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<document id="174DA0BB38DA267577A13F84328D2CA1" ID-CLB-Dataset="8626" ID-DOI="10.5281/zenodo.4476396" ID-GBIF-Dataset="f638455f-2539-4bb0-816f-bafbc0b7c1ac" ID-ISSN="ISSN 0374-1036" ID-Zenodo-Dep="4476396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1611663472831" checkinUser="felipe" docAuthor="Kment, Petr" docDate="2005" docId="039887C4A30DF4657D89793A354A56E2" docLanguage="en" docName="ActaEntMusNatPra.45.21-50.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosomatidae Signoret 1863" docType="key" docVersion="6" lastPageNumber="41" masterDocId="FFA1FFBCA31EF4717D327C42312D533A" masterDocTitle="Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles" masterLastPageNumber="50" masterPageNumber="21" pageNumber="40" updateTime="1698925359088" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2F92972FC1ECF0111D145C28EDF6D501">Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles</mods:title>
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<mods:namePart id="89909F5A77954A8C76F1741E0CB25651">Kment, Petr</mods:namePart>
<mods:affiliation id="8A461DB9446D7E5D006AC8C59104B3FC">) &amp;) Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: sigara @ post. cz &amp;) &amp;) Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:title id="E3EEDB54408679D6BA2735410927BD0A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="086E70E522ADD1D704CF91FED58A972D">2005</mods:date>
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<mods:title id="4A71548497018BB3763E275C793DE5D3">suppl.</mods:title>
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<mods:number id="EE35E49247591F2CE6BF24D69BF465DD">2005-12-31</mods:number>
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<treatment id="039887C4A30DF4657D89793A354A56E2" ID-DOI="http://doi.org/10.5281/zenodo.4476468" ID-GBIF-Taxon="177135171" ID-Zenodo-Dep="4476468" LSID="urn:lsid:plazi:treatment:039887C4A30DF4657D89793A354A56E2" httpUri="http://treatment.plazi.org/id/039887C4A30DF4657D89793A354A56E2" lastPageId="20" lastPageNumber="41" pageId="19" pageNumber="40">
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<paragraph id="8B8E36D2A30DF4627D89793A32DA56AB" blockId="19.[187,1015,1399,1426]" box="[187,1015,1399,1426]" pageId="19" pageNumber="40">
<heading id="D0C681BEA30DF4627D89793A32DA56AB" bold="true" box="[187,1015,1399,1426]" centered="true" fontSize="11" level="1" pageId="19" pageNumber="40" reason="2">
<emphasis id="B945EAC0A30DF4627D89793A32DA56AB" bold="true" box="[187,1015,1399,1426]" pageId="19" pageNumber="40">
Key to the
<taxonomicName id="4C314D51A30DF4627C7D7935331B56AB" ID-CoL="7NFSF" authorityName="Signoret" authorityYear="1863" box="[335,566,1399,1425]" class="Insecta" family="Acanthosomatidae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="40" phylum="Arthropoda" rank="family">Acanthosomatidae</taxonomicName>
of
<collectingCountry id="F3267642A30DF4627F54793A33D256A8" box="[614,767,1400,1426]" name="Madagascar" pageId="19" pageNumber="40">Madagascar</collectingCountry>
and the
<collectingCountry id="F3267642A30DF4627E4B793532DA56AB" box="[889,1015,1399,1425]" name="Seychelles" pageId="19" pageNumber="40">Seychelles</collectingCountry>
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<paragraph id="8B8E36D2A30DF4627D4379F3356C56C0" blockId="19.[113,1089,1457,1530]" pageId="19" pageNumber="40">
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1. Mesosternal carina not developed. Antennae 5-segmented, antennomere 2 not exceptionally long. Metathoracic scent gland complex as in
<figureCitation id="130A2A57A30DF4627C46798E309E56E5" box="[372,435,1484,1503]" captionStart="Figs" captionStartId="17.[113,152,1206,1225]" captionTargetBox="[113,1088,220,1175]" captionTargetId="figure-16@17.[113,1088,220,1175]" captionTargetPageId="17" captionText="Figs. 31-34. Pleura and metathoracic scent gland opening, most exposed view. 31 – Mahea sexualis Distant, 1909; 32 – Catadipson imernensis (Cachan, 1952); 33 – Noualhieridia rufa Cachan, 1952; 30 – Ibocoris ficivora Roche, 1947." figureDoi="http://doi.org/10.5281/zenodo.4476428" httpUri="https://zenodo.org/record/4476428/files/figure.png" pageId="19" pageNumber="40">Fig. 33</figureCitation>
; mesopleura with lateral round black spot. Broadly oval bugs of pentatomid appearance.
<taxonomicName id="4C314D51A30DF4627DCE79A430D256C3" ID-CoL="7PMQV" authority="Breddin, 1898" authorityName="Breddin" authorityYear="1898" box="[252,511,1510,1529]" class="Insecta" family="Acanthosomatidae" genus="Noualhieridia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAC0A30DF4627DCE79A430D256C3" bold="true" box="[252,511,1510,1529]" pageId="19" pageNumber="40">
<emphasis id="B945EAC0A30DF4627DCE79A4305B56C3" bold="true" box="[252,374,1510,1529]" italics="true" pageId="19" pageNumber="40">Noualhieridia</emphasis>
<bibRefCitation id="EFA04B23A30DF4627C4C79A430D256C3" author="BREDDIN G." box="[382,511,1510,1529]" pageId="19" pageNumber="40" pagination="262 - 268" refId="ref13049" refString="BREDDIN G. 1898: Studia hemipterologica III. Entomologische Nachrichten (Berlin) 24: 262 - 268." type="journal article" year="1898">Breddin, 1898</bibRefCitation>
</emphasis>
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........................................................................................................ 2
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<paragraph id="8B8E36D2A30AF4657DA57CA2354B5212" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57CA2354B5212" pageId="20" pageNumber="41">
– Mesosternal carina well developed. Antennae 4-segmented, antennomere 2 very long. Metathoracic scent gland complex different (
<figureCitation id="130A2A57A30AF4657C6E7CB830EC5237" box="[348,449,250,269]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStart-7="Figs" captionStart-8="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionStartId-2="8.[151,190,917,936]" captionStartId-3="10.[151,190,1518,1537]" captionStartId-4="11.[113,152,838,857]" captionStartId-5="13.[113,152,1072,1091]" captionStartId-6="14.[151,190,1263,1282]" captionStartId-7="16.[151,190,737,756]" captionStartId-8="17.[113,152,1206,1225]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetBox-2="[151,1126,220,883]" captionTargetBox-3="[151,1124,361,1478]" captionTargetBox-4="[113,1088,220,800]" captionTargetBox-5="[115,1088,220,1043]" captionTargetBox-6="[151,1126,220,1229]" captionTargetBox-7="[151,1126,220,706]" captionTargetBox-8="[113,1088,220,1175]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetId-2="figure-211@8.[151,1126,220,883]" captionTargetId-3="figure-11@10.[151,1124,361,1478]" captionTargetId-4="figure-281@11.[113,1088,220,800]" captionTargetId-5="figure-165@13.[113,1088,220,1043]" captionTargetId-6="figure-112@14.[151,1126,220,1229]" captionTargetId-7="figure-268@16.[151,1126,220,706]" captionTargetId-8="figure-16@17.[113,1088,220,1175]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="16" captionTargetPageId-8="17" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." captionText-2="Figs. 8-10. Mahea andriai (Cachan, 1952). 8 – pronotum, dorsal view; 9 – pronotum, posterior view; 10 – abdomen, ventral view." captionText-3="Figs. 11-16. Mahea distanti sp. nov. 11 – antenna; 12 – pronotum, dorsal view; 13 – pronotum, posterior view; 14 – pleural margin and connexiva, dorsal view; 15 – hind leg (tarsomeres and claws missing); 16 – abdomen, ventral view." captionText-4="Figs. 17-19. Mahea distanti sp. nov. 17 – pygophore, ventral view; 18 – right paramere, dorsal view; 19 – aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." captionText-5="Figs. 20-23. Mahea durrelli sp. nov. 20 – pronotum, dorsal view; 21 – pronotum, posterior view; 22 – hind leg (claws missing); 23 – abdomen, ventral view." captionText-6="Figs. 24-27. Mahea durrelli sp. nov. 24 – pygophore, posterior view; 25 – pygophore, ventral view; 26 – left paramere, ventral view; 27 – aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." captionText-7="Figs. 28-30. Mahea parvula sp. nov. 28 – pronotum, dorsal view; 29 – pronotum, posterior view; 30 – abdomen, ventral view." captionText-8="Figs. 31-34. Pleura and metathoracic scent gland opening, most exposed view. 31 – Mahea sexualis Distant, 1909; 32 – Catadipson imernensis (Cachan, 1952); 33 – Noualhieridia rufa Cachan, 1952; 30 – Ibocoris ficivora Roche, 1947." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" figureDoi-2="http://doi.org/10.5281/zenodo.4476408" figureDoi-3="http://doi.org/10.5281/zenodo.4476412" figureDoi-4="http://doi.org/10.5281/zenodo.4476414" figureDoi-5="http://doi.org/10.5281/zenodo.4476418" figureDoi-6="http://doi.org/10.5281/zenodo.4476422" figureDoi-7="http://doi.org/10.5281/zenodo.4476424" figureDoi-8="http://doi.org/10.5281/zenodo.4476428" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" httpUri-2="https://zenodo.org/record/4476408/files/figure.png" httpUri-3="https://zenodo.org/record/4476412/files/figure.png" httpUri-4="https://zenodo.org/record/4476414/files/figure.png" httpUri-5="https://zenodo.org/record/4476418/files/figure.png" httpUri-6="https://zenodo.org/record/4476422/files/figure.png" httpUri-7="https://zenodo.org/record/4476424/files/figure.png" httpUri-8="https://zenodo.org/record/4476428/files/figure.png" pageId="20" pageNumber="41">Figs. 31-32</figureCitation>
), mesopleura without lateral round black spot. Slender species, body more or less parallel. ......................................................................................................................................................... 4
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<keyStep id="30C52E77A30AF4657DA57D6D354A5297" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57D6D354B5267" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57D6D354B5267" pageId="20" pageNumber="41">
2. Body dorsally (including connexiva), antennae and legs entirely terracotta orange to red. Antero-lateral margins of pronotum convex. ......................................................................................
<taxonomicName id="4C314D51A30AF4657E097D08354B5267" ID-CoL="7W3TY" authority="Cachan, 1952" authorityName="Cachan" authorityYear="1952" box="[827,1126,330,349]" class="Insecta" family="Acanthosomatidae" genus="Noualhieridia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B945EAC0A30AF4657E097D08354B5267" bold="true" box="[827,1126,330,349]" pageId="20" pageNumber="41">
<emphasis id="B945EAC0A30AF4657E097D0832CF5267" bold="true" box="[827,994,330,349]" italics="true" pageId="20" pageNumber="41">Noualhieridia rufa</emphasis>
<bibRefCitation id="EFA04B23A30AF4657ED87D08354B5267" author="CACHAN P." box="[1002,1126,330,349]" pageId="20" pageNumber="41" pagination="231 - 462" refId="ref13113" refString="CACHAN P. 1952. Les Pentatomidae de Madagascar (Hemipteres Heteropteres). Memoires de l'Institut Scientifique de Madagascar, Serie E 1 (2): 231 - 462." type="journal article" year="1952">Cachan, 1952</bibRefCitation>
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<paragraph id="8B8E36D2A30AF4657DA57D27354A5297" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57D27354A5297" pageId="20" pageNumber="41">– Body dorsally, antennae and legs multicoloured. Antero-lateral margins of pronotum convex or concave with pale spot; base of corium with another pale spot laterally; postero-lateral angles of connexiva with black spots. ............................................................................................................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E77A30AF4657DA57DF7354A5108" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57DF7354B52D8" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57DF7354B52D8" pageId="20" pageNumber="41">
3. Antero-lateral pronotal margins convex, with oval pale spot overlapping to pronotal disc; trochanters, apices of femora and entire tibiae, tarsi and antennae black. ...............................
<taxonomicName id="4C314D51A30AF4657E227D8D354B52D8" ID-CoL="7W3TX" authority="Breddin, 1898" authorityName="Breddin" authorityYear="1898" box="[784,1126,463,482]" class="Insecta" family="Acanthosomatidae" genus="Noualhieridia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="ornatula">
<emphasis id="B945EAC0A30AF4657E227D8D354B52D8" bold="true" box="[784,1126,463,482]" pageId="20" pageNumber="41">
<emphasis id="B945EAC0A30AF4657E227D8D32F052D8" bold="true" box="[784,989,463,482]" italics="true" pageId="20" pageNumber="41">Noualhieridia ornatula</emphasis>
<bibRefCitation id="EFA04B23A30AF4657ED77D8D354B52D8" author="BREDDIN G." box="[997,1126,463,482]" pageId="20" pageNumber="41" pagination="262 - 268" refId="ref13049" refString="BREDDIN G. 1898: Studia hemipterologica III. Entomologische Nachrichten (Berlin) 24: 262 - 268." type="journal article" year="1898">Breddin, 1898</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57DA8354A5108" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57DA8354A5108" pageId="20" pageNumber="41">
– Antero-lateral pronotal margins slightly concave, with long narrow pale spot not overlapping to pronotal disc; legs and antennae entirely pale, only apical halves of antennomeres 3-5 blackened. ..........................................
<emphasis id="B945EAC0A30AF4657D8F7E6233D05108" box="[189,765,544,562]" italics="true" pageId="20" pageNumber="41">............................................................................................................</emphasis>
<taxonomicName id="4C314D51A30AF4657E367E5D354A5108" ID-CoL="7W3TW" authority="Cachan, 1952" authorityName="Cachan" authorityYear="1952" box="[772,1127,543,562]" class="Insecta" family="Acanthosomatidae" genus="Noualhieridia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B945EAC0A30AF4657E367E5D32CC5108" bold="true" box="[772,993,543,562]" italics="true" pageId="20" pageNumber="41">Noualhieridia marginata</emphasis>
<bibRefCitation id="EFA04B23A30AF4657EDB7E5D354A5108" author="CACHAN P." box="[1001,1127,543,562]" pageId="20" pageNumber="41" pagination="231 - 462" refId="ref13113" refString="CACHAN P. 1952. Les Pentatomidae de Madagascar (Hemipteres Heteropteres). Memoires de l'Institut Scientifique de Madagascar, Serie E 1 (2): 231 - 462." type="journal article" year="1952">Cachan, 1952</bibRefCitation>
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<keyStep id="30C52E77A30AF4657DA57E78354B5032" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57E78354B51A7" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57E78354B51A7" pageId="20" pageNumber="41">
4. Elongate oval species. Head very broad, slightly broader than two thirds of pronotal width.Apex of clypeus free, mandibular plates projected as conspicuous apical spines. Humeral angles of pronotum not produced, rounded. Metathoracic scent gland complex as in
<figureCitation id="130A2A57A30AF4657F307E2D331251B8" box="[514,575,623,642]" captionStart="Figs" captionStartId="17.[113,152,1206,1225]" captionTargetBox="[113,1088,220,1175]" captionTargetId="figure-16@17.[113,1088,220,1175]" captionTargetPageId="17" captionText="Figs. 31-34. Pleura and metathoracic scent gland opening, most exposed view. 31 – Mahea sexualis Distant, 1909; 32 – Catadipson imernensis (Cachan, 1952); 33 – Noualhieridia rufa Cachan, 1952; 30 – Ibocoris ficivora Roche, 1947." figureDoi="http://doi.org/10.5281/zenodo.4476428" httpUri="https://zenodo.org/record/4476428/files/figure.png" pageId="20" pageNumber="41">Fig. 32</figureCitation>
.Connexival margins without apparent spines.Head and pronotum with very large black punctures.
<emphasis id="B945EAC0A30AF4657CFF7EC9332551A7" box="[461,520,651,669]" italics="true" pageId="20" pageNumber="41">...........</emphasis>
<taxonomicName id="4C314D51A30AF4657F217EC833D251A7" ID-CoL="7NS2D" authority="Breddin, 1903" authorityName="Breddin" authorityYear="1903" box="[531,767,650,669]" class="Insecta" family="Acanthosomatidae" genus="Catadipson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAC0A30AF4657F217EC8335B51A7" bold="true" box="[531,630,650,669]" italics="true" pageId="20" pageNumber="41">Catadipson</emphasis>
<bibRefCitation id="EFA04B23A30AF4657F4E7EC833D251A7" author="BREDDIN G." box="[636,767,650,669]" pageId="20" pageNumber="41" pagination="90 - 91" refId="ref13069" refString="BREDDIN G. 1903: Catadipson n. gen. Acanthosominorum (Hemiptera Heteroptera). Societas Entomologica 18: 90 - 91." type="journal article" year="1903">Breddin, 1903</bibRefCitation>
</taxonomicName>
:
<taxonomicName id="4C314D51A30AF4657E3B7EC8354B51A7" ID-CoL="7RZBG" authority="(Cachan, 1952)" baseAuthorityName="Cachan" baseAuthorityYear="1952" box="[777,1126,650,669]" class="Insecta" family="Acanthosomatidae" genus="Catadipson" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="imernensis">
<emphasis id="B945EAC0A30AF4657E3B7EC832FF51A7" bold="true" box="[777,978,650,669]" italics="true" pageId="20" pageNumber="41">Catadipson imernensis</emphasis>
(
<bibRefCitation id="EFA04B23A30AF4657ED27EC8354C51A7" author="CACHAN P." box="[992,1121,650,669]" pageId="20" pageNumber="41" pagination="231 - 462" refId="ref13113" refString="CACHAN P. 1952. Les Pentatomidae de Madagascar (Hemipteres Heteropteres). Memoires de l'Institut Scientifique de Madagascar, Serie E 1 (2): 231 - 462." type="journal article" year="1952">Cachan, 1952</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57EE7354B5032" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57EE7354B5032" pageId="20" pageNumber="41">
– Not conspicuously elongate, parallel species. Head only as broad as ca one half to one third of pronotal width). Mandibular plates without apical spines, meeting or not in front of clypeus. Humeral angles of pronotum each with conspicuous spine. Metathoracic scent gland complex as in
<figureCitation id="130A2A57A30AF4657FDC7E98320151D7" box="[750,812,730,749]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 31</figureCitation>
. Connexival margins spinous. Head and pronotum with small dark punctures.
<taxonomicName id="4C314D51A30AF4657F2D7EB633F65032" ID-CoL="e7fdc867-b6fc-48cf-921a-89904d66d8e7" authority="Distant, 1908" authorityName="Distant" authorityYear="1908" box="[543,731,756,776]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAC0A30AF4657F2D7EB633F65032" bold="true" box="[543,731,756,776]" pageId="20" pageNumber="41">
<emphasis id="B945EAC0A30AF4657F2D7EB63371503D" bold="true" box="[543,604,756,775]" italics="true" pageId="20" pageNumber="41">Mahea</emphasis>
Distant, 1908
</emphasis>
</taxonomicName>
...................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E77A30AF4657DA57F4D354B5007" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57F4D354B5018" blockId="20.[151,1128,223,1496]" box="[151,1126,783,803]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57F4D354B5018" box="[151,1126,783,803]" pageId="20" pageNumber="41">5. Humeral angles gradually narrowed, each produced into conical spine, not constricted basally. ...................... 6</keyLead>
</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57F68354B5007" blockId="20.[151,1128,223,1496]" box="[151,1126,810,829]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57F68354B5007" box="[151,1126,810,829]" pageId="20" pageNumber="41">– Humeral angles abruptly narrowed, constricted, each with long needle-like spine. ........................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E77A30AF4657DA57F07354A50E7" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57F07354A50B7" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57F07354A50B7" pageId="20" pageNumber="41">
6. Smaller species (
<quantity id="4CC99B37A30AF4657C7A7F0730BD5062" box="[328,400,837,856]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="20" pageNumber="41" unit="mm" value="7.3">7.3 mm</quantity>
). Humeral angles of pronotum less prominent (
<figureCitation id="130A2A57A30AF4657E187F0732BD5062" box="[810,912,837,856]" captionStart="Figs" captionStartId="16.[151,190,737,756]" captionTargetBox="[151,1126,220,706]" captionTargetId="figure-268@16.[151,1126,220,706]" captionTargetPageId="16" captionText="Figs. 28-30. Mahea parvula sp. nov. 28 – pronotum, dorsal view; 29 – pronotum, posterior view; 30 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476424" httpUri="https://zenodo.org/record/4476424/files/figure.png" pageId="20" pageNumber="41">Figs. 28-29</figureCitation>
). Head slightly narrower than half of pronotal width. Clypeus narrowly free apically. Lateral connexival spines as in
<figureCitation id="130A2A57A30AF4657EFA7F1D352A5048" box="[968,1031,863,882]" captionStart="Figs" captionStartId="16.[151,190,737,756]" captionTargetBox="[151,1126,220,706]" captionTargetId="figure-268@16.[151,1126,220,706]" captionTargetPageId="16" captionText="Figs. 28-30. Mahea parvula sp. nov. 28 – pronotum, dorsal view; 29 – pronotum, posterior view; 30 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476424" httpUri="https://zenodo.org/record/4476424/files/figure.png" pageId="20" pageNumber="41">Fig. 30</figureCitation>
. ................
