162 lines
20 KiB
XML
162 lines
20 KiB
XML
<document id="A09AC658196D55A7596863EAAF01A414" ID-DOI="10.1111/j.1096-3642.2008.00439.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630459907035" checkinUser="felipe" docAuthor="Kriwet, Jürgen, Nunn, Elizabeth V. & Klug, Stefanie" docDate="2009" docId="038887F10F63FFBF8975AA13FD64FA32" docLanguage="en" docName="j.1096-3642.2008.00439.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Ptychotrygon striata Kriwet & Nunn & Klug 2009, SP. NOV." docType="treatment" docUuid="BC153283-FD2C-47DC-9F33-8CAED05F5A5C" docUuidSource="ZooBank" docVersion="6" lastPageNumber="338" masterDocId="FFB1FF890F77FFA98A26A94BFFB8FFBB" masterDocTitle="Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain" masterLastPageNumber="347" masterPageNumber="316" pageNumber="336" updateTime="1699278836986" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B3CBC1076C9CA90B0FA055680BE2B692">Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain</mods:title>
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<mods:namePart id="9FFC23E0D7570740ED9AD93C043C6B37">Kriwet, Jürgen</mods:namePart>
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<mods:namePart id="0049D93F0FF14832F2696C623E94529E">Nunn, Elizabeth V.</mods:namePart>
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<mods:roleTerm id="3FAC442BD0BB1975E01DCB44151C654B">Author</mods:roleTerm>
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<mods:namePart id="9D0AC2A7BEF00F574F5124DCC357F4D3">Klug, Stefanie</mods:namePart>
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<mods:typeOfResource id="FD6A78101DAE145CFB39E29AC5D1CDFE">text</mods:typeOfResource>
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<mods:title id="118F48D75842656223C58D32D7BE9854">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="8884B50643E83B45B3698C2756435381">
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<mods:date id="8AD0EDADFC2DD458EE101E0A1BC6AEF8">2009</mods:date>
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<mods:number id="E2E6586704CE5770FE5390B6AE7D69D3">2009-02-28</mods:number>
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<mods:number id="1ED0A5B67C87C17C869B92B4339C097E">155</mods:number>
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<mods:url id="F214C564BA38E37D701DC27F02446108">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x</mods:url>
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<mods:identifier id="511F266EAE9A51A0ECD43F1EBA9A4777" type="DOI">10.1111/j.1096-3642.2008.00439.x</mods:identifier>
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<treatment id="038887F10F63FFBF8975AA13FD64FA32" ID-DOI="http://doi.org/10.5281/zenodo.5492457" ID-Zenodo-Dep="5492457" LSID="urn:lsid:zoobank.org:act:BC153283-FD2C-47DC-9F33-8CAED05F5A5C" httpUri="http://treatment.plazi.org/id/038887F10F63FFBF8975AA13FD64FA32" lastPageId="22" lastPageNumber="338" pageId="20" pageNumber="336">
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<subSubSection id="C33B656C0F63FFBD8975AA13FBF6FC6A" pageId="20" pageNumber="336" type="nomenclature">
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<paragraph id="8B9E36E70F63FFBD8975AA13FA31FCD6" blockId="20.[851,1417,855,877]" box="[851,1417,855,877]" pageId="20" pageNumber="336">
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<heading id="D0D6818B0F63FFBD8975AA13FA31FCD6" box="[851,1417,855,877]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="336" reason="2">
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<taxonomicName id="4C214D640F63FFBD8975AA13FBC4FCD7" authority="Kriwet & Nunn & Klug, 2009" authorityName="Kriwet & Nunn & Klug" authorityYear="2009" box="[851,1148,856,877]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="20" pageNumber="341" phylum="Chordata" rank="species" species="striata" status="sp. nov.">PTYCHOTRYGON STRIATA</taxonomicName>
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<emphasis id="B955EAF50F63FFBD8EA0AA10FB53FCD7" bold="true" box="[1158,1259,855,876]" pageId="20" pageNumber="336">
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<taxonomicNameLabel id="A266578E0F63FFBD8EA0AA10FB53FCD7" box="[1158,1259,855,876]" pageId="20" pageNumber="336" rank="species">SP. NOV.</taxonomicNameLabel>
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||
</emphasis>
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||
(
