcol="1" gridrow="9" pageId="3" pageNumber="31">0.3287</td>
<td id="76D0785DFFBD0061FD1026584A0F5551" box="[756,829,592,615]" gridcol="2" gridrow="9" pageId="3" pageNumber="31">0.0303</td>
<td id="76D0785DFFBD0061FC3E26584D7B5551" box="[986,1097,592,615]" gridcol="3" gridrow="9" pageId="3" pageNumber="31">0.3868</td>
<td id="76D0785DFFBD0061FB2426584C195551" box="[1216,1323,592,615]" gridcol="4" gridrow="9" pageId="3" pageNumber="31">0.2581</td>
</tr>
<tr id="35011121FFBD0061FF7B26724C1955A7" box="[159,1323,634,657]" gridrow="10" pageId="3" pageNumber="31">
<th id="76D0785DFFBD0061FF7B2672481C55A7" box="[159,302,634,657]" gridcol="0" gridrow="10" pageId="3" pageNumber="31">Lmer5/Lpro5</th>
<td id="76D0785DFFBD0061FDEA26724B5455A7" box="[526,614,634,657]" gridcol="1" gridrow="10" pageId="3" pageNumber="31">0.9936</td>
<td id="76D0785DFFBD0061FD1026724A0F55A7" box="[756,829,634,657]" gridcol="2" gridrow="10" pageId="3" pageNumber="31">0.0684</td>
<td id="76D0785DFFBD0061FC3E26724D7B55A7" box="[986,1097,634,657]" gridcol="3" gridrow="10" pageId="3" pageNumber="31">1.1949</td>
<td id="76D0785DFFBD0061FB2426724C1955A7" box="[1216,1323,634,657]" gridcol="4" gridrow="10" pageId="3" pageNumber="31">0.9027</td>
</tr>
</table>
</paragraph>
<subSubSection id="C32B40E8FFBDFF9DFF2226F74C785304" pageId="3" pageNumber="31" type="description">
<paragraph id="8B8E1363FFBDFF9DFF2226F74C785304" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF2226F7486C542F" bold="true" box="[198,350,767,793]" pageId="3" pageNumber="31">Description.</emphasis>
Carapace laterally compressed, surface smooth, with small pores and scattered hairs. Subtriangular rostrum, usually extending up to the second segment of the antennula. Rostrum surface flat, edged by crests that converge to the apex, ending in a short and depressed spine. Long hairs, growing down to the orbit from the rostral margins. The aureole is narrower at the centre, with branchial crests anteriorly and posteriorly diverging. Palm long. Fingers are almost the same size, straight but converging distally, with small gap between them. Both fingers finish in a spinous processes and have small teeth on the cutting edges, with a larger tooth appearing on the proximal third. Abdomen smooth, with rounded abdominal pleura. Telson with lateral margins converging distally to a pair of spines and then finishing in a rounded posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C32B40E8FFBDFF9DFF2220374BFA5244" pageId="3" pageNumber="31" type="materials_examined">
<paragraph id="8B8E1363FFBDFF9DFF22203748A75214" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF22203748CE536F" bold="true" box="[198,508,1087,1113]" pageId="3" pageNumber="31">
<typeStatus id="548AADC1FFBDFF9DFF222037480A536F" box="[198,312,1087,1113]" pageId="3" pageNumber="31" type="holotype">Holotype</typeStatus>
measurements.
</emphasis>
Height of right cheliped 
<quantity id="4CC9BE86FFBDFF9DFCCF20484AAA536C" box="[811,920,1088,1114]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="31" unit="mm" value="13.5">13.5 mm</quantity>
, thickness of right cheliped 
<quantity id="4CC9BE86FFBDFF9DFB0C20484C74536C" box="[1256,1350,1088,1114]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="3" pageNumber="31" unit="mm" value="7.2">7.2 mm</quantity>
, length of rigth cheliped 
<quantity id="4CC9BE86FFBDFF9DFEBA206048FB53B4" box="[350,457,1128,1154]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="3" pageNumber="31" unit="mm" value="32.4">32.4 mm</quantity>
, length of carpus (5th rigth pereiopod) 
<quantity id="4CC9BE86FFBDFF9DFC7320604AC153B4" box="[919,1011,1128,1154]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="31" unit="mm" value="6.3">6.3 mm</quantity>
, length of dactylus (5th right pereiopod) 
<quantity id="4CC9BE86FFBDFF9DFF3020984803539C" box="[212,305,1168,1194]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="31" unit="mm" value="3.9">3.9 mm</quantity>
, length of merus (5th right pereiopod) 
<quantity id="4CC9BE86FFBDFF9DFD1C20984A67539C" box="[760,853,1168,1194]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="3" pageNumber="31" unit="mm" value="8.8">8.8 mm</quantity>
, length of propodus (5th right pereiopod) 
<quantity id="4CC9BE86FFBDFF9DFAA620984CAA539C" box="[1346,1432,1168,1194]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="3" pageNumber="31" unit="mm" value="8.9">8.9mm</quantity>
, length of the carapace 
<quantity id="4CC9BE86FFBDFF9DFE7920B04B3A53E4" box="[413,520,1208,1234]" metricMagnitude="-2" metricUnit="m" metricValue="3.67" pageId="3" pageNumber="31" unit="mm" value="36.7">36.7 mm</quantity>
, length of the abdomen 
<quantity id="4CC9BE86FFBDFF9DFCFA20B04AB853E4" box="[798,906,1208,1234]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="3" pageNumber="31" unit="mm" value="40.6">40.6 mm</quantity>
, length of the rostrum (to the posterior orbital edge) 
<quantity id="4CC9BE86FFBDFF9DFF0020E8487753CC" box="[228,325,1248,1274]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="31" unit="mm" value="5.9">5.9 mm</quantity>
, length of telson 
<quantity id="4CC9BE86FFBDFF9DFDC420E84BA353CC" box="[544,657,1248,1274]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="31" unit="mm" value="12.6">12.6 mm</quantity>
, length of uropod 1 (endopodite) 
<quantity id="4CC9BE86FFBDFF9DFBD120E84D9753CC" box="[1077,1189,1248,1274]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="3" pageNumber="31" unit="mm" value="10.1">10.1 mm</quantity>
, length of uropod 2 (exopodite) 
<quantity id="4CC9BE86FFBDFF9DFEC1210048A05214" box="[293,402,1288,1314]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="3" pageNumber="31" unit="mm" value="12.1">12.1 mm</quantity>
.