<emphasis id="B945EAC0A30AF4657D8F7F39353750B7" box="[189,1050,890,909]" italics="true" pageId="20" pageNumber="41">
.......................................................................................................................................
<taxonomicName id="4C314D51A30AF4657EA67F38353750B7" ID-CoL="7VRC6" box="[916,1050,890,909]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="parvula" status="sp. nov.">
<emphasis id="B945EAC0A30AF4657EA67F38353750B7" bold="true" box="[916,1050,890,909]" pageId="20" pageNumber="41">Mahea parvula</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A27657BBA30AF46579137F38354A50B7" box="[1057,1127,890,909]" pageId="20" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57FD7354A50E7" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57FD7354A50E7" pageId="20" pageNumber="41">
– Larger species (&gt;
<quantity id="4CC99B37A30AF4657C677FD730A15092" box="[341,396,917,936]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="41" unit="mm" value="8.0">8 mm</quantity>
). Humeral angles of pronotum more prominent (
<figureCitation id="130A2A57A30AF4657E047FD732AB5092" box="[822,902,917,936]" captionStart="Figs" captionStartId="8.[151,190,917,936]" captionTargetBox="[151,1126,220,883]" captionTargetId="figure-211@8.[151,1126,220,883]" captionTargetPageId="8" captionText="Figs. 8-10. Mahea andriai (Cachan, 1952). 8 – pronotum, dorsal view; 9 – pronotum, posterior view; 10 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476408" httpUri="https://zenodo.org/record/4476408/files/figure.png" pageId="20" pageNumber="41">Figs. 8-9</figureCitation>
,
<figureCitation id="130A2A57A30AF4657EA37FD732E95092" box="[913,964,917,936]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">12-13</figureCitation>
). Head as wide as about one third of pronotal width. Clypeus completely enclosed apically. Lateral connexival spines different (
<figureCitation id="130A2A57A30AF4657D8E7F8831D150E7" box="[188,252,970,989]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 10</figureCitation>
). .............................................................................................................................................................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E77A30AF4657DA57FA7354A5638" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57FA7354A5758" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57FA7354A5758" pageId="20" pageNumber="41">
7. Body almost uniformly yellowish ochraceous with small dark punctures (except darkened lateral corners of pronotum, apex of scutellum, antennae and paired black spots laterally on abdominal venter). Head with vague impressed rows of punctures. Apex of scutellum more broadly rounded. Sternum without black punctures. Hairs on antennomere 2 shorter than its diameter. Metapleura not swollen. Lateral connexival spines as in
<figureCitation id="130A2A57A30AF46579217877357F5772" box="[1043,1106,1077,1096]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 10</figureCitation>
.. ...........................................................................................................................