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<figureCitation id="131A2A620F63FFBD8EDBAA1CFA39FCD6" box="[1277,1409,855,877]" captionStart="Figure 11" captionStartId="21.[144,223,1664,1683]" captionTargetBox="[184,1384,195,1634]" captionTargetId="figure-0@21.[184,1384,195,1634]" captionTargetPageId="21" captionText="Figure 11. A–P, Ptychotrygon striata sp. nov. A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, antero-lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–X, Iberotrygon plagiolophus gen. et sp. nov. Q–T, antero-lateral tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, anterior juvenile(?) tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445980" httpUri="https://zenodo.org/record/5445980/files/figure.png" pageId="20" pageNumber="336">FIG. 11A–P</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="8B9E36E70F63FFBD891CAA35FBF6FC6A" blockId="20.[826,1441,894,977]" pageId="20" pageNumber="336">
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<emphasis id="B955EAF50F63FFBD891CAA35FB9BFC2F" box="[826,1059,894,916]" italics="true" pageId="20" pageNumber="336">Derivation of name:</emphasis>
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From the Latin word ‘striatus’, ‘striated’, in allusion to the ridges above the lingual interlocking depression.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B656C0F63FFBF891CAAB1FD64FA32" lastPageId="22" lastPageNumber="338" pageId="20" pageNumber="336" type="materials_examined">
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<paragraph id="8B9E36E70F63FFBD891CAAB1FCE5FB95" blockId="20.[826,1441,1018,1070]" pageId="20" pageNumber="336">
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<materialsCitation id="3B493CBA0F63FFBD891CAAB1FCE5FB95" collectionCode="MPZ" pageId="20" pageNumber="336" specimenCode="MPZ 200-31" specimenCount="1" typeStatus="holotype">
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<emphasis id="B955EAF50F63FFBD891CAAB1FC1CFBB4" box="[826,932,1018,1039]" italics="true" pageId="20" pageNumber="336">
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<typeStatus id="549A88450F63FFBD891CAAB1FC26FBB4" box="[826,926,1018,1039]" pageId="20" pageNumber="336" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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An anterior tooth,
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<specimenCode id="DB879E9C0F63FFBD8EB4AAB1FA99FBB4" box="[1170,1313,1018,1039]" collectionCode="MPZ" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="20" pageNumber="336">MPZ 200-31</specimenCode>
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(
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<figureCitation id="131A2A620F63FFBD8F12AAB1FCE8FB95" captionStart="Figure 11" captionStartId="21.[144,223,1664,1683]" captionTargetBox="[184,1384,195,1634]" captionTargetId="figure-0@21.[184,1384,195,1634]" captionTargetPageId="21" captionText="Figure 11. A–P, Ptychotrygon striata sp. nov. A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, antero-lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–X, Iberotrygon plagiolophus gen. et sp. nov. Q–T, antero-lateral tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, anterior juvenile(?) tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445980" httpUri="https://zenodo.org/record/5445980/files/figure.png" pageId="20" pageNumber="336">Fig. 11A– D</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9E36E70F63FFBD891CAD13FC24FB37" blockId="20.[826,1441,1111,1164]" pageId="20" pageNumber="336">
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<emphasis id="B955EAF50F63FFBD891CAD13FC0AFBD6" box="[826,946,1112,1133]" italics="true" pageId="20" pageNumber="336">
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<typeStatus id="549A88450F63FFBD891CAD13FC13FBD6" box="[826,939,1112,1133]" pageId="20" pageNumber="336" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Nine generally well-preserved teeth;
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<materialsCitation id="3B493CBA0F63FFBD8F4CAD13FC20FB37" collectionCode="MPZ" pageId="20" pageNumber="336" specimenCode="MPZ 2005-32" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DB879E9C0F63FFBD8F4CAD13FC20FB37" collectionCode="MPZ" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="20" pageNumber="336">MPZ 2005-32</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B9E36E70F63FFBD891CADFEFC1FFA9D" blockId="20.[826,1441,1204,1318]" pageId="20" pageNumber="336">
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<emphasis id="B955EAF50F63FFBD891CADFEFB96FB72" box="[826,1070,1204,1226]" italics="true" pageId="20" pageNumber="336">Horizon and locality:</emphasis>
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Sample 4 from floodplain environments with channel deposits (facies 2) of the lower Mosqueruela Formation (lower Cenomanian), Aliaga subbasin.