</paragraph>
<paragraph id="8B8E1363FFBDFF9DFF2221274BFA5244" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF22212748B6527F" bold="true" box="[198,388,1327,1353]" pageId="3" pageNumber="31">Maximum size.</emphasis>
The largest of 750 captured and released animals was 
<quantity id="4CC9BE86FFBDFF9DFBE721384D61527C" box="[1027,1107,1328,1354]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="31" unit="cm" value="9.7">9.7 cm</quantity>
long (total length measured from the tip of the rostrum to end of the telson).
</paragraph>
</subSubSection>
<subSubSection id="C32B40E8FFBDFF98FF22217749FE5404" lastPageId="6" lastPageNumber="34" pageId="3" pageNumber="31" type="description">
<paragraph id="8B8E1363FFBDFF9DFF2221774B6952DC" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF222177481152AF" bold="true" box="[198,291,1407,1433]" pageId="3" pageNumber="31">Colour.</emphasis>
The colour varies according to size and substrate. Small animals are usually sand-coloured. Large animals are darker, ranging from brown to dark olive, sometimes almost purple. Fingertips of the first pereopod are red or orange in large animals.
</paragraph>
<paragraph id="8B8E1363FFBDFF9DFF2221FF4C335127" blockId="3.[151,1437,767,2003]" box="[198,1281,1527,1553]" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF2221FF4C335127" bold="true" box="[198,1281,1527,1553]" pageId="3" pageNumber="31">
Morphometric Comparison of 
<taxonomicName id="4C3168E0FFBDFF9DFDA321FF4AB55127" box="[583,903,1527,1553]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBDFF9DFDA321FF4AB55127" bold="true" box="[583,903,1527,1553]" italics="true" pageId="3" pageNumber="31">P. brasiliensis promatensis</emphasis>
</taxonomicName>
and 
<emphasis id="B945CF71FFBDFF9DFC2021FF4C335127" bold="true" box="[964,1281,1527,1553]" italics="true" pageId="3" pageNumber="31">
<taxonomicName id="4C3168E0FFBDFF9DFC2021FF4DCF5127" box="[964,1277,1527,1553]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">P. brasiliensis brasiliensis</taxonomicName>
.
</emphasis>
</emphasis>
</paragraph>
<paragraph id="8B8E1363FFBDFF9DFF2222284C0351BA" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
Mean residuals and residual standard deviations derived from the adjusted power regressions (y=a.Carapace_Length b) are presented at Table 4. Comparison of mean residuals (ANOVA) between 
<taxonomicName id="4C3168E0FFBDFF9DFAD52243485D51BA" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBDFF9DFAD52243485D51BA" italics="true" pageId="3" pageNumber="31">P. brasiliensis promatensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3168E0FFBDFF9DFE4C227B4BE851BA" box="[424,730,1651,1676]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBDFF9DFE4C227B4BE851BA" box="[424,730,1651,1676]" italics="true" pageId="3" pageNumber="31">P. brasiliensis brasiliensis</emphasis>
</taxonomicName>
resulted significant for 11 different measurements.
</paragraph>
<paragraph id="8B8E1363FFBDFF9DFF2222924851504A" blockId="3.[151,1437,767,2003]" pageId="3" pageNumber="31">
Discriminant analysis was applied to compare both species. All variance was explained by function 1 (P&lt;0.001), with a Canonical Correlation of 0.948 and perfect (100%) membership identification. Centroid values for 
<taxonomicName id="4C3168E0FFBDFF9DFEF622E34B785032" box="[274,586,1771,1796]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBDFF9DFEF622E34B785032" box="[274,586,1771,1796]" italics="true" pageId="3" pageNumber="31">P. brasiliensis promatensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3168E0FFBDFF9DFD6722E34A865032" box="[643,948,1771,1796]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBDFF9DFD6722E34A865032" box="[643,948,1771,1796]" italics="true" pageId="3" pageNumber="31">P. brasiliensis brasiliensis</emphasis>
</taxonomicName>
was 3.04 and -2.78 respectively. 
<figureCitation id="130A0FE6FFBDFF9DFADE22E24CAE5032" box="[1338,1436,1770,1796]" captionStart="FIGURE 2" captionStartId="5.[151,255,1110,1134]" captionTargetBox="[351,1236,482,1080]" captionTargetId="figure@5.[342,1245,474,1086]" captionTargetPageId="5" captionText="FIGURE 2. Frequency distribution of the Discriminant scores for individuals of Parastacus brasiliensis promatensis subsp. n. and Parastacus brasiliensis brasiliensis." httpUri="https://zenodo.org/record/183371/files/figure.png" pageId="3" pageNumber="31">Figure 2</figureCitation>
shows frequency distribution of the Discriminant scores, showing that each animal could be perfectly identified through morphometrical data. Discriminant scores are also no affected by animal size, as could be seen through 
<figureCitation id="130A0FE6FFBDFF9DFF1D236A486C504A" box="[249,350,1890,1916]" captionStart="FIGURE 3" captionStartId="5.[151,255,1854,1878]" captionTargetBox="[310,1276,1227,1826]" captionTargetId="figure@5.[307,1281,1222,1830]" captionTargetPageId="5" captionText="FIGURE 3. Discriminant scores for individuals of Parastacus brasiliensis promatensis subsp. n. and Parastacus brasiliensis brasiliensis as a function of the carapace length." httpUri="https://zenodo.org/record/183372/files/figure.png" pageId="3" pageNumber="31">Figure 3</figureCitation>
.
</paragraph>
<paragraph id="8B8E1363FFBDFF9AFF2223854DBE5634" blockId="3.[151,1437,767,2003]" lastBlockId="4.[151,1436,152,258]" lastPageId="4" lastPageNumber="32" pageId="3" pageNumber="31">
<emphasis id="B945CF71FFBDFF9DFF222385481F5091" bold="true" box="[198,301,1933,1959]" pageId="3" pageNumber="31">Habitat.</emphasis>
Garapiá stream drains a small basin with a thin acid soil layer over basalt rocks, covered by grass, natural and 
<emphasis id="B945CF71FFBDFF9DFE9423B148F350E4" box="[368,449,1977,2002]" italics="true" pageId="3" pageNumber="31">Pinnus</emphasis>
forests. Basin area is about 
<quantity id="4CC9BE86FFBDFF9DFCF723B14A7D50E5" box="[787,847,1977,2003]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="31" unit="km" value="3.0">3 km</quantity>
2 through an altitude of 
<quantity id="4CC9BE86FFBDFF9DFB8E23B14D9E50E5" box="[1130,1196,1977,2003]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="31" unit="m" value="850.0">850m</quantity>
before a high water fall to the coastal plains. The species was not found in lower areas of the same basin. The width of the stream ranges from 
<quantity id="4CC9BE86FFBAFF9AFECB24C8484657EC" box="[303,372,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="32" unit="m" value="0.5">0.5 m</quantity>
to about 
<quantity id="4CC9BE86FFBAFF9AFE0324C84B2B57EC" box="[487,537,192,218]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="32" unit="m" value="3.0">3 m</quantity>
, with a maximum depth of 
<quantity id="4CC9BE86FFBAFF9AFC8224C84AE957EC" box="[870,987,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="4" pageNumber="32" unit="cm" value="75.0" valueMax="80.0" valueMin="70.0">70-80 cm</quantity>
. The substrate varies from fine silt to sand and small rocks. Animals are found in hollows in the stream bank during the day.
</paragraph>
<paragraph id="8B8E1363FFBAFF9AFF7325324843568F" blockId="4.[151,1436,312,441]" pageId="4" pageNumber="32">
<emphasis id="B945CF71FFBAFF9AFF732532483E5664" bold="true" box="[151,268,314,338]" pageId="4" pageNumber="32">TABLE 4.</emphasis>
Mean residuals from power regressions (y=a.Carapace_Length b) of pooled data of 
<taxonomicName id="4C3168E0FFBAFF9AFB442532482A5642" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBAFF9AFB442532482A5642" italics="true" pageId="4" pageNumber="32">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3168E0FFBAFF9AFEAA25554BF95642" box="[334,715,349,372]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBAFF9AFEAA25554BF95642" box="[334,715,349,372]" italics="true" pageId="4" pageNumber="32">Parastacus brasiliensis brasiliensis</emphasis>
</taxonomicName>
. Regressions were performed using SPSS (11.0) non linear regression routine, applying Levenberg-Marquardt algorithm. Coefficients of the Canonical Discriminant function 1 are also presented (r=0.948).
</paragraph>
<paragraph id="8B8E1363FFBAFF9AFEC525D24CBD56C7" blockId="4.[159,1423,474,609]" box="[289,1423,474,497]" pageId="4" pageNumber="32">
<emphasis id="B945CF71FFBAFF9AFEC525D24A6656C7" box="[289,852,474,497]" italics="true" pageId="4" pageNumber="32">
<taxonomicName id="4C3168E0FFBAFF9AFEC525D2488556C7" box="[289,439,474,497]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="species" species="brasiliensis">P. brasiliensis</taxonomicName>
promaten- 
<taxonomicName id="4C3168E0FFBAFF9AFDD925D24A6656C7" authority="Significance" authorityName="Significance" box="[573,852,474,497]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">P. brasiliensis brasiliensis</taxonomicName>
</emphasis>
Significance for Coefficients of the Canonical
</paragraph>
<paragraph id="8B8E1363FFBAFF9AFEC525F54C695522" blockId="4.[159,1423,474,609]" box="[289,1371,509,532]" pageId="4" pageNumber="32">
<emphasis id="B945CF71FFBAFF9AFEC525F5480F5522" box="[289,317,509,532]" italics="true" pageId="4" pageNumber="32">sis</emphasis>
comparing means Discriminant Function 1
</paragraph>
<paragraph id="8B8E1363FFBAFF9AFC8D26284AE55501" blockId="4.[159,1423,474,609]" box="[873,983,544,567]" pageId="4" pageNumber="32">(ANOVA)</paragraph>
<paragraph id="8B8E1363FFBAFF9AFF7B26424A785557" blockId="4.[159,1423,474,609]" box="[159,842,586,609]" pageId="4" pageNumber="32">Character Mean Std. Deviation Mean Std. Deviation</paragraph>
<paragraph id="8B8E1363FFBAFF9AFF7B267D482951C2" pageId="4" pageNumber="32">
<table id="F931E1C3FFBA0061FF73267D4CBD51C2" box="[151,1423,629,1780]" gridcols="4" gridrows="17" pageId="4" pageNumber="32">
<tr id="35011121FFBA0061FF73267D4CBD55BA" box="[151,1423,629,652]" gridrow="0" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73267D48DB55BA" box="[151,489,629,652]" gridcol="0" gridrow="0" pageId="4" pageNumber="32">HCHE -0.01638 0.15623</th>
<th id="76D0785DFFBA0061FDD9267D4A3A55BA" box="[573,776,629,652]" gridcol="1" gridrow="0" pageId="4" pageNumber="32">0.02266 0.16803</th>
<th id="76D0785DFFBA0061FC8D267D4AF255BA" box="[873,960,629,652]" gridcol="2" gridrow="0" pageId="4" pageNumber="32">0.43060</th>
<th id="76D0785DFFBA0061FBB3267D4CBD55BA" box="[1111,1423,629,652]" gridcol="3" gridrow="0" pageId="4" pageNumber="32">3.18313</th>
</tr>
<tr id="35011121FFBA0061FF7326A84CBD55D7" box="[151,1423,672,737]" gridrow="1" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7326A848DB55D7" box="[151,489,672,737]" gridcol="0" gridrow="1" pageId="4" pageNumber="32">HPRO5 -0.01293 0.07031 LABD 0.02278 0.03526</th>
<td id="76D0785DFFBA0061FDD926A84A3A55D7" box="[573,776,672,737]" gridcol="1" gridrow="1" pageId="4" pageNumber="32">0.00739 0.10926 -0.02387 0.04456</td>
<td id="76D0785DFFBA0061FC8D26A84AF255D7" box="[873,960,672,737]" gridcol="2" gridrow="1" pageId="4" pageNumber="32">0.47216 0.00043</td>
<td id="76D0785DFFBA0061FBB326A84CBD55D7" box="[1111,1423,672,737]" gridcol="3" gridrow="1" pageId="4" pageNumber="32">-1.31597 11.44408</td>
</tr>
<tr id="35011121FFBA0061FF7326FD4CBD543A" box="[151,1423,757,780]" gridrow="2" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7326FD48DB543A" box="[151,489,757,780]" gridcol="0" gridrow="2" pageId="4" pageNumber="32">LACAR 0.00085 0.03247</th>
<td id="76D0785DFFBA0061FDD926FD4A3A543A" box="[573,776,757,780]" gridcol="1" gridrow="2" pageId="4" pageNumber="32">-0.00077 0.01440</td>
<td id="76D0785DFFBA0061FC8D26FD4AF2543A" box="[873,960,757,780]" gridcol="2" gridrow="2" pageId="4" pageNumber="32">0.82937</td>
<td id="76D0785DFFBA0061FBB326FD4CBD543A" box="[1111,1423,757,780]" gridcol="3" gridrow="2" pageId="4" pageNumber="32">-35.58309</td>
</tr>
<tr id="35011121FFBA0061FF7327284CBD5457" box="[151,1423,800,865]" gridrow="3" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73272848DB5457" box="[151,489,800,865]" gridcol="0" gridrow="3" pageId="4" pageNumber="32">LCAR5 -0.03060 0.11806 LCHE 0.00852 0.11491</th>
<td id="76D0785DFFBA0061FDD927284A3A5457" box="[573,776,800,865]" gridcol="1" gridrow="3" pageId="4" pageNumber="32">0.03315 0.11929 -0.00363 0.14961</td>
<td id="76D0785DFFBA0061FC8D27284AF25457" box="[873,960,800,865]" gridcol="2" gridrow="3" pageId="4" pageNumber="32">0.08244 0.76572</td>
<td id="76D0785DFFBA0061FBB327284CBD5457" box="[1111,1423,800,865]" gridcol="3" gridrow="3" pageId="4" pageNumber="32">-2.01322 -8.31548</td>
</tr>
<tr id="35011121FFBA0061FF73277D4CBD54BA" box="[151,1423,885,908]" gridrow="4" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73277D48DB54BA" box="[151,489,885,908]" gridcol="0" gridrow="4" pageId="4" pageNumber="32">LDAC 0.01672 0.10657</th>
<td id="76D0785DFFBA0061FDD9277D4A3A54BA" box="[573,776,885,908]" gridcol="1" gridrow="4" pageId="4" pageNumber="32">-0.00887 0.12076</td>
<td id="76D0785DFFBA0061FC8D277D4AF254BA" box="[873,960,885,908]" gridcol="2" gridrow="4" pageId="4" pageNumber="32">0.46208</td>
<td id="76D0785DFFBA0061FBB3277D4CBD54BA" box="[1111,1423,885,908]" gridcol="3" gridrow="4" pageId="4" pageNumber="32">12.15535</td>
</tr>
<tr id="35011121FFBA0061FF7327A84CBD54D7" box="[151,1423,928,993]" gridrow="5" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7327A848DB54D7" box="[151,489,928,993]" gridcol="0" gridrow="5" pageId="4" pageNumber="32">LDAC5 0.02801 0.10222 LMER5 -0.01828 0.07660</th>
<td id="76D0785DFFBA0061FDD927A84A3A54D7" box="[573,776,928,993]" gridcol="1" gridrow="5" pageId="4" pageNumber="32">-0.02654 0.09230 0.01787 0.08211</td>
<td id="76D0785DFFBA0061FC8D27A84AF254D7" box="[873,960,928,993]" gridcol="2" gridrow="5" pageId="4" pageNumber="32">0.06981 0.13956</td>
<td id="76D0785DFFBA0061FBB327A84CBD54D7" box="[1111,1423,928,993]" gridcol="3" gridrow="5" pageId="4" pageNumber="32">-0.53563 -5.49672</td>
</tr>
<tr id="35011121FFBA0061FF7327FD4CBD533A" box="[151,1423,1013,1036]" gridrow="6" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7327FD48DB533A" box="[151,489,1013,1036]" gridcol="0" gridrow="6" pageId="4" pageNumber="32">LPCAR -0.00105 0.03757</th>
<td id="76D0785DFFBA0061FDD927FD4A3A533A" box="[573,776,1013,1036]" gridcol="1" gridrow="6" pageId="4" pageNumber="32">-0.00024 0.07585</td>
<td id="76D0785DFFBA0061FC8D27FD4AF2533A" box="[873,960,1013,1036]" gridcol="2" gridrow="6" pageId="4" pageNumber="32">0.96491</td>
<td id="76D0785DFFBA0061FBB327FD4CBD533A" box="[1111,1423,1013,1036]" gridcol="3" gridrow="6" pageId="4" pageNumber="32">1.49908</td>
</tr>
<tr id="35011121FFBA0061FF7320284CBD5357" box="[151,1423,1056,1121]" gridrow="7" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73202848DB5357" box="[151,489,1056,1121]" gridcol="0" gridrow="7" pageId="4" pageNumber="32">LPRO5 0.02896 0.04731 LROS 0.05558 0.05321</th>
<td id="76D0785DFFBA0061FDD920284A3A5357" box="[573,776,1056,1121]" gridcol="1" gridrow="7" pageId="4" pageNumber="32">-0.03079 0.07761 -0.05122 0.10055</td>
<td id="76D0785DFFBA0061FC8D20284AF25357" box="[873,960,1056,1121]" gridcol="2" gridrow="7" pageId="4" pageNumber="32">0.00399 0.00009</td>
<td id="76D0785DFFBA0061FBB320284CBD5357" box="[1111,1423,1056,1121]" gridcol="3" gridrow="7" pageId="4" pageNumber="32">12.47622 8.57633</td>
</tr>
<tr id="35011121FFBA0061FF73207D4CBD53BA" box="[151,1423,1141,1164]" gridrow="8" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73207D48DB53BA" box="[151,489,1141,1164]" gridcol="0" gridrow="8" pageId="4" pageNumber="32">LSTEL 0.04448 0.04459</th>
<td id="76D0785DFFBA0061FDD9207D4A3A53BA" box="[573,776,1141,1164]" gridcol="1" gridrow="8" pageId="4" pageNumber="32">-0.03912 0.04313</td>
<td id="76D0785DFFBA0061FC8D207D4AF253BA" box="[873,960,1141,1164]" gridcol="2" gridrow="8" pageId="4" pageNumber="32">0.00000</td>
<td id="76D0785DFFBA0061FBB3207D4CBD53BA" box="[1111,1423,1141,1164]" gridcol="3" gridrow="8" pageId="4" pageNumber="32">21.80923</td>
</tr>
<tr id="35011121FFBA0061FF7320A84CBD53D7" box="[151,1423,1184,1249]" gridrow="9" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7320A848DB53D7" box="[151,489,1184,1249]" gridcol="0" gridrow="9" pageId="4" pageNumber="32">LTEL 0.04269 0.03550 LURO1 0.01185 0.06364</th>
<td id="76D0785DFFBA0061FDD920A84A3A53D7" box="[573,776,1184,1249]" gridcol="1" gridrow="9" pageId="4" pageNumber="32">-0.04025 0.04229 -0.00673 0.06116</td>
<td id="76D0785DFFBA0061FC8D20A84AF253D7" box="[873,960,1184,1249]" gridcol="2" gridrow="9" pageId="4" pageNumber="32">0.00000 0.32905</td>
<td id="76D0785DFFBA0061FBB320A84CBD53D7" box="[1111,1423,1184,1249]" gridcol="3" gridrow="9" pageId="4" pageNumber="32">11.45896 8.83343</td>
</tr>
<tr id="35011121FFBA0061FF7320FD4CBD523A" box="[151,1423,1269,1292]" gridrow="10" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7320FD48DB523A" box="[151,489,1269,1292]" gridcol="0" gridrow="10" pageId="4" pageNumber="32">LURO2 0.01878 0.04321</th>
<td id="76D0785DFFBA0061FDD920FD4A3A523A" box="[573,776,1269,1292]" gridcol="1" gridrow="10" pageId="4" pageNumber="32">-0.01148 0.05277</td>
<td id="76D0785DFFBA0061FC8D20FD4AF2523A" box="[873,960,1269,1292]" gridcol="2" gridrow="10" pageId="4" pageNumber="32">0.04472</td>
<td id="76D0785DFFBA0061FBB320FD4CBD523A" box="[1111,1423,1269,1292]" gridcol="3" gridrow="10" pageId="4" pageNumber="32">-12.50582</td>
</tr>
<tr id="35011121FFBA0061FF7321284CBD5257" box="[151,1423,1312,1377]" gridrow="11" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73212848DB5257" box="[151,489,1312,1377]" gridcol="0" gridrow="11" pageId="4" pageNumber="32">TCHE -0.02001 0.18231 WCAR 0.00776 0.03775</th>
<td id="76D0785DFFBA0061FDD921284A3A5257" box="[573,776,1312,1377]" gridcol="1" gridrow="11" pageId="4" pageNumber="32">0.02447 0.19067 -0.00513 0.04852</td>
<td id="76D0785DFFBA0061FC8D21284AF25257" box="[873,960,1312,1377]" gridcol="2" gridrow="11" pageId="4" pageNumber="32">0.43446 0.33404</td>
<td id="76D0785DFFBA0061FBB321284CBD5257" box="[1111,1423,1312,1377]" gridcol="3" gridrow="11" pageId="4" pageNumber="32">-4.15076 7.09544</td>
</tr>
<tr id="35011121FFBA0061FF73217D4CBD52BA" box="[151,1423,1397,1420]" gridrow="12" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73217D48DB52BA" box="[151,489,1397,1420]" gridcol="0" gridrow="12" pageId="4" pageNumber="32">WROS 0.03735 0.05910</th>
<td id="76D0785DFFBA0061FDD9217D4A3A52BA" box="[573,776,1397,1420]" gridcol="1" gridrow="12" pageId="4" pageNumber="32">-0.03384 0.08647</td>
<td id="76D0785DFFBA0061FC8D217D4AF252BA" box="[873,960,1397,1420]" gridcol="2" gridrow="12" pageId="4" pageNumber="32">0.00294</td>
<td id="76D0785DFFBA0061FBB3217D4CBD52BA" box="[1111,1423,1397,1420]" gridcol="3" gridrow="12" pageId="4" pageNumber="32">-0.05134</td>
</tr>
<tr id="35011121FFBA0061FF7321A84CBD52D7" box="[151,1423,1440,1505]" gridrow="13" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7321A848DB52D7" box="[151,489,1440,1505]" gridcol="0" gridrow="13" pageId="4" pageNumber="32">WTEL 0.04248 0.04534 WTES 0.05249 0.08400</th>
<td id="76D0785DFFBA0061FDD921A84A3A52D7" box="[573,776,1440,1505]" gridcol="1" gridrow="13" pageId="4" pageNumber="32">-0.03411 0.04396 -0.05430 0.15678</td>
<td id="76D0785DFFBA0061FC8D21A84AF252D7" box="[873,960,1440,1505]" gridcol="2" gridrow="13" pageId="4" pageNumber="32">0.00000 0.00817</td>
<td id="76D0785DFFBA0061FBB321A84CBD52D7" box="[1111,1423,1440,1505]" gridcol="3" gridrow="13" pageId="4" pageNumber="32">13.29373 -1.99503</td>
</tr>
<tr id="35011121FFBA0061FF7321FD4CBD513A" box="[151,1423,1525,1548]" gridrow="14" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF7321FD48DB513A" box="[151,489,1525,1548]" gridcol="0" gridrow="14" pageId="4" pageNumber="32">WURO1 0.00234 0.04338</th>
<td id="76D0785DFFBA0061FDD921FD4A3A513A" box="[573,776,1525,1548]" gridcol="1" gridrow="14" pageId="4" pageNumber="32">-0.00212 0.05863</td>
<td id="76D0785DFFBA0061FC8D21FD4AF2513A" box="[873,960,1525,1548]" gridcol="2" gridrow="14" pageId="4" pageNumber="32">0.77738</td>
<td id="76D0785DFFBA0061FBB321FD4CBD513A" box="[1111,1423,1525,1548]" gridcol="3" gridrow="14" pageId="4" pageNumber="32">-23.42223</td>
</tr>
<tr id="35011121FFBA0061FF7322284CBD5157" box="[151,1423,1568,1633]" gridrow="15" pageId="4" pageNumber="32">
<th id="76D0785DFFBA0061FF73222848DB5157" box="[151,489,1568,1633]" gridcol="0" gridrow="15" pageId="4" pageNumber="32">WURO2 0.01424 0.06554 Constant</th>
<td id="76D0785DFFBA0061FDD922284A3A5157" box="[573,776,1568,1633]" gridcol="1" gridrow="15" pageId="4" pageNumber="32">-0.01407 0.06075</td>
<td id="76D0785DFFBA0061FC8D22284AF25157" box="[873,960,1568,1633]" gridcol="2" gridrow="15" pageId="4" pageNumber="32">0.14453</td>
<td id="76D0785DFFBA0061FBB322284CBD5157" box="[1111,1423,1568,1633]" gridcol="3" gridrow="15" pageId="4" pageNumber="32">-8.97476 -0.00886</td>
</tr>
<tr id="35011121FFBA0061FF7322D24CBD51C2" box="[151,1423,1754,1780]" gridrow="16" pageId="4" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76D0785DFFBA0061FF7322D248DB51C2" box="[151,489,1754,1780]" gridcol="0" gridrow="16" pageId="4" pageNumber="32">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="8B8E1363FFBAFF9AFF73232D4DEA50B9" blockId="4.[151,1436,1829,2015]" pageId="4" pageNumber="32">
Using the identification keys provided by 
<bibRefCitation id="EFA06E92FFBAFF9AFD6A232D4A8F5009" author="Buckup" box="[654,957,1829,1855]" pageId="4" pageNumber="34" refString="Buckup, L. &amp; Rossi, A. (1980) O genero Parastacus no Brasil (Crustacea, Decapoda, Parastacidae). Revista Brasileira de Biologia, 40, 663 - 681." type="journal article" year="1980">Buckup and Rossi (1980)</bibRefCitation>
and 
<bibRefCitation id="EFA06E92FFBAFF9AFC1E232D4D9D5009" author="Buckup" box="[1018,1199,1829,1855]" pageId="4" pageNumber="34" refString="Buckup, L. (1999) Familia Parastacidae. In: Buckup, L. &amp; Bond-Buckup, G. (Ed.), Os Crustaceos do Rio Grande do Sul. Editora da UFRGS, Porto Alegre, p. 319 - 327." type="book chapter" year="1999">Buckup (1999)</bibRefCitation>
, 
<taxonomicName id="4C3168E0FFBAFF9AFB58232D49CC5050" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBAFF9AFB58232D49CC5050" italics="true" pageId="4" pageNumber="32">P. brasiliensis promatensis</emphasis>
</taxonomicName>
is identified as 
<taxonomicName id="4C3168E0FFBAFF9AFE2023454B595050" box="[452,619,1869,1894]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B945CF71FFBAFF9AFE2023454B595050" box="[452,619,1869,1894]" italics="true" pageId="4" pageNumber="32">P. brasiliensis</emphasis>
</taxonomicName>
. However, Discriminant analysis (Table 4, 
<figureCitation id="130A0FE6FFBAFF9AFB9E23454DF35051" box="[1146,1217,1869,1895]" captionStart="FIGURE 2" captionStartId="5.[151,255,1110,1134]" captionTargetBox="[351,1236,482,1080]" captionTargetId="figure@5.[342,1245,474,1086]" captionTargetPageId="5" captionText="FIGURE 2. Frequency distribution of the Discriminant scores for individuals of Parastacus brasiliensis promatensis subsp. n. and Parastacus brasiliensis brasiliensis." httpUri="https://zenodo.org/record/183371/files/figure.png" pageId="4" pageNumber="32">Fig. 2</figureCitation>
and 
<figureCitation id="130A0FE6FFBAFF9AFAE523454C225051" box="[1281,1296,1869,1895]" captionStart="FIGURE 3" captionStartId="5.[151,255,1854,1878]" captionTargetBox="[310,1276,1227,1826]" captionTargetId="figure@5.[307,1281,1222,1830]" captionTargetPageId="5" captionText="FIGURE 3. Discriminant scores for individuals of Parastacus brasiliensis promatensis subsp. n. and Parastacus brasiliensis brasiliensis as a function of the carapace length." httpUri="https://zenodo.org/record/183372/files/figure.png" pageId="4" pageNumber="32">3</figureCitation>
) are strong enough for a clear distinction between them, with perfect (100%) membership identification.
</paragraph>
<paragraph id="8B8E1363FFBAFF9BFF2123954B2A5694" blockId="4.[151,1436,1829,2015]" lastBlockId="5.[151,1436,152,418]" lastPageId="5" lastPageNumber="33" pageId="4" pageNumber="32">
The relationship between both subspecies is very intriguing. Between them there is a geographical (
<quantity id="4CC9BE86FFBAFF9AFAAA23954CA15081" box="[1358,1427,1949,1975]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="32" unit="km" value="60.0">60km</quantity>
) and altitudinal (
<quantity id="4CC9BE86FFBAFF9AFEAA23CD48A650E9" box="[334,404,1989,2015]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="32" unit="m" value="650.0">650m</quantity>
) gap with no registered occurrence of any Parastacid. 
<taxonomicName id="4C3168E0FFBAFF9BFBF423CD49845787" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" lastPageId="5" lastPageNumber="33" order="Decapoda" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBAFF9BFBF423CD49845787" italics="true" lastPageId="5" lastPageNumber="33" pageId="4" pageNumber="32">Parastacus brasiliensis brasiliensis</emphasis>
</taxonomicName>
is found in the central depression of Rio Grande do Sul (Jacuí Drainage), at altitudes of no more then 
<quantity id="4CC9BE86FFBBFF9BFABD24904CAE5784" box="[1369,1436,152,178]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="33" unit="m" value="200.0">200m</quantity>
(estimated), characterized by sandstone sediments from Triassic. By the other hand, 
<taxonomicName id="4C3168E0FFBBFF9BFB9324C849845637" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFBBFF9BFB9324C849845637" italics="true" pageId="5" pageNumber="33">P. brasiliensis promatensis</emphasis>
</taxonomicName>
is found 
<quantity id="4CC9BE86FFBBFF9BFEC224E0485B5634" box="[294,361,232,258]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="33" unit="m" value="850.0">850m</quantity>
high (Litoral Drainage), on a basalt basin formed during the Gondwana breakdown (lower Cretaceous). Of course, both subspecies are not so old. Cretaceous geological events may be related to vicariant process related to South American parastacoid genus formation, as reported by 
<bibRefCitation id="EFA06E92FFBBFF9BFB6A25304CAA5664" box="[1166,1432,312,338]" pageId="5" pageNumber="34" refString="Crandall, K. A., Fetzer J. W., Jara, C. G. &amp; Buckup, L. (2000) On the phylogenetic positioning of the freshwater crayfish genera (Decapoda: Parastacidae). Journal of Crustacean Biology, 20, 530 - 540." type="journal article">
Crandall 
<emphasis id="B945CF71FFBBFF9BFB1925304C095667" box="[1277,1339,312,337]" italics="true" pageId="5" pageNumber="33">et al.</emphasis>
(2000)
</bibRefCitation>
. Also, no major geological event is known from Cretaceous and Pleistocene climatic changes apparently do not justify this geographical gap.
</paragraph>
<caption id="DF4E43EBFFBBFF9BFF73205E4B8353A6" httpUri="https://zenodo.org/record/183371/files/figure.png" pageId="5" pageNumber="33" targetBox="[351,1236,482,1080]" targetPageId="5">
<paragraph id="8B8E1363FFBBFF9BFF73205E4B8353A6" blockId="5.[151,1435,1110,1169]" pageId="5" pageNumber="33">
<emphasis id="B945CF71FFBBFF9BFF73205E48295358" bold="true" box="[151,283,1110,1134]" pageId="5" pageNumber="33">FIGURE 2.</emphasis>
Frequency distribution of the Discriminant scores for individuals of 
<taxonomicName id="4C3168E0FFBBFF9BFBF1205F4CA95358" box="[1045,1435,1111,1134]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBBFF9BFBF1205F4CA95358" box="[1045,1435,1111,1134]" italics="true" pageId="5" pageNumber="33">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
<emphasis id="B945CF71FFBBFF9BFF73207149C953A7" bold="true" box="[151,251,1145,1169]" pageId="5" pageNumber="33">
<taxonomicNameLabel id="A276720AFFBBFF9BFF73207149C953A7" box="[151,251,1145,1169]" pageId="5" pageNumber="33" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C3168E0FFBBFF9BFECA20714B9B53A6" box="[302,681,1145,1168]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBBFF9BFECA20714B9B53A6" box="[302,681,1145,1168]" italics="true" pageId="5" pageNumber="33">Parastacus brasiliensis brasiliensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF4E43EBFFBBFF9BFF7323364BEF504E" httpUri="https://zenodo.org/record/183372/files/figure.png" pageId="5" pageNumber="33" targetBox="[310,1276,1227,1826]" targetPageId="5">
<paragraph id="8B8E1363FFBBFF9BFF7323364BEF504E" blockId="5.[151,1436,1854,1913]" pageId="5" pageNumber="33">
<emphasis id="B945CF71FFBBFF9BFF732336482A5060" bold="true" box="[151,280,1854,1878]" pageId="5" pageNumber="33">FIGURE 3.</emphasis>
Discriminant scores for individuals of 
<taxonomicName id="4C3168E0FFBBFF9BFD5C23374D0B5060" box="[696,1081,1855,1878]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" status="subsp. nov." subSpecies="promatensis">
<emphasis id="B945CF71FFBBFF9BFD5C23374D0B5060" box="[696,1081,1855,1878]" italics="true" pageId="5" pageNumber="33">Parastacus brasiliensis promatensis</emphasis>
</taxonomicName>
<emphasis id="B945CF71FFBBFF9BFBA423364D915060" bold="true" box="[1088,1187,1854,1878]" pageId="5" pageNumber="33">
<taxonomicNameLabel id="A276720AFFBBFF9BFBA423364D915060" box="[1088,1187,1854,1878]" pageId="5" pageNumber="33" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C3168E0FFBBFF9BFB3223374867504F" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFBBFF9BFB3223374867504F" italics="true" pageId="5" pageNumber="33">Parastacus brasiliensis brasiliensis</emphasis>
</taxonomicName>
as a function of the carapace length.
</paragraph>
</caption>
<paragraph id="8B8E1363FFBBFF98FF2123A448245634" blockId="5.[151,1436,1964,2030]" lastBlockId="6.[151,1437,152,818]" lastPageId="6" lastPageNumber="34" pageId="5" pageNumber="33">
Parastacids are known to migrate outside water and to resist desiccation (
<bibRefCitation id="EFA06E92FFBBFF9BFBC323A44C7650F0" author="Hughes" box="[1063,1348,1964,1990]" pageId="5" pageNumber="34" refString="Hughes J. M. &amp; Hillyer M. (2003) Patterns of connectivity among populations of Cherax destructor (Decapoda: Parastacidae) in Western Queensland, Australia. Journal of Marine and Freshwater Research, 54, 1 - 10." type="journal article" year="2003">Hughes &amp; Hillyer 2003</bibRefCitation>
). Some years ago, an individual of 
<taxonomicName id="4C3168E0FFBBFF9BFE3023DC4BC850DB" box="[468,762,2004,2029]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasilienis">
<emphasis id="B945CF71FFBBFF9BFE3023DC4BC850DB" box="[468,762,2004,2029]" italics="true" pageId="5" pageNumber="33">P. brasiliensis brasilienis</emphasis>
</taxonomicName>
escaped from an aquarium in our lab and was found alive some (2-3) days before. The animal was so dehydrated that when returned to the water was unable to sink. Nevertheless, confined inside a hollow under water, in just some hours the animal was behaving as a healthy individual.
</paragraph>
<paragraph id="8B8E1363FFB8FF98FF2125184D595694" blockId="6.[151,1437,152,818]" pageId="6" pageNumber="34">
Parastacids can also move large distances using constructed or natural passages under the ground. We personally known a case of a very shallow (1,2m deep) water well constructed on a hill (
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) in sandstone sediments. The nearest creek was at least 
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distant and 
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bellow. After one year, the well was opened for cleaning and there were crayfishes (
<taxonomicName id="4C3168E0FFB8FF98FD2625804AF35697" box="[706,961,392,417]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="34" phylum="Arthropoda" rank="species" species="pilimanus">
<emphasis id="B945CF71FFB8FF98FD2625804AF35697" box="[706,961,392,417]" italics="true" pageId="6" pageNumber="34">Parastacus pilimanus</emphasis>
</taxonomicName>
) living inside.
</paragraph>
<paragraph id="8B8E1363FFB8FF98FF2125B84DC25574" blockId="6.[151,1437,152,818]" pageId="6" pageNumber="34">
<bibRefCitation id="EFA06E92FFB8FF98FF2125B848DD56FC" author="Hughes" box="[197,495,432,458]" pageId="6" pageNumber="34" refString="Hughes J. M. &amp; Hillyer M. (2003) Patterns of connectivity among populations of Cherax destructor (Decapoda: Parastacidae) in Western Queensland, Australia. Journal of Marine and Freshwater Research, 54, 1 - 10." type="journal article" year="2003">Hughes &amp; Hillyer (2003)</bibRefCitation>
analysed, using molecular data, the patterns of connectivity among populations of 
<taxonomicName id="4C3168E0FFB8FF98FF5325D04B0956C4" box="[183,571,472,498]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="34" phylum="Arthropoda" rank="species" species="destructor">
<emphasis id="B945CF71FFB8FF98FF5325D048BF56C7" box="[183,397,472,497]" italics="true" pageId="6" pageNumber="34">Cherax destructor</emphasis>
(Parastacidae)
</taxonomicName>
in Western Queensland, 
<collectingCountry id="F32653F3FFB8FF98FC8525D04AE256C4" box="[865,976,472,498]" name="Australia" pageId="6" pageNumber="34">Australia</collectingCountry>
. Although this species is able to terrestrial migration, these authors found almost no sharing of haplotypes across drainage boundaries, indicating limited terrestrial dispersal across them. Similar results were obtained by 
<bibRefCitation id="EFA06E92FFB8FF98FC1026204DDF5574" box="[1012,1261,552,578]" pageId="6" pageNumber="34" refString="Nguyen, T. T. T., Burridge, C. P. &amp; Austin C. M. (2005) Population genetic studies on the Australian freshwater crayfish, Cherax destructor (Crustacea: Parastacidae) using allozyme and RAPD markers. Aquatic Living Resources, 18, 55 - 64." type="journal article">
Nguyen e 
<emphasis id="B945CF71FFB8FF98FB8126204DA15577" box="[1125,1171,552,577]" italics="true" pageId="6" pageNumber="34">t al.</emphasis>
(2005)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B8E1363FFB8FF98FF2126584B01558C" blockId="6.[151,1437,152,818]" pageId="6" pageNumber="34">
Crayfish dispersal between drainages was reported by 
<bibRefCitation id="EFA06E92FFB8FF98FCAB26584D40555C" author="Horwitz" box="[847,1138,592,618]" pageId="6" pageNumber="34" refString="Horwitz P. &amp; Knott B. (1995) The distribution and spread of the yabby Cherax complex in Australia: Speculations, hypotheses and the need for research. Freshwater Crayfish 10, 81 - 91." type="journal article" year="1995">Horwitz &amp; Knott (1995)</bibRefCitation>
as food items frequently translocated by Australian aborigines. Amerinds also use crustaceans as regular food items, although with no record of 
<taxonomicName id="4C3168E0FFB8FF98FEED26A848BF558F" box="[265,397,672,697]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B945CF71FFB8FF98FEED26A848BF558F" box="[265,397,672,697]" italics="true" pageId="6" pageNumber="34">Parastacus</emphasis>
</taxonomicName>
consumption.
</paragraph>
<paragraph id="8B8E1363FFB8FF98FF2126C049FE5404" blockId="6.[151,1437,152,818]" pageId="6" pageNumber="34">
So, the distribution of 
<taxonomicName id="4C3168E0FFB8FF98FE2926C04A3255D7" box="[461,768,712,737]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="brasiliensis">
<emphasis id="B945CF71FFB8FF98FE2926C04A3255D7" box="[461,768,712,737]" italics="true" pageId="6" pageNumber="34">P. brasiliensis brasiliensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3168E0FFB8FF98FCDD26C04D4055D7" box="[825,1138,712,737]" class="Malacostraca" family="Parastacidae" genus="Parastacus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="brasiliensis" subSpecies="promatensis">
<emphasis id="B945CF71FFB8FF98FCDD26C04D4055D7" box="[825,1138,712,737]" italics="true" pageId="6" pageNumber="34">P. brasiliensis promatensis</emphasis>
</taxonomicName>
so far apart is very interesting and maybe a molecular approach will be necessary to solve this biogeographical and evolutive question.
</paragraph>
</subSubSection>
</treatment>
</document>