treatments-xml/data/03/89/41/038941033818FF89FCD2F9C92FF40509.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.3681897" ID-GBIF-Dataset="f68fe85c-2a4c-4625-b684-74f61398d6c2" ID-GBIF-Taxon="162245528" ID-ISSN="1982-0224" ID-Zenodo-Dep="3681897" checkinTime="1582294307033" checkinUser="carolina" docAuthor="Ota, Renata Rúbia, Deprá, Gabriel de Carvalho, Graça, Weferson Júnio da &amp; Pavanelli, Carla Simone" docDate="2018" docId="038941033818FF89FCD2F9C92FF40509" docLanguage="en" docName="NeotropIchthyol.16.2.e170094.pdf.imf" docOrigin="Neotropical Ichthyology 16 (2)" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Geophagus sveni Lucinda, Lucena, Assis 2010" docType="treatment" docVersion="9" lastPageId="96" lastPageNumber="97" masterDocId="FFB0397B3847FFE9FFB4FFDA2B5A0600" masterDocTitle="Peixes da planície de inundação do alto rio Paraná e áreas adjacentes: revised, annotated and updated" masterLastPageNumber="111" masterPageNumber="1" pageId="95" pageNumber="96" updateTime="1634709611861" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Peixes da planície de inundação do alto rio Paraná e áreas adjacentes: revised, annotated and updated</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ota, Renata Rúbia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deprá, Gabriel de Carvalho</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Graça, Weferson Júnio da</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pavanelli, Carla Simone</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-06-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>111</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/1982-0224-20170094</mods:identifier>
<mods:identifier type="GBIF-Dataset">f68fe85c-2a4c-4625-b684-74f61398d6c2</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">3678395</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3681897" ID-GBIF-Taxon="162245528" ID-Zenodo-Dep="3681897" LSID="urn:lsid:plazi:treatment:038941033818FF89FCD2F9C92FF40509" httpUri="http://treatment.plazi.org/id/038941033818FF89FCD2F9C92FF40509" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<subSubSection box="[870,1393,1555,1579]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph blockId="95.[870,1393,1555,1579]" box="[870,1393,1555,1579]" pageId="95" pageNumber="96">
<heading bold="true" box="[870,1393,1555,1579]" centered="true" fontSize="10" level="2" pageId="95" pageNumber="96" reason="2">
<taxonomicName authority="Lucinda, Lucena, Assis, 2010" authorityName="Lucinda, Lucena, Assis" authorityYear="2010" box="[870,1393,1555,1579]" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="sveni">
<emphasis bold="true" box="[870,1393,1555,1579]" pageId="95" pageNumber="96">
<emphasis bold="true" box="[870,1058,1555,1579]" italics="true" pageId="95" pageNumber="96">Geophagus sveni</emphasis>
<bibRefCitation author="Lucinda, Lucena, Assis" box="[1065,1393,1555,1579]" firstAuthor="Lucinda" pageId="95" pageNumber="96" pagination="29 - 42" refId="ref82394" refString="Lucinda PHF, Lucena CAS, Assis NC. Two new species of cichlid fish genus Geophagus Heckel from the rio Tocantins drainage (Perciformes: Cichlidae). Zootaxa. 2010; 2429: 29 - 42." type="journal article" year="2010">Lucinda, Lucena, Assis, 2010</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1093,1171,1619,1643]" pageId="95" pageNumber="96" type="description">
<paragraph blockId="95.[1093,1171,1619,1643]" box="[1093,1171,1619,1643]" pageId="95" pageNumber="96">
<figureCitation box="[1093,1171,1619,1643]" captionStart="Fig" captionStartId="96.[136,175,1808,1832]" captionTargetBox="[136,1459,825,1793]" captionTargetId="figure@96.[136,1459,825,1793]" captionTargetPageId="96" captionText="Fig. 35. a. Geophagus brasiliensis, 148.0 mm SL, fresh specimen, uncat. b. Geophagus sveni, NUP 18698, 115.0 mm SL, rio Ivinheima, Naviraí, State of Mato Grosso do Sul. c. Gymnogeophagus setequedas, 81.3 mm SL, fresh specimen, uncat. d. Laetacara araguaiae, NUP 11315, 43.5 mm SL, upper rio Sucuriú, Paraíso, State of Mato Grosso do Sul. e. Oreochromis niloticus, NUP 1132, 185.0 mm SL, Corumbá Reservoir, Caldas Novas, State of Goiás. d. Satanoperca sp., 150.0 mm SL, fresh specimen, uncat." figureDoi="http://doi.org/10.5281/zenodo.3678469" httpUri="https://zenodo.org/record/3678469/files/figure.png" pageId="95" pageNumber="96">
<emphasis bold="true" box="[1093,1171,1619,1643]" pageId="95" pageNumber="96">Fig. 35</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="description">
<paragraph blockId="95.[813,1452,1684,1964]" lastBlockId="96.[136,776,176,456]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
Body deep; greatest depth contained 2.1 to 2.5 times in SL; head length 3.0 to
<quantity box="[1071,1136,1716,1740]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" pageId="95" pageNumber="96" unit="in" value="3.3">3.3 in</quantity>
SL; snout length 1.5 to 2.4, horizontal orbital diameter 3.2 to 4.7 and least interorbital width
<quantity box="[885,1030,1780,1804]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" metricValueMax="8.89" metricValueMin="6.858" pageId="95" pageNumber="96" unit="in" value="3.1" valueMax="3.5" valueMin="2.7">2.7 to 3.5 in</quantity>
HL. Mouth terminal; premaxilla and dentary with 2-4 teeth rows. Upper lateral line with 20-24 pored scales, lower lateral line with 17-19 pored scales, and longitudinal series with 32-34 scales. Transversal series above upper lateral line with 4 scale rows and below lower lateral line with 5 scale rows. Dorsal fin with XVI- XVIII,10-13 rays, pectoral fin with 15-16, pelvic fin with I,5 and anal fin with III,7-8 rays. Ground color greenish or silvery; iridescent marks on lachrymal, preopercle and opercle; five inconspicuous dark-brown transverse bars on flank and caudal peduncle; black rounded blotch on flank; six to 12 longitudinal series of orange spots on flank. Caudal fin reddish with four to seven iridescent blue stripes, occasionally broken into spots. Dorsal and anal fins with iridescent blue spots, sometimes forming horizontal stripes.
</paragraph>
<paragraph blockId="96.[136,570,497,521]" box="[136,570,497,521]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[136,441,497,521]" pageId="96" pageNumber="97">Maximum standard length</emphasis>
. 200.0 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="96" pageNumber="97" type="biology_ecology">
<paragraph blockId="96.[136,775,561,777]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[136,333,561,586]" pageId="96" pageNumber="97">Biological data.</emphasis>
Feeds on sediment, decomposing organic matter, allochthonous plant fragments, mollusks, crustaceans, cladocerans, copepods and insects. Nests in the substrate, presents external fertilization and displays parental care. Both sexes may care for eggs and juveniles, and males usually defend the territory, while the female cares for the brood (
<bibRefCitation author="Moretto" box="[350,569,753,777]" etAl="et al." firstAuthor="Moretto" pageId="96" pageNumber="97" pagination="3013 - 25" refId="ref83279" refString="Moretto EM, Marciano FT, Velludo MR, Fenerich-Verani N, Espindola ELG, Rocha O. The recent occurrence, establishment and potential impact of Geophagus proximus (Cichlidae: Perciformes) in the Tiete River reservoirs: an Amazonian fish species introduced in the Parana basin (Brazil). Biodivers Conserv. 2008; 17: 3013 - 25." type="journal article" year="2008">
Moretto
<emphasis box="[445,492,753,777]" italics="true" pageId="96" pageNumber="97">et al</emphasis>
., 2008
</bibRefCitation>
;
<bibRefCitation author="Gois" box="[579,761,753,777]" etAl="et al." firstAuthor="Gois" pageId="96" pageNumber="97" pagination="401 - 13" refId="ref81255" refString="Gois KS, Pelicice FM, Gomes LC, Agostinho AA. Invasion of an Amazonian cichlid in the upper Parana River: facilitation by dams and decline of a phylogenetically related species. Hydrobiologia. 2015; 746 (1): 401 - 13." type="journal article" year="2015">
Gois
<emphasis box="[636,690,753,777]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
, 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[820,1298,176,200]" pageId="96" pageNumber="97" type="distribution">
<paragraph blockId="96.[820,1298,176,200]" box="[820,1298,176,200]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[820,967,176,200]" pageId="96" pageNumber="97">Distribution.</emphasis>
Rio Tocantins-Araguaia basin.
</paragraph>
</subSubSection>
<subSubSection pageId="96" pageNumber="97" type="discussion">
<paragraph blockId="96.[820,1460,241,777]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[820,931,241,265]" pageId="96" pageNumber="97">Remarks.</emphasis>
<taxonomicName authorityName="Lucinda, Lucena, Assis" authorityYear="2010" box="[950,1145,241,265]" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="sveni">
<emphasis box="[950,1145,241,265]" italics="true" pageId="96" pageNumber="97">Geophagus sveni</emphasis>
</taxonomicName>
was identified as
<taxonomicName authorityName="Lucinda et al." authorityYear="2010" baseAuthorityName="Castelnau" baseAuthorityYear="1855" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="proximus">
<emphasis box="[1386,1413,241,264]" italics="true" pageId="96" pageNumber="97">G.</emphasis>
cf.
<emphasis box="[820,918,274,297]" italics="true" pageId="96" pageNumber="97">proximus</emphasis>
</taxonomicName>
by
<bibRefCitation author="Graca, Pavanelli" box="[960,1220,273,297]" firstAuthor="Graca" pageId="96" pageNumber="97" pagination="58 - 64" refId="ref84115" refString="Pavanelli CS, Graca WJ, Zawadzki CH, Britski HA, Vidotti AP, Avelino GS, Verissimo S. Fishes from the Corumba Reservoir, Paranaiba River drainage, upper rio Parana basin, State of Goias, Brazil. Check List. 2007; 3 (1): 58 - 64." type="journal article" year="2007">Graça, Pavanelli (2007)</bibRefCitation>
.
<bibRefCitation author="Lucinda" box="[1231,1459,273,297]" etAl="et al." firstAuthor="Lucinda" pageId="96" pageNumber="97" pagination="29 - 42" refId="ref82394" refString="Lucinda PHF, Lucena CAS, Assis NC. Two new species of cichlid fish genus Geophagus Heckel from the rio Tocantins drainage (Perciformes: Cichlidae). Zootaxa. 2010; 2429: 29 - 42." type="journal article" year="2010">
Lucinda
<emphasis box="[1326,1381,273,297]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
(2010)
</bibRefCitation>
described the new species from the rio
<collectingRegion box="[1286,1389,305,329]" country="Brazil" name="Tocantins" pageId="96" pageNumber="97">Tocantins</collectingRegion>
basin and distinguished it from
<taxonomicName authorityName="Lucinda et al." authorityYear="2010" baseAuthorityName="Castelnau" baseAuthorityYear="1855" box="[1105,1238,337,361]" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="proximus">
<emphasis box="[1105,1238,337,361]" italics="true" pageId="96" pageNumber="97">G. proximus</emphasis>
</taxonomicName>
by not presenting a dark-grey preopercular mark.
<taxonomicName authority="Lucinda, Lucena, Assis, 2010" authorityName="Lucinda, Lucena, Assis" authorityYear="2010" box="[1150,1324,369,393]" class="Actinopterygii" family="Cichlidae" genus="Geophaus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="sveni">
<emphasis box="[1150,1324,369,393]" italics="true" pageId="96" pageNumber="97">Geophaus sveni</emphasis>
</taxonomicName>
can also be distinguished by presenting four or five transverse parallel white stripes on the caudal fin, which can be broken into several spots (
<emphasis box="[995,1024,466,489]" italics="true" pageId="96" pageNumber="97">vs.</emphasis>
parallel white stripes complete and horizontally directed on the caudal fin, in
<taxonomicName authorityName="Lucinda et al." authorityYear="2010" baseAuthorityName="Castelnau" baseAuthorityYear="1855" box="[1313,1450,497,521]" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="proximus">
<emphasis box="[1313,1450,497,521]" italics="true" pageId="96" pageNumber="97">G. proximus</emphasis>
</taxonomicName>
) (
<bibRefCitation author="Lucinda" box="[827,1068,529,553]" etAl="et al." firstAuthor="Lucinda" pageId="96" pageNumber="97" pagination="29 - 42" refId="ref82394" refString="Lucinda PHF, Lucena CAS, Assis NC. Two new species of cichlid fish genus Geophagus Heckel from the rio Tocantins drainage (Perciformes: Cichlidae). Zootaxa. 2010; 2429: 29 - 42." type="journal article" year="2010">
Lucinda
<emphasis box="[930,991,529,553]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
, 2010
</bibRefCitation>
). A manuscript on the geographic distribution of
<taxonomicName authorityName="Lucinda, Lucena, Assis" authorityYear="2010" box="[980,1068,561,585]" class="Actinopterygii" family="Cichlidae" genus="Geophaus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="sveni">
<emphasis box="[980,1068,561,585]" italics="true" pageId="96" pageNumber="97">G. sveni</emphasis>
</taxonomicName>
, with a genetic comparison between specimens from the rio
<collectingRegion box="[1108,1211,593,617]" country="Brazil" name="Tocantins" pageId="96" pageNumber="97">Tocantins</collectingRegion>
and from the upper rio
<collectingRegion box="[862,935,625,649]" country="Brazil" name="Parana" pageId="96" pageNumber="97">Paraná</collectingRegion>
floodplain and an analysis of the ontogenetic development of color patterns in this species is being prepared by GCD.
<taxonomicName authorityName="Lucinda, Lucena, Assis" authorityYear="2010" box="[1025,1209,689,713]" class="Actinopterygii" family="Cichlidae" genus="Geophagus" kingdom="Animalia" order="Perciformes" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="sveni">
<emphasis box="[1025,1209,689,713]" italics="true" pageId="96" pageNumber="97">Geophagus sveni</emphasis>
</taxonomicName>
is a non-native species from the upper rio
<collectingRegion box="[1029,1102,721,745]" country="Brazil" name="Parana" pageId="96" pageNumber="97">Paraná</collectingRegion>
basin, and its occurrence can be associated with the aquarium trade.
</paragraph>
</subSubSection>
</treatment>
</document>