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</paragraph>
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<paragraph id="8B9E36E70F63FFBD891CAC1BFA19F980" blockId="20.[826,1441,1359,1595]" pageId="20" pageNumber="336">
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<emphasis id="B955EAF50F63FFBD891CAC1BFC09FADE" box="[826,945,1360,1381]" italics="true" pageId="20" pageNumber="336">Diagnosis:</emphasis>
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A species of
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<taxonomicName id="4C214D640F63FFBD8E61AC04FB63FADF" authorityName="Jaekel" authorityYear="1894" box="[1095,1243,1359,1380]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="20" pageNumber="336" phylum="Chordata" rank="genus">
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<emphasis id="B955EAF50F63FFBD8E61AC04FB63FADF" box="[1095,1243,1359,1380]" italics="true" pageId="20" pageNumber="336">Ptychotrygon</emphasis>
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</taxonomicName>
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characterized by the following dental features: high and well-cuspidate tooth crown; short and convex labial crown face in profile view; several labial ridges originating from a vertical labial crest; additional irregular and short ridges present; transversal labial and lingual crests on the same; lingual depression well marked; short and flexuous ridges above lingual depression present.
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</paragraph>
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<paragraph id="8B9E36E70F63FFBF891CAF2EFE6CFD5F" blockId="20.[826,1441,1636,1903]" lastBlockId="22.[164,779,198,740]" lastPageId="22" lastPageNumber="338" pageId="20" pageNumber="336">
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<emphasis id="B955EAF50F63FFBD891CAF2EFC7AF9C1" box="[826,962,1637,1658]" italics="true" pageId="20" pageNumber="336">Description:</emphasis>
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The teeth are small and broader than long, with a high cuspidate and pyramidal crown. The central cusp is well developed, and is upright and lingually placed, with a well-marked transverse occlusal crest running through the apex. This crest is bent lingually towards the crown edge in
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<specimenCount id="9D27FD6E0F63FFBD8F08AFB6FCD7F88A" pageId="20" pageNumber="336" type="generic">one specimen</specimenCount>
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. In occlusal view the teeth are heavily ornamented, with ridges radiating from the apex. The labial face has a longitudinal centro-labial ridge, which converges at the apex, and divides into several v-shapes towards the apron. There are several short sub-vertical folds originating at the crown base, and extending perpendicularly towards the lingual face in profile view. The apron is rather long and prominent, with a rounded anterior extremity that is bent slightly basally, and is well detached from the labial contour. The uvula is short and broad with a rounded extremity and pronounced central depression. The top of the central depression is bounded by strong vertical ridges converging towards the apex. In profile view the lingual face is much more abrupt than the labial one. The root is low, slightly displaced lingually, and does not extend laterally beyond the crown. There are two sub-triangular root lobes, separated by a broad and deep groove with a central foramen. The margino-lingual faces have one pair of well-marked margino-lingual foramina.
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</paragraph>
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<caption id="DF5E666F0F62FFBC8AB6AFCBFCD2F89D" ID-DOI="http://doi.org/10.5281/zenodo.5445980" ID-Zenodo-Dep="5445980" httpUri="https://zenodo.org/record/5445980/files/figure.png" pageId="21" pageNumber="337" startId="21.[144,223,1664,1683]" targetBox="[184,1384,195,1634]" targetPageId="21">
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<paragraph id="8B9E36E70F62FFBC8AB6AFCBFCD2F89D" blockId="21.[144,1422,1664,1830]" pageId="21" pageNumber="337">
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<emphasis id="B955EAF50F62FFBC8AB6AFCBFEBCF928" bold="true" box="[144,260,1664,1683]" pageId="21" pageNumber="337">Figure 11.</emphasis>
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A–P,
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<emphasis id="B955EAF50F62FFBC8B6EAFCBFD22F92F" bold="true" box="[328,666,1664,1684]" pageId="21" pageNumber="337">
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||
<taxonomicName id="4C214D640F62FFBC8B6EAFCBFD82F928" authority="Kriwet & Nunn & Klug, 2009" authorityName="Kriwet & Nunn & Klug" authorityYear="2009" box="[328,570,1664,1683]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="21" pageNumber="337" phylum="Chordata" rank="species" species="striata" status="sp. nov.">
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||