<taxonomicName id="4C314D51A30AF4657E62780D354A5758" ID-CoL="7VRC3" authority="(Cachan, 1952)" baseAuthorityName="Cachan" baseAuthorityYear="1952" box="[848,1127,1103,1122]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="andriai">
<emphasis id="B945EAC0A30AF4657E62780D354A5758" bold="true" box="[848,1127,1103,1122]" pageId="20" pageNumber="41">
<emphasis id="B945EAC0A30AF4657E62780D32F95758" bold="true" box="[848,980,1103,1122]" italics="true" pageId="20" pageNumber="41">Mahea andriai</emphasis>
(
<bibRefCitation id="EFA04B23A30AF4657ED0780D354D5758" author="CACHAN P." box="[994,1120,1103,1122]" pageId="20" pageNumber="41" pagination="231 - 462" refId="ref13113" refString="CACHAN P. 1952. Les Pentatomidae de Madagascar (Hemipteres Heteropteres). Memoires de l'Institut Scientifique de Madagascar, Serie E 1 (2): 231 - 462." type="journal article" year="1952">Cachan, 1952</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57828354A5638" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57828354A5638" pageId="20" pageNumber="41">
– Body more colourful, covered with prominent black punctures. Basic colour ochraceous, posterior part of pronotum with reddish tones. Anterior part of scutellum reddish brown, antero-lateral angles and posterior part contrastly pale, apex of scutellum black. Dark punctures on head, pleura and abdominal venter coalescent, forming conspicuous rows (usually impressed and darkened). Apex of scutellum narrowly rounded. Sternum with black punctures. Hairs on antennomere 2 as long as or slightly longer than its diameter. Metapleura conspicuously swollen. Lateral connexival spines as in
<figureCitation id="130A2A57A30AF4657F2378AD337D5638" box="[529,592,1263,1282]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[113,152,1493,1512]" captionStartId-1="6.[151,190,654,673]" captionTargetBox-0="[114,1065,227,1462]" captionTargetBox-1="[151,1126,220,628]" captionTargetId-0="figure-16@5.[114,1065,220,1467]" captionTargetId-1="figure-323@6.[151,1126,220,628]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." captionText-1="Figs. 6-7. Mahea sexualis Distant, 1909. Pygophore (lectotype). 6 – posterior view; 7 – ventral view. Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476402" figureDoi-1="http://doi.org/10.5281/zenodo.4476404" httpUri-0="https://zenodo.org/record/4476402/files/figure.png" httpUri-1="https://zenodo.org/record/4476404/files/figure.png" pageId="20" pageNumber="41">Fig. 16</figureCitation>
. ...........................................................
<emphasis id="B945EAC0A30AF4657EA478AD354A5638" bold="true" box="[918,1127,1263,1282]" pageId="20" pageNumber="41">
<taxonomicName id="4C314D51A30AF4657EA478AD35375638" ID-CoL="7VRC4" box="[918,1050,1263,1282]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="distanti" status="sp. nov.">
<emphasis id="B945EAC0A30AF4657EA478AD35375638" bold="true" box="[918,1050,1263,1282]" italics="true" pageId="20" pageNumber="41">Mahea distanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657BBA30AF465791378AD354A5638" box="[1057,1127,1263,1282]" pageId="20" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E77A30AF4657DA57948354A56E2" pageId="20" pageNumber="41">
<paragraph id="8B8E36D2A30AF4657DA57948354B5657" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57948354B5657" pageId="20" pageNumber="41">
8. Larger, more robust species (
<quantity id="4CC99B37A30AF4657C8A7948333E5627" box="[440,531,1290,1309]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="20" pageNumber="41" unit="mm" value="7.75" valueMax="8.0" valueMin="7.5">7.5-8 mm</quantity>
). Humeral angles not raised upwards (
<figureCitation id="130A2A57A30AF4657E58794832965627" box="[874,955,1290,1309]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Figs. 2-3</figureCitation>
). Male: metapleura swollen; hind femora and tibiae curved inwards; abdomen ventrally with only one row of black spots mediolaterally on sterna 3-7; connexiva as in
<figureCitation id="130A2A57A30AF4657F34797D33175668" box="[518,570,1343,1362]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 4</figureCitation>
; postero-lateral angles of sternum 7 triangular, not curved laterad (
<figureCitation id="130A2A57A30AF4657D8E791831DC5657" box="[188,241,1370,1389]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 4</figureCitation>
). Female: connexiva as in
<figureCitation id="130A2A57A30AF4657CE5791833215657" box="[471,524,1370,1389]" captionStart="Figs" captionStartId="5.[113,152,1493,1512]" captionTargetBox="[114,1065,227,1462]" captionTargetId="figure-16@5.[114,1065,220,1467]" captionTargetPageId="5" captionText="Figs. 1-5. Mahea sexualis Distant, 1909. 1 – head, dorsal view; 2 – pronotum, dorsal view; 3 – pronotum, posterior view; 4 – abdomen of male (lectotype), ventral view; 5 – abdomen of female (paralectotype), ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476402" httpUri="https://zenodo.org/record/4476402/files/figure.png" pageId="20" pageNumber="41">Fig. 5</figureCitation>
. .............................................................
<taxonomicName id="4C314D51A30AF4657E697918354B5657" ID-CoL="7VRC7" authority="Distant, 1908" authorityName="Distant" authorityYear="1908" box="[859,1126,1370,1389]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="sexualis">
<emphasis id="B945EAC0A30AF4657E697918354B5657" bold="true" box="[859,1126,1370,1389]" pageId="20" pageNumber="41">
<emphasis id="B945EAC0A30AF4657E69791832CB5657" bold="true" box="[859,998,1370,1389]" italics="true" pageId="20" pageNumber="41">Mahea sexualis</emphasis>
Distant, 1908
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36D2A30AF4657DA57937354A56E2" blockId="20.[151,1128,223,1496]" pageId="20" pageNumber="41">
<keyLead id="30C095E7A30AF4657DA57937354A56E2" pageId="20" pageNumber="41">
– Smaller, more slender species (
<quantity id="4CC99B37A30AF4657CF77937333356B2" box="[453,542,1397,1416]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.0" metricValueMin="5.9" pageId="20" pageNumber="41" unit="mm" value="5.95" valueMax="6.0" valueMin="5.9">5.9-6 mm</quantity>
). Humeral angles raised askew upwards (
<figureCitation id="130A2A57A30AF4657EB5793732C656B2" box="[903,1003,1397,1416]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 – pronotum, dorsal view; 21 – pronotum, posterior view; 22 – hind leg (claws missing); 23 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="20" pageNumber="41">Figs. 20-21</figureCitation>
). Male: metapleura not swollen; hind femora and tibiae not curved inwards (
<figureCitation id="130A2A57A30AF4657FE779CD323B5698" box="[725,790,1423,1442]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 – pronotum, dorsal view; 21 – pronotum, posterior view; 22 – hind leg (claws missing); 23 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="20" pageNumber="41">Fig. 22</figureCitation>
); abdomen ventrally with three rows of black spots on each side of sterna 3-6; connexiva as in
<figureCitation id="130A2A57A30AF4657F9079E833CE5687" box="[674,739,1450,1469]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 – pronotum, dorsal view; 21 – pronotum, posterior view; 22 – hind leg (claws missing); 23 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="20" pageNumber="41">Fig. 23</figureCitation>
; postero-lateral angles of sternum 7 broadly lanceolate, distinctly curved laterad (
<figureCitation id="130A2A57A30AF4657CC17987331956E2" box="[499,564,1477,1496]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 – pronotum, dorsal view; 21 – pronotum, posterior view; 22 – hind leg (claws missing); 23 – abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="20" pageNumber="41">Fig. 23</figureCitation>
). Female: unknown. ................................
<emphasis id="B945EAC0A30AF4657EA77986354A56E2" bold="true" box="[917,1127,1476,1496]" pageId="20" pageNumber="41">
<taxonomicName id="4C314D51A30AF4657EA77986353756ED" ID-CoL="7VRC5" box="[917,1050,1476,1495]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="41" phylum="Arthropoda" rank="species" species="durrelli" status="sp. nov.">
<emphasis id="B945EAC0A30AF4657EA77986353756ED" bold="true" box="[917,1050,1476,1495]" italics="true" pageId="20" pageNumber="41">Mahea durrelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657BBA30AF46579137987354A56E2" box="[1057,1127,1477,1496]" pageId="20" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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