<emphasis id="B955EAF50F62FFBC8B6EAFCBFD82F928" bold="true" box="[328,570,1664,1683]" italics="true" pageId="21" pageNumber="337">Ptychotrygon striata</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A266578E0F62FFBC8864AFCAFD22F92F" box="[578,666,1665,1684]" pageId="21" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, antero-lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–X,
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<emphasis id="B955EAF50F62FFBC8880AF93FBC4F950" bold="true" box="[678,1148,1752,1771]" pageId="21" pageNumber="337">
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<taxonomicName id="4C214D640F62FFBC8880AF93FC73F950" authority="Kriwet & Nunn & Klug, 2009" authorityName="Kriwet & Nunn & Klug" authorityYear="2009" box="[678,971,1752,1771]" class="Elasmobranchii" genus="Iberotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="21" pageNumber="337" phylum="Chordata" rank="species" species="plagiolophus" status="gen. et sp. nov.">
|
||
<emphasis id="B955EAF50F62FFBC8880AF93FC73F950" bold="true" box="[678,971,1752,1771]" italics="true" pageId="21" pageNumber="337">Iberotrygon plagiolophus</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A266578E0F62FFBC89F4AF93FBC4F950" box="[978,1148,1752,1771]" pageId="21" pageNumber="337" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Q–T, antero-lateral tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, anterior juvenile(?) tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="8B9E36E70F61FFBF8A82AA5DFEEBFA8F" blockId="22.[164,779,790,1332]" pageId="22" pageNumber="338">
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||
<emphasis id="B955EAF50F61FFBF8A82AA5DFEB7FC90" box="[164,271,790,811]" italics="true" pageId="22" pageNumber="338">Remarks:</emphasis>
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This species is similar to
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<taxonomicName id="4C214D640F61FFBF8851AA5DFEAFFCF2" authorityName="Cappetta & Case" authorityYear="1975" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="22" pageNumber="338" phylum="Chordata" rank="species" species="slaughteri">
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<emphasis id="B955EAF50F61FFBF8851AA5DFEAFFCF2" italics="true" pageId="22" pageNumber="338">Ptychotrygon slaughteri</emphasis>
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</taxonomicName>
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, from the Cretaceous of
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<collectingRegion id="49E5F8050F61FFBF8812AA7EFDCDFCF1" box="[564,629,821,842]" country="United States of America" name="Texas" pageId="22" pageNumber="338">Texas</collectingRegion>
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||
(
|
||
<bibRefCitation id="EFB04B160F61FFBF88A3AA7EFE9DFCD2" author="Cappetta H & Case GR" pageId="22" pageNumber="338" pagination="1 - 112" refId="ref17599" refString="Cappetta H, Case GR. 1999. Additions aux faunes de selaciens du Cretace du Texas. Palaeo Ichthyologica 9: 1 - 112." type="journal article" year="1999">Cappetta & Case, 1999</bibRefCitation>
|
||
), especially in terms of its occlusal ornamentation. However,
|
||
<taxonomicName id="4C214D640F61FFBF8B88AA39FDFFFC3C" authorityName="Cappetta & Case" authorityYear="1975" box="[430,583,882,903]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="22" pageNumber="338" phylum="Chordata" rank="species" species="slaughteri">
|
||
<emphasis id="B955EAF50F61FFBF8B88AA39FDFFFC3C" box="[430,583,882,903]" italics="true" pageId="22" pageNumber="338">P. slaughteri</emphasis>
|
||
</taxonomicName>
|
||
has a marked step-like appearance in occlusal view as a result of a rounded, convex crest that borders the lingual depression. This step-like aspect is not seen in this species due to the absence of the lingual crest. Instead, the central depression of this species is bordered by strong lingual ridges that converge towards the apex. It differs from all other species of
|
||
<taxonomicName id="4C214D640F61FFBF880DAD03FD07FBE6" authorityName="Jaekel" authorityYear="1894" box="[555,703,1096,1117]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="22" pageNumber="338" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F61FFBF880DAD03FD07FBE6" box="[555,703,1096,1117]" italics="true" pageId="22" pageNumber="338">Ptychotrygon</emphasis>
|
||
</taxonomicName>
|
||
in the combination of a well-cuspidate crown, a short and convex labial crown face in profile view, several labial ridges originating from a vertical labial fold, and additional irregular and short ridges, a transversal lingual crest on the same level as the labial one and additional vertical ridges above the lingual interlocking depression.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F61FFBF8AF4AC38FD64FA32" blockId="22.[210,732,1395,1417]" box="[210,732,1395,1417]" pageId="22" pageNumber="338">
|
||
<collectionCode id="ED30AE220F61FFBF8AF4AC38FE99FA3C" box="[210,289,1395,1416]" pageId="22" pageNumber="338">ORDER</collectionCode>
|
||
<taxonomicName id="4C214D640F61FFBF8B0DAC38FD64FA32" authority=", FAM. INDET." authorityName="FAM. INDET." box="[299,732,1395,1417]" class="Elasmobranchii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sclerorhynchiformes" pageId="22" pageNumber="338" phylum="Chordata" rank="order">SCLERORHYNCHIFORMES, FAM. INDET.</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |