418 lines
55 KiB
XML
418 lines
55 KiB
XML
<document id="A5DA7524462B033D348986FAA21F17DB" ID-CLB-Dataset="22253" ID-DOI="10.11646/zootaxa.4772.3.2" ID-GBIF-Dataset="983953d9-e7f1-48e8-b24d-b004ada2d986" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819706" ID-ZooBank="403F169F-DD66-4A64-8887-571D55066E33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175668640" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="038887EDFFF0FFDAFF08188A04F8FA1E" docLanguage="en" docName="zootaxa.4772.3.2.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triacanthoneus toro in Anker 2010" docType="treatment" docVersion="10" lastPageNumber="466" masterDocId="FFB1FF95FFFDFFCAFF9F1A53077EFFA8" masterDocTitle="Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)" masterLastPageNumber="468" masterPageNumber="450" pageNumber="463" updateTime="1698839100595" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="41A347A9CB40C026323177B1C682AD93">Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="368B273C09845FE13F1A8153E48BE356">Anker, Arthur</mods:namePart>
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<mods:title id="0ACAA37AEDEA89926699554F1A9DE81D">Zootaxa</mods:title>
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<mods:part id="E7A3D11B9BE6EB999C77F82F9BF007FB">
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<mods:date id="F7A7765580DB7EC7362790F76D438C40">2020</mods:date>
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<mods:number id="FC992029B50A96164A1D13B4F3A6D795">2020-05-11</mods:number>
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<mods:number id="9480A894EA7D799C323CDCE45062406D">4772</mods:number>
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<treatment id="038887EDFFF0FFDAFF08188A04F8FA1E" ID-DOI="http://doi.org/10.5281/zenodo.3843779" ID-GBIF-Taxon="164035968" ID-Zenodo-Dep="3843779" LSID="urn:lsid:plazi:treatment:038887EDFFF0FFDAFF08188A04F8FA1E" httpUri="http://treatment.plazi.org/id/038887EDFFF0FFDAFF08188A04F8FA1E" lastPageId="16" lastPageNumber="466" pageId="13" pageNumber="463">
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<subSubSection id="C33B6570FFF0FFC7FF08188A0555FD5B" box="[151,555,729,756]" pageId="13" pageNumber="463" type="nomenclature">
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<paragraph id="8B9E36FBFFF0FFC7FF08188A0555FD5B" blockId="13.[151,555,729,791]" box="[151,555,729,756]" pageId="13" pageNumber="463">
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<heading id="D0D68197FFF0FFC7FF08188A0555FD5B" bold="true" box="[151,555,729,756]" fontSize="11" level="1" pageId="13" pageNumber="463" reason="1">
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<taxonomicName id="4C214D78FFF0FFC7FF08188A0555FD5B" ID-CoL="85ZFV" authority="Anker, 2010" authorityName="in Anker" authorityYear="2010" box="[151,555,729,756]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
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<emphasis id="B955EAE9FFF0FFC7FF08188A0555FD5B" bold="true" box="[151,555,729,756]" pageId="13" pageNumber="463">
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<emphasis id="B955EAE9FFF0FFC7FF08188A06F1FD5B" bold="true" box="[151,399,729,756]" italics="true" pageId="13" pageNumber="463">Triacanthoneus toro</emphasis>
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Anker, 2010
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6570FFF0FFC7FF0818AE0672FCBF" box="[151,268,765,791]" pageId="13" pageNumber="463" type="description">
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<paragraph id="8B9E36FBFFF0FFC7FF0818AE0672FCBF" blockId="13.[151,555,729,791]" box="[151,268,765,791]" pageId="13" pageNumber="463">
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<figureCitation id="131A2A7EFFF0FFC7FF0818AE0672FCBF" box="[151,268,765,791]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="14.[151,250,1822,1847]" captionStartId-1="15.[151,250,1855,1880]" captionStartId-2="16.[151,250,753,778]" captionTargetBox-0="[151,1436,181,1797]" captionTargetBox-1="[151,1436,181,1830]" captionTargetBox-2="[355,1235,192,717]" captionTargetId-0="figure@14.[151,1436,181,1797]" captionTargetId-1="figure@15.[151,1436,181,1830]" captionTargetId-2="figure@16.[344,1243,181,727]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURE 9. Triacanthoneus toro Anker, 2010: non-ovigerous specimen (cl 4.6 mm) from Bocas del Toro, Panama (MZUSP 33675); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." captionText-1="FIGURE 10. Triacanthoneus toroAnker, 2010: ovigerous specimen (cl 3.8 mm) from Bocas del Toro, Panama (MZUSP 33674); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." captionText-2="FIGURE 11. Triacanthoneus cf. toro Anker, 2010, ovigerous specimen (cl 4.5 mm) from Sisal, Mexico (OUMNH.ZC. 2016- 01-0087), anterior portion of carapace and frontal appendages, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3819728" figureDoi-1="http://doi.org/10.5281/zenodo.3819730" figureDoi-2="http://doi.org/10.5281/zenodo.3819732" httpUri-0="https://zenodo.org/record/3819728/files/figure.png" httpUri-1="https://zenodo.org/record/3819730/files/figure.png" httpUri-2="https://zenodo.org/record/3819732/files/figure.png" pageId="13" pageNumber="463">Figs. 9–11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6570FFF0FFC7FF08191205A8FCF1" box="[151,726,832,857]" pageId="13" pageNumber="463" type="reference_group">
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<paragraph id="8B9E36FBFFF0FFC7FF08191205A8FCF1" blockId="13.[151,726,832,857]" box="[151,726,832,857]" pageId="13" pageNumber="463">
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<treatmentCitationGroup id="AB3111D5FFF0FFC7FF08191205A8FCF1" box="[151,726,832,857]" pageId="13" pageNumber="463">
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<treatmentCitation id="0A8010EAFFF0FFC7FF0819120568FCF1" ID-DOI="http://doi.org/10.5281/zenodo.6206916" ID-Zenodo-Dep="6206916" author="Anker, A." box="[151,534,832,857]" httpUri="http://treatment.plazi.org/id/039F5848FF9AFFE5E5F3F955FECDCD29" page="49" pageId="13" pageNumber="463" year="2010">
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<taxonomicName id="4C214D78FFF0FFC7FF0819120568FCF1" ID-CoL="85ZFV" authority="Anker 2010 a: 49" authorityName="in Anker" authorityPageNumber="49" authorityYear="2010" box="[151,534,832,857]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
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||
<emphasis id="B955EAE9FFF0FFC7FF081912061AFCF1" box="[151,356,833,857]" italics="true" pageId="13" pageNumber="463">Triacanthoneus toro</emphasis>
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<bibRefCitation id="EFB04B0AFFF0FFC7FEF619130568FCF1" author="Anker, A." box="[361,534,832,857]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a: 49</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs. 1–4, 10A–C.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33B6570FFF0FFC7FF0819D206CBFB30" pageId="13" pageNumber="463" type="materials_examined">
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<paragraph id="8B9E36FBFFF0FFC7FF0819D20389FB84" blockId="13.[151,1437,897,1969]" pageId="13" pageNumber="463">
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<emphasis id="B955EAE9FFF0FFC7FF0819D20606FC34" bold="true" box="[151,376,897,924]" pageId="13" pageNumber="463">Material examined</emphasis>
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.
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<taxonomicName id="4C214D78FFF0FFC7FE1A19D1046EFC34" authority="Anker, 2010: 1" authorityName="in Anker" authorityPageNumber="1" authorityYear="2010" box="[389,784,898,924]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
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||
<emphasis id="B955EAE9FFF0FFC7FE1A19D10519FC34" box="[389,615,898,924]" italics="true" pageId="13" pageNumber="463">Triacanthoneus toro</emphasis>
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||
<materialsCitation id="3B493CA6FFF0FFC7FDF419D1046EFC34" ID-GBIF-Occurrence="2611567303" box="[619,784,898,924]" collectingDate="2015-04-30" collectionCode="MZUSP" collectorName="A. Anker" country="Panama" elevation="1" location="Popa" municipality="Caribbean" pageId="13" pageNumber="463" specimenCode="MZUSP 33674" specimenCount="1" stateProvince="Bocas del Toro">Anker, 2010: 1</materialsCitation>
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</taxonomicName>
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ovigerous specimen (cl
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<quantity id="4CD99B1EFFF0FFC7FB8219D10308FC35" box="[1053,1142,898,925]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="463" unit="mm" value="3.8">3.8 mm</quantity>
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),
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<specimenCode id="DB879E80FFF0FFC7FB1719D1024DFC34" box="[1160,1331,898,924]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="13" pageNumber="463">MZUSP 33674</specimenCode>
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||
,
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||
<collectingCountry id="F336766BFFF0FFC7FAA219D002E7FC35" box="[1341,1433,899,925]" name="Panama" pageId="13" pageNumber="463">Panama</collectingCountry>
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,
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||
<collectingMunicipality id="6BFAAC81FFF0FFC7FF0819F50674FC68" box="[151,266,934,960]" pageId="13" pageNumber="463">Caribbean</collectingMunicipality>
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||
coast,
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||
<collectingRegion id="49E5F819FFF0FFC7FEC719F4057DFC69" box="[344,515,934,961]" country="Panama" name="Bocas del Toro" pageId="13" pageNumber="463">Bocas del Toro</collectingRegion>
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||
, Isla
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||
<location id="8EFE6020FFF0FFC7FDA119F40507FC69" LSID="urn:lsid:plazi:treatment:038887EDFFF0FFDAFF08188A04F8FA1E:8EFE6020FFF0FFC7FDA119F40507FC69" box="[574,633,935,961]" country="Panama" municipality="Caribbean" name="Popa" pageId="13" pageNumber="463" stateProvince="Bocas del Toro">Popa</location>
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||
, shallow sea grass flat, with rubble, sponges, sea plumes, and some corals, depth:
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||
<quantity id="4CD99B1EFFF0FFC7FE8D199A060FFC4D" box="[274,369,969,997]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="13" pageNumber="463" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">
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<elevation id="000CD1C8FFF0FFC7FE8D199A060FFC4D" box="[274,369,969,997]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="13" pageNumber="463" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 m</elevation>
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</quantity>
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||
, leg.
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||
<collectorName id="26D4532DFFF0FFC7FE331999056DFC4C" box="[428,531,970,996]" pageId="13" pageNumber="463">A. Anker</collectorName>
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<emphasis id="B955EAE9FFF0FFC7FD841998052AFC4C" box="[539,596,970,996]" italics="true" pageId="13" pageNumber="463">et al.</emphasis>
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,
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<date id="FF9F103BFFF0FFC7FDFD1999047FFC4C" box="[610,769,969,996]" pageId="13" pageNumber="463" value="2015-04-30">
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<collectingDate id="EFDBE9D3FFF0FFC7FDFD1999047FFC4C" box="[610,769,969,996]" pageId="13" pageNumber="463" value="2015-04-30">30 April 2015</collectingDate>
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</date>
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||
;
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||
<materialsCitation id="3B493CA6FFF0FFC7FC911999038AFB84" ID-GBIF-Occurrence="2611567307" collectingDate="2015-04-28" collectionCode="MZUSP" collectorName="A. Anker" country="Panama" county="Caribbean" elevation="8" location="Cayo Coral" municipality="Bastimentos" pageId="13" pageNumber="463" specimenCode="MZUSP 33675" specimenCount="1" stateProvince="Bocas del Toro">
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||
2 non-ovigerous specimens (cl
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||
<quantity id="4CD99B1EFFF0FFC7FBF2199903B8FC4D" box="[1133,1222,970,997]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="463" unit="mm" value="4.4">4.4 mm</quantity>
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,
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<quantity id="4CD99B1EFFF0FFC7FB4D19990252FC4D" box="[1234,1324,970,997]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="13" pageNumber="463" unit="mm" value="4.6">4.6 mm</quantity>
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||
),
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||
<specimenCode id="DB879E80FFF0FFC7FADF1999079EFBA0" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="13" pageNumber="463">MZUSP 33675</specimenCode>
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||
,
|
||
<collectingCountry id="F336766BFFF0FFC7FF7419BC0639FBA1" box="[235,327,1007,1033]" name="Panama" pageId="13" pageNumber="463">Panama</collectingCountry>
|
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,
|
||
<collectingCounty id="62FF4E77FFF0FFC7FECE19BD06BAFBA0" box="[337,452,1006,1032]" pageId="13" pageNumber="463">Caribbean</collectingCounty>
|
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coast,
|
||
<collectingRegion id="49E5F819FFF0FFC7FD8D19BC05C3FBA1" box="[530,701,1006,1033]" country="Panama" name="Bocas del Toro" pageId="13" pageNumber="463">Bocas del Toro</collectingRegion>
|
||
, Isla
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||
<collectingMunicipality id="6BFAAC81FFF0FFC7FD6819BD04FAFBA0" box="[759,900,1006,1032]" pageId="13" pageNumber="463">Bastimentos</collectingMunicipality>
|
||
,
|
||
<location id="8EFE6020FFF0FFC7FC1019BD036CFBA0" LSID="urn:lsid:plazi:treatment:038887EDFFF0FFDAFF08188A04F8FA1E:8EFE6020FFF0FFC7FC1019BD036CFBA0" box="[911,1042,1006,1032]" country="Panama" county="Caribbean" municipality="Bastimentos" name="Cayo Coral" pageId="13" pageNumber="463" stateProvince="Bocas del Toro">Cayo Coral</location>
|
||
, shallow subtidal sand flat with sea grass and coral rubble, in burrow, suction pump, depth:
|
||
<quantity id="4CD99B1EFFF0FFC7FC971E420418FB85" box="[776,870,1041,1069]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="13" pageNumber="463" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
|
||
<elevation id="000CD1C8FFF0FFC7FC971E420418FB85" box="[776,870,1041,1069]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="13" pageNumber="463" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</elevation>
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||
</quantity>
|
||
, leg.
|
||
<collectorName id="26D4532DFFF0FFC7FC3E1E410379FB84" box="[929,1031,1042,1068]" pageId="13" pageNumber="463">A. Anker</collectorName>
|
||
<emphasis id="B955EAE9FFF0FFC7FB911E400339FB84" box="[1038,1095,1042,1068]" italics="true" pageId="13" pageNumber="463">et al.</emphasis>
|
||
,
|
||
<date id="FF9F103BFFF0FFC7FBCB1E41038AFB84" box="[1108,1268,1041,1068]" pageId="13" pageNumber="463" value="2015-04-28">
|
||
<collectingDate id="EFDBE9D3FFF0FFC7FBCB1E41038AFB84" box="[1108,1268,1041,1068]" pageId="13" pageNumber="463" value="2015-04-28">28 April 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36FBFFF0FFC7FF581E6506CBFB30" blockId="13.[151,1437,897,1969]" pageId="13" pageNumber="463">
|
||
<materialsCitation id="3B493CA6FFF0FFC7FF581E6506CCFB30" ID-GBIF-Occurrence="2611567301" collectingDate="2016-10-18" collectionCode="OUMNH, ZC" collectorName="El Casquito & J. Duarte" country="Mexico" county="Tentative" latitude="21.20782" location="Sisal" longLatPrecision="1" longitude="-90.05507" municipality="Peninsula" pageId="13" pageNumber="463" specimenCount="1" stateProvince="Yucatan">
|
||
<collectingCounty id="62FF4E77FFF0FFC7FF581E650651FBF8" box="[199,303,1078,1104]" pageId="13" pageNumber="463">Tentative</collectingCounty>
|
||
identification:
|
||
<taxonomicName id="4C214D78FFF0FFC7FE471E6504F9FBF8" authority="Anker, 2010: 1" authorityName="in Anker" authorityPageNumber="1" authorityYear="2010" box="[472,903,1078,1104]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FE471E6505F8FBF8" box="[472,646,1078,1104]" italics="true" pageId="13" pageNumber="463">Triacanthoneus</emphasis>
|
||
cf.
|
||
<emphasis id="B955EAE9FFF0FFC7FD301E6405A3FBF8" box="[687,733,1079,1104]" italics="true" pageId="13" pageNumber="463">toro</emphasis>
|
||
Anker, 2010: 1
|
||
</taxonomicName>
|
||
ovigerous specimen (cl
|
||
<quantity id="4CD99B1EFFF0FFC7FB091E660391FBF9" box="[1174,1263,1077,1105]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="463" unit="mm" value="4.5">4.5 mm</quantity>
|
||
),
|
||
<collectionCode id="ED30AE3EFFF0FFC7FA9E1E65020FFBF8" box="[1281,1393,1078,1104]" pageId="13" pageNumber="463">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="ED30AE3EFFF0FFC7FAEB1E6502E7FBF8" box="[1396,1433,1078,1104]" country="Austria" name="Zoological Collection, University of Vienna" pageId="13" pageNumber="463" type="University or college">ZC</collectionCode>
|
||
. 2016-01-0087,
|
||
<collectingCountry id="F336766BFFF0FFC7FEDA1E0906E3FBDC" box="[325,413,1114,1140]" name="Mexico" pageId="13" pageNumber="463">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E5F819FFF0FFC7FE361E08057DFBDD" box="[425,515,1115,1141]" country="Mexico" name="Yucatan" pageId="13" pageNumber="463">Yucatan</collectingRegion>
|
||
<collectingMunicipality id="6BFAAC81FFF0FFC7FD931E090502FBDC" box="[524,636,1114,1140]" pageId="13" pageNumber="463">Peninsula</collectingMunicipality>
|
||
,
|
||
<location id="8EFE6020FFF0FFC7FD171E0905BFFBDC" LSID="urn:lsid:plazi:treatment:038887EDFFF0FFDAFF08188A04F8FA1E:8EFE6020FFF0FFC7FD171E0905BFFBDC" box="[648,705,1114,1140]" country="Mexico" county="Tentative" latitude="21.20782" longLatPrecision="1" longitude="-90.05507" municipality="Peninsula" name="Sisal" pageId="13" pageNumber="463" stateProvince="Yucatan">Sisal</location>
|
||
,
|
||
<collectorName id="26D4532DFFF0FFC7FD511E090428FBDC" box="[718,854,1114,1140]" pageId="13" pageNumber="463">
|
||
<location id="8EFE6020FFF0FFC7FD511E090428FBDC" LSID="urn:lsid:plazi:treatment:038887EDFFF0FFDAFF08188A04F8FA1E:8EFE6020FFF0FFC7FD511E090428FBDC" box="[718,854,1114,1140]" country="Mexico" county="Tentative" latitude="21.20782" longLatPrecision="1" longitude="-90.05507" municipality="Peninsula" name="El Casquito" pageId="13" pageNumber="463" stateProvince="Yucatan">El Casquito</location>
|
||
</collectorName>
|
||
,
|
||
<geoCoordinate id="EE15503CFFF0FFC7FCFC1E0A0371FBDC" box="[867,1039,1113,1140]" degrees="21" direction="north" minutes="12" orientation="latitude" pageId="13" pageNumber="463" precision="1" seconds="28.15" value="21.20782">21°12’28.15”N</geoCoordinate>
|
||
<geoCoordinate id="EE15503CFFF0FFC7FB871E0A03C2FBDC" box="[1048,1212,1113,1140]" degrees="90" direction="west" minutes="3" orientation="longitude" pageId="13" pageNumber="463" precision="1" seconds="18.25" value="-90.05507">90°3’18.25”W</geoCoordinate>
|
||
, under coral, leg.
|
||
<collectorName id="26D4532DFFF0FFC7FA151E080798FB31" pageId="13" pageNumber="463">J. Duarte</collectorName>
|
||
,
|
||
<date id="FF9F103BFFF0FFC7FF6E1E2D06CCFB30" box="[241,434,1150,1176]" pageId="13" pageNumber="463" value="2016-10-18">
|
||
<collectingDate id="EFDBE9D3FFF0FFC7FF6E1E2D06CCFB30" box="[241,434,1150,1176]" pageId="13" pageNumber="463" value="2016-10-18">18 October 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B6570FFF0FFDAFF581EF104F8FA1E" lastPageId="16" lastPageNumber="466" pageId="13" pageNumber="463" type="discussion">
|
||
<paragraph id="8B9E36FBFFF0FFC7FF581EF1079BFAD9" blockId="13.[151,1437,897,1969]" pageId="13" pageNumber="463">
|
||
<emphasis id="B955EAE9FFF0FFC7FF581EF1064AFB14" bold="true" box="[199,308,1186,1212]" pageId="13" pageNumber="463">Remarks</emphasis>
|
||
. The three Panamanian specimens agree very well with the
|
||
<typeStatus id="549A8859FFF0FFC7FC7B1EF0036AFB15" box="[996,1044,1187,1213]" pageId="13" pageNumber="463">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C214D78FFF0FFC7FB191EF003AFFB14" authorityName="in Anker" authorityYear="2010" box="[1158,1233,1187,1212]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FB191EF003AFFB14" box="[1158,1233,1187,1212]" italics="true" pageId="13" pageNumber="463">T. toro</emphasis>
|
||
</taxonomicName>
|
||
, as described and illustrated by
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FEAB1E9506A0FB48" author="Anker, A." box="[308,478,1222,1248]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker (2010a)</bibRefCitation>
|
||
. The variation in the position of the carapacial teeth seems to be minimal (cf.
|
||
<figureCitation id="131A2A7EFFF0FFC7FAFA1E9507D9FAAC" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1847]" captionTargetBox="[151,1436,181,1797]" captionTargetId="figure@14.[151,1436,181,1797]" captionTargetPageId="14" captionText="FIGURE 9. Triacanthoneus toro Anker, 2010: non-ovigerous specimen (cl 4.6 mm) from Bocas del Toro, Panama (MZUSP 33675); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819728" httpUri="https://zenodo.org/record/3819728/files/figure.png" pageId="13" pageNumber="463">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation id="131A2A7EFFF0FFC7FF2B1EB907ADFAAC" box="[180,211,1258,1284]" captionStart="FIGURE 10" captionStartId="15.[151,250,1855,1880]" captionTargetBox="[151,1436,181,1830]" captionTargetId="figure@15.[151,1436,181,1830]" captionTargetPageId="15" captionText="FIGURE 10. Triacanthoneus toroAnker, 2010: ovigerous specimen (cl 3.8 mm) from Bocas del Toro, Panama (MZUSP 33674); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819730" httpUri="https://zenodo.org/record/3819730/files/figure.png" pageId="13" pageNumber="463">10</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FF411EB9060BFAAC" author="Anker, A." box="[222,373,1258,1284]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: figs. 1, 2B, 10A). Interestingly, the species is known with certainty only from
|
||
<collectingRegion id="49E5F819FFF0FFC7FB6D1EB802E2FAAD" box="[1266,1436,1258,1285]" country="Panama" name="Bocas del Toro" pageId="13" pageNumber="463">Bocas del Toro</collectingRegion>
|
||
in
|
||
<collectingCountry id="F336766BFFF0FFC7FF281F5C066EFA81" box="[183,272,1295,1321]" name="Panama" pageId="13" pageNumber="463">Panama</collectingCountry>
|
||
(
|
||
<collectingCountry id="F336766BFFF0FFC7FEBE1F5D0635FA80" box="[289,331,1294,1320]" name="Iceland" pageId="13" pageNumber="463">Isla</collectingCountry>
|
||
Colón,
|
||
<collectingCountry id="F336766BFFF0FFC7FE361F5D0507FA80" box="[425,633,1294,1320]" name="Isla San Cristobal" pageId="13" pageNumber="463">Isla San Cristobal</collectingCountry>
|
||
,
|
||
<collectingCountry id="F336766BFFF0FFC7FD181F5D05D1FA80" box="[647,687,1294,1320]" name="Iceland" pageId="13" pageNumber="463">Isla</collectingCountry>
|
||
Popa and
|
||
<collectingCountry id="F336766BFFF0FFC7FCB31F5D042AFA80" box="[812,852,1294,1320]" name="Iceland" pageId="13" pageNumber="463">Isla</collectingCountry>
|
||
Bastimentos), where it inhabits subtidal sand flats with abundant rubble and sea grass, often near mangroves, at depths ranging from
|
||
<quantity id="4CD99B1EFFF0FFC7FBA41F6103E7FAE5" box="[1083,1177,1330,1357]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="7.0" metricValueMin="1.0" pageId="13" pageNumber="463" unit="m" value="4.0" valueMax="7.0" valueMin="1.0">1 to 7 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FB371F610241FAE4" author="Anker, A." box="[1192,1343,1330,1356]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
; present study).
|
||
</paragraph>
|
||
<paragraph id="8B9E36FBFFF0FFC7FF581F290546F819" blockId="13.[151,1437,897,1969]" pageId="13" pageNumber="463">
|
||
The specimen from Sisal,
|
||
<collectingCountry id="F336766BFFF0FFC7FE6D1F290534FA3C" box="[498,586,1402,1428]" name="Mexico" pageId="13" pageNumber="463">Mexico</collectingCountry>
|
||
, herein tentatively identified as
|
||
<taxonomicName id="4C214D78FFF0FFC7FC281F280359FA3C" authorityName="in Anker" authorityYear="2010" box="[951,1063,1402,1428]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FC281F2804B2FA3C" box="[951,972,1403,1428]" italics="true" pageId="13" pageNumber="463">T.</emphasis>
|
||
cf.
|
||
<emphasis id="B955EAE9FFF0FFC7FC661F280359FA3C" box="[1017,1063,1403,1428]" italics="true" pageId="13" pageNumber="463">toro</emphasis>
|
||
</taxonomicName>
|
||
differs in some aspects from the
|
||
<typeStatus id="549A8859FFF0FFC7FF081FCD0787FA10" box="[151,249,1438,1464]" pageId="13" pageNumber="463" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C214D78FFF0FFC7FEBD1FCC0613FA10" authorityName="in Anker" authorityYear="2010" box="[290,365,1439,1464]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FEBD1FCC0613FA10" box="[290,365,1439,1464]" italics="true" pageId="13" pageNumber="463">T. toro</emphasis>
|
||
</taxonomicName>
|
||
, for instance, in the development of the conical anteromesial tubercle on the eyestalks, which seems to be much smaller in the former specimen and not visible in lateral view (cf.
|
||
<figureCitation id="131A2A7EFFF0FFC7FBD51F9103E5FA74" box="[1098,1179,1474,1500]" captionStart="FIGURE 11" captionStartId="16.[151,250,753,778]" captionTargetBox="[355,1235,192,717]" captionTargetId="figure@16.[344,1243,181,727]" captionTargetPageId="16" captionText="FIGURE 11. Triacanthoneus cf. toro Anker, 2010, ovigerous specimen (cl 4.5 mm) from Sisal, Mexico (OUMNH.ZC. 2016- 01-0087), anterior portion of carapace and frontal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3819732" httpUri="https://zenodo.org/record/3819732/files/figure.png" pageId="13" pageNumber="463">Fig. 11</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FB391F910246FA74" author="Anker, A." box="[1190,1336,1474,1500]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: fig. 2B; this tubercle being also clearly visible in the close-up views of the cephalic region in
|
||
<figureCitation id="131A2A7EFFF0FFC7FBCB1FB503C8F9A8" box="[1108,1206,1510,1536]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1847]" captionTargetBox="[151,1436,181,1797]" captionTargetId="figure@14.[151,1436,181,1797]" captionTargetPageId="14" captionText="FIGURE 9. Triacanthoneus toro Anker, 2010: non-ovigerous specimen (cl 4.6 mm) from Bocas del Toro, Panama (MZUSP 33675); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819728" httpUri="https://zenodo.org/record/3819728/files/figure.png" pageId="13" pageNumber="463">Figs. 9B</figureCitation>
|
||
,
|
||
<figureCitation id="131A2A7EFFF0FFC7FB5D1FB5038AF9A8" box="[1218,1268,1510,1536]" captionStart="FIGURE 10" captionStartId="15.[151,250,1855,1880]" captionTargetBox="[151,1436,181,1830]" captionTargetId="figure@15.[151,1436,181,1830]" captionTargetPageId="15" captionText="FIGURE 10. Triacanthoneus toroAnker, 2010: ovigerous specimen (cl 3.8 mm) from Bocas del Toro, Panama (MZUSP 33674); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819730" httpUri="https://zenodo.org/record/3819730/files/figure.png" pageId="13" pageNumber="463">10B</figureCitation>
|
||
). The Mexican specimen also has three pairs of very robust cuspidate setae inserted at approximately 0.3, 0.5 and 0.7 of the telson length, respectively (Dr. Sammy De Grave, pers. comm.). However, the presence of additional cuspidate setae, as well as additional posterior spiniform setae on the telson might be an aberrant condition, which is occasionally also seen in other genera, especially in
|
||
<taxonomicName id="4C214D78FFF0FFC7FDBB1C2504FFF938" authority="Spence Bate, 1888" authorityName="Spence Bate" authorityYear="1888" box="[548,897,1654,1681]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF0FFC7FDBB1C2505DAF938" box="[548,676,1654,1680]" italics="true" pageId="13" pageNumber="463">Synalpheus</emphasis>
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FD331C2504FFF938" author="Spence Bate, C." box="[684,897,1654,1681]" pageId="13" pageNumber="463" pagination="1 - 942" refId="ref11713" refString="Spence Bate, C. (1888) Report on the Crustacea Macrura collected by the Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. " Challenger " during the years 1873 - 76, 24, i-xc, 1 - 942, pls. 1 - 157." type="journal article" year="1888">Spence Bate, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FC531C2403A7F938" author="Banner, D. M. & Banner, A. H." box="[972,1241,1654,1681]" pageId="13" pageNumber="463" pagination="215 - 257" refId="ref11222" refString="Banner, D. M. & Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner & Banner 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FB791C25025FF938" author="Banner, A. H. & Banner, D. M." box="[1254,1313,1654,1680]" pageId="13" pageNumber="463" pagination="25 - 35" refId="ref11177" refString="Banner, A. H. & Banner, D. M. (1979) Some small collections of alpheid shrimp from the Indian Ocean, including two new species of the genus Synalpheus. Pacific Science, 33, 25 - 35." type="journal article" year="1979">1979</bibRefCitation>
|
||
; A. Anker, pers. obs.). Most of the other morphological characters of the Mexican specimen seem to be congruent with the description of
|
||
<taxonomicName id="4C214D78FFF0FFC7FEA51CEC0531F970" authority="in Anker (2010 a)" authorityName="in Anker" authorityYear="2010" box="[314,591,1726,1752]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FEA51CEC06FAF970" box="[314,388,1727,1752]" italics="true" pageId="13" pageNumber="463">T. toro</emphasis>
|
||
in
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FE381CED0531F970" author="Anker, A." box="[423,591,1726,1752]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker (2010a)</bibRefCitation>
|
||
</taxonomicName>
|
||
. Most importantly, in the specimen from Sisal, the mid-dorsal tooth is situated at about half of the carapace length, whilst the lateral teeth arise in the hepatic area, well below and well more anterior relative to the mid-dorsal tooth (
|
||
<figureCitation id="131A2A7EFFF0FFC7FDC51D5505D5F888" box="[602,683,1798,1824]" captionStart="FIGURE 11" captionStartId="16.[151,250,753,778]" captionTargetBox="[355,1235,192,717]" captionTargetId="figure@16.[344,1243,181,727]" captionTargetPageId="16" captionText="FIGURE 11. Triacanthoneus cf. toro Anker, 2010, ovigerous specimen (cl 4.5 mm) from Sisal, Mexico (OUMNH.ZC. 2016- 01-0087), anterior portion of carapace and frontal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3819732" httpUri="https://zenodo.org/record/3819732/files/figure.png" pageId="13" pageNumber="463">Fig. 11</figureCitation>
|
||
), exactly as in the
|
||
<typeStatus id="549A8859FFF0FFC7FCE71D5504A4F888" box="[888,986,1798,1824]" pageId="13" pageNumber="463" type="holotype">holotype</typeStatus>
|
||
and the topotypical specimens of
|
||
<taxonomicName id="4C214D78FFF0FFC7FACB1D5402E3F888" authorityName="in Anker" authorityYear="2010" box="[1364,1437,1799,1824]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FACB1D5402E3F888" box="[1364,1437,1799,1824]" italics="true" pageId="13" pageNumber="463">T. toro</emphasis>
|
||
</taxonomicName>
|
||
reported above (
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FED31D79069AF8EC" author="Anker, A." box="[332,484,1834,1860]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: fig. 2A, B; see also
|
||
<figureCitation id="131A2A7EFFF0FFC7FD571D790468F8EC" box="[712,790,1834,1860]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1847]" captionTargetBox="[151,1436,181,1797]" captionTargetId="figure@14.[151,1436,181,1797]" captionTargetPageId="14" captionText="FIGURE 9. Triacanthoneus toro Anker, 2010: non-ovigerous specimen (cl 4.6 mm) from Bocas del Toro, Panama (MZUSP 33675); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819728" httpUri="https://zenodo.org/record/3819728/files/figure.png" pageId="13" pageNumber="463">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation id="131A2A7EFFF0FFC7FCBD1D79043FF8EC" box="[802,833,1834,1860]" captionStart="FIGURE 10" captionStartId="15.[151,250,1855,1880]" captionTargetBox="[151,1436,181,1830]" captionTargetId="figure@15.[151,1436,181,1830]" captionTargetPageId="15" captionText="FIGURE 10. Triacanthoneus toroAnker, 2010: ovigerous specimen (cl 3.8 mm) from Bocas del Toro, Panama (MZUSP 33674); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3819730" httpUri="https://zenodo.org/record/3819730/files/figure.png" pageId="13" pageNumber="463">10</figureCitation>
|
||
). Furthermore, in the specimen from Sisal, the postorbital margin margin is broadly rounded, without any trace of orbital teeth (
|
||
<figureCitation id="131A2A7EFFF0FFC7FC451D1D0352F8C0" box="[986,1068,1870,1896]" captionStart="FIGURE 11" captionStartId="16.[151,250,753,778]" captionTargetBox="[355,1235,192,717]" captionTargetId="figure@16.[344,1243,181,727]" captionTargetPageId="16" captionText="FIGURE 11. Triacanthoneus cf. toro Anker, 2010, ovigerous specimen (cl 4.5 mm) from Sisal, Mexico (OUMNH.ZC. 2016- 01-0087), anterior portion of carapace and frontal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3819732" httpUri="https://zenodo.org/record/3819732/files/figure.png" pageId="13" pageNumber="463">Fig. 11</figureCitation>
|
||
), as in
|
||
<taxonomicName id="4C214D78FFF0FFC7FBE31D1C03B8F8C0" authorityName="in Anker" authorityYear="2010" box="[1148,1222,1871,1896]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF0FFC7FBE31D1C03B8F8C0" box="[1148,1222,1871,1896]" italics="true" pageId="13" pageNumber="463">T. toro</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB04B0AFFF0FFC7FB4B1D1D0212F8C0" author="Anker, A." box="[1236,1388,1870,1896]" pageId="13" pageNumber="463" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: fig. 2A, B). On the other hand, the general shape of the rostrum and the size of cornea of the Mexican specimen appear to be more like those of
|
||
<taxonomicName id="4C214D78FFF0FFC7FE381DC4054FF818" authorityName="Anker" authorityYear="2010" box="[423,561,1942,1968]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFF0FFC7FE381DC4054FF818" box="[423,561,1942,1968]" italics="true" pageId="13" pageNumber="463">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5E6673FFF3FFC4FF081D4D048DF8F3" ID-DOI="http://doi.org/10.5281/zenodo.3819728" ID-Zenodo-Dep="3819728" httpUri="https://zenodo.org/record/3819728/files/figure.png" pageId="14" pageNumber="464" startId="14.[151,250,1822,1847]" targetBox="[151,1436,181,1797]" targetPageId="14">
|
||
<paragraph id="8B9E36FBFFF3FFC4FF081D4D048DF8F3" blockId="14.[151,1436,1822,1883]" pageId="14" pageNumber="464">
|
||
<emphasis id="B955EAE9FFF3FFC4FF081D4D0671F89F" bold="true" box="[151,271,1822,1847]" pageId="14" pageNumber="464">FIGURE 9</emphasis>
|
||
.
|
||
<taxonomicName id="4C214D78FFF3FFC4FE821D4C0509F89F" authority="Anker, 2010" authorityName="in Anker" authorityYear="2010" box="[285,631,1822,1847]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF3FFC4FE821D4C0695F89F" box="[285,491,1823,1847]" italics="true" pageId="14" pageNumber="464">Triacanthoneus toro</emphasis>
|
||
Anker, 2010
|
||
</taxonomicName>
|
||
: non-ovigerous specimen (cl 4.6 mm) from Bocas del Toro, Panama (MZUSP 33675); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5E6673FFF2FFC5FF081D6C04E0F8D4" ID-DOI="http://doi.org/10.5281/zenodo.3819730" ID-Zenodo-Dep="3819730" httpUri="https://zenodo.org/record/3819730/files/figure.png" pageId="15" pageNumber="465" startId="15.[151,250,1855,1880]" targetBox="[151,1436,181,1830]" targetPageId="15">
|
||
<paragraph id="8B9E36FBFFF2FFC5FF081D6C04E0F8D4" blockId="15.[151,1436,1855,1916]" pageId="15" pageNumber="465">
|
||
<emphasis id="B955EAE9FFF2FFC5FF081D6C0666F8F0" bold="true" box="[151,280,1855,1880]" pageId="15" pageNumber="465">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C214D78FFF2FFC5FEBC1D13050DF8F0" authority="Anker, 2010" authorityName="in Anker" authorityYear="2010" box="[291,627,1855,1880]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="465" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF2FFC5FEBC1D130691F8F0" box="[291,495,1856,1880]" italics="true" pageId="15" pageNumber="465">Triacanthoneus toro</emphasis>
|
||
Anker, 2010
|
||
</taxonomicName>
|
||
: ovigerous specimen (cl 3.8 mm) from Bocas del Toro, Panama (MZUSP 33674); living shrimp in dorsal [A] and lateral [B] views. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5E6673FFEDFFDAFF0818A204EAFC86" ID-DOI="http://doi.org/10.5281/zenodo.3819732" ID-Zenodo-Dep="3819732" httpUri="https://zenodo.org/record/3819732/files/figure.png" pageId="16" pageNumber="466" startId="16.[151,250,753,778]" targetBox="[355,1235,192,717]" targetPageId="16">
|
||
<paragraph id="8B9E36FBFFEDFFDAFF0818A204EAFC86" blockId="16.[151,1437,753,814]" pageId="16" pageNumber="466">
|
||
<emphasis id="B955EAE9FFEDFFDAFF0818A20667FCA2" bold="true" box="[151,281,753,778]" pageId="16" pageNumber="466">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName id="4C214D78FFEDFFDAFEB918A105E3FCA2" authority="Anker, 2010" authorityName="in Anker" authorityYear="2010" box="[294,669,753,778]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFEDFFDAFEB918A106BAFCA2" box="[294,452,754,778]" italics="true" pageId="16" pageNumber="466">Triacanthoneus</emphasis>
|
||
cf.
|
||
<emphasis id="B955EAE9FFEDFFDAFE7318A1056BFCA1" box="[492,533,754,777]" italics="true" pageId="16" pageNumber="466">toro</emphasis>
|
||
Anker, 2010
|
||
</taxonomicName>
|
||
, ovigerous specimen (cl 4.5 mm) from Sisal, Mexico (OUMNH.ZC. 2016- 01-0087), anterior portion of carapace and frontal appendages, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9E36FBFFEDFFDAFF58190E05EFFB77" blockId="16.[151,1437,860,1463]" pageId="16" pageNumber="466">
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFF58190E06B8FCDE" author="De Grave, S. & Anker, A. & Pindar, C. & Sandell, J. & Johnson, M. L." box="[199,454,860,887]" pageId="16" pageNumber="466" pagination="617 - 624" refId="ref11336" refString="De Grave, S., Anker, A., Pindar, C., Sandell, J. & Johnson, M. L. (2017) Eleven new records of caridean shrimps from Cuban waters (Decapoda, Caridea). Crustaceana, 90, 617 - 624. https: // doi. org / 10.1163 / 15685403 - 00003675" type="journal article" year="2017">
|
||
De Grave
|
||
<emphasis id="B955EAE9FFEDFFDAFEA5190E060CFCDE" box="[314,370,860,886]" italics="true" pageId="16" pageNumber="466">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
, while reporting
|
||
<taxonomicName id="4C214D78FFEDFFDAFD1B190E0473FCDE" authorityName="Anker" authorityYear="2010" box="[644,781,860,886]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFD1B190E0473FCDE" box="[644,781,860,886]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49E5F819FFEDFFDAFCCD190F035BFCDE" box="[850,1061,860,886]" country="Cuba" name="Isla de la Juventud" pageId="16" pageNumber="466">Isla de la Juventud</collectingRegion>
|
||
,
|
||
<collectingCountry id="F336766BFFEDFFDAFBAF190F0310FCDE" box="[1072,1134,860,886]" name="Cuba" pageId="16" pageNumber="466">Cuba</collectingCountry>
|
||
, noted some important differences between their single Cuban specimen and the
|
||
<typeStatus id="549A8859FFEDFFDAFD6219D30421FC32" box="[765,863,896,922]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C214D78FFEDFFDAFC1A19D20370FC32" authorityName="Anker" authorityYear="2010" box="[901,1038,896,922]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFC1A19D20370FC32" box="[901,1038,896,922]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
from Alacranes Reef. In the Cuban specimen, which is noticeably smaller than the
|
||
<typeStatus id="549A8859FFEDFFDAFD3619F70470FC16" box="[681,782,932,958]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
, the dorsolateral carapacial teeth are much more advanced compared to those of the
|
||
<typeStatus id="549A8859FFEDFFDAFE27199B0564FC4A" box="[440,538,968,994]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFDB5199B0460FC4A" author="De Grave, S. & Anker, A. & Pindar, C. & Sandell, J. & Johnson, M. L." box="[554,798,968,994]" pageId="16" pageNumber="466" pagination="617 - 624" refId="ref11336" refString="De Grave, S., Anker, A., Pindar, C., Sandell, J. & Johnson, M. L. (2017) Eleven new records of caridean shrimps from Cuban waters (Decapoda, Caridea). Crustaceana, 90, 617 - 624. https: // doi. org / 10.1163 / 15685403 - 00003675" type="journal article" year="2017">
|
||
De Grave
|
||
<emphasis id="B955EAE9FFEDFFDAFD3E199A05A4FC4A" box="[673,730,968,994]" italics="true" pageId="16" pageNumber="466">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
: fig. 1A), almost approaching their position in
|
||
<taxonomicName id="4C214D78FFEDFFDAFAAC199A0203FC4A" authorityName="in Anker" authorityYear="2010" box="[1331,1405,969,994]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFEDFFDAFAAC199A0203FC4A" box="[1331,1405,969,994]" italics="true" pageId="16" pageNumber="466">T. toro</emphasis>
|
||
</taxonomicName>
|
||
. A further difference is that the postorbital margin of the Cuban specimen is much less pronounced and not as angular as in the
|
||
<typeStatus id="549A8859FFEDFFDAFF651E430622FB82" box="[250,348,1040,1066]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C214D78FFEDFFDAFE1F1E420577FB82" authorityName="Anker" authorityYear="2010" box="[384,521,1040,1066]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFE1F1E420577FB82" box="[384,521,1040,1066]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
(Dr. Sammy De Grave, pers. comm.), in which it forms subtriangular orbital teeth (cf.
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFF5B1E670625FBE6" author="Anker, A." box="[196,347,1076,1102]" pageId="16" pageNumber="466" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: fig. 8A, B;
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFE781E6605A7FBE6" author="De Grave, S. & Anker, A. & Pindar, C. & Sandell, J. & Johnson, M. L." box="[487,729,1076,1103]" pageId="16" pageNumber="466" pagination="617 - 624" refId="ref11336" refString="De Grave, S., Anker, A., Pindar, C., Sandell, J. & Johnson, M. L. (2017) Eleven new records of caridean shrimps from Cuban waters (Decapoda, Caridea). Crustaceana, 90, 617 - 624. https: // doi. org / 10.1163 / 15685403 - 00003675" type="journal article" year="2017">
|
||
De Grave
|
||
<emphasis id="B955EAE9FFEDFFDAFDC31E6605E8FBE6" box="[604,662,1076,1102]" italics="true" pageId="16" pageNumber="466">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
: fig. 1A). However, the two pairs of the dorsal cuspidate setae of the telson of the Cuban specimen are small and situated in the posterior third of the telson, i.e. they agree in both their relative size and position with those of the
|
||
<typeStatus id="549A8859FFEDFFDAFD5E1E2F045DFB3E" box="[705,803,1148,1174]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFCAA1E2F04B3FB3E" author="Anker, A." box="[821,973,1148,1174]" pageId="16" pageNumber="466" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
|
||
: fig. 8L). Based on the evaluation of the above features, the Cuban specimen indeed is best assigned to
|
||
<taxonomicName id="4C214D78FFEDFFDAFCC71EF2049CFB12" authorityName="Anker" authorityYear="2010" box="[856,994,1184,1210]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFCC71EF2049CFB12" box="[856,994,1184,1210]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
, whereas the Mexican specimen seems to be morphologically much closer to
|
||
<taxonomicName id="4C214D78FFEDFFDAFDDE1E9605F5FB76" authorityName="in Anker" authorityYear="2010" box="[577,651,1221,1246]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFEDFFDAFDDE1E9605F5FB76" box="[577,651,1221,1246]" italics="true" pageId="16" pageNumber="466">T. toro</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36FBFFEDFFDAFF581EBB04F8FA1E" blockId="16.[151,1437,860,1463]" pageId="16" pageNumber="466">
|
||
With only three specimens of
|
||
<taxonomicName id="4C214D78FFEDFFDAFD881EBB05BBFAAA" authorityName="Anker" authorityYear="2010" box="[535,709,1256,1282]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFEDFFDAFD881EBB05BBFAAA" box="[535,709,1256,1282]" italics="true" pageId="16" pageNumber="466">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
from the southern Gulf of Mexico and
|
||
<collectingCountry id="F336766BFFEDFFDAFB1F1EBB03C5FAAA" box="[1152,1211,1256,1282]" name="Cuba" pageId="16" pageNumber="466">Cuba</collectingCountry>
|
||
presently available, i.e. the
|
||
<typeStatus id="549A8859FFEDFFDAFF731F5F0630FA8E" box="[236,334,1292,1318]" pageId="16" pageNumber="466" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C214D78FFEDFFDAFEE81F5E057CFA8E" authorityName="Anker" authorityYear="2010" box="[375,514,1292,1318]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFEE81F5E057CFA8E" box="[375,514,1292,1318]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
from Alacranes Reef, the specimen from Sisal herein assigned to
|
||
<taxonomicName id="4C214D78FFEDFFDAFB6A1F5E021BFA8E" authorityName="in Anker" authorityYear="2010" box="[1269,1381,1292,1318]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFEDFFDAFB6A1F5E0274FA8E" box="[1269,1290,1293,1318]" italics="true" pageId="16" pageNumber="466">T.</emphasis>
|
||
cf.
|
||
<emphasis id="B955EAE9FFEDFFDAFAA81F5E021BFA8E" box="[1335,1381,1293,1318]" italics="true" pageId="16" pageNumber="466">toro</emphasis>
|
||
</taxonomicName>
|
||
, and the specimen from
|
||
<collectingRegion id="49E5F819FFEDFFDAFEF11F630541FAE2" box="[366,575,1328,1354]" country="Cuba" name="Isla de la Juventud" pageId="16" pageNumber="466">Isla de la Juventud</collectingRegion>
|
||
identified as
|
||
<taxonomicName id="4C214D78FFEDFFDAFD481F620318FAE2" authority="(De Grave et al. 2017)" baseAuthorityName="De Grave" baseAuthorityYear="2017" box="[727,1126,1328,1354]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFD481F62041EFAE2" box="[727,864,1328,1354]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB04B0AFFEDFFDAFCF01F630320FAE2" author="De Grave, S. & Anker, A. & Pindar, C. & Sandell, J. & Johnson, M. L." box="[879,1118,1328,1354]" pageId="16" pageNumber="466" pagination="617 - 624" refId="ref11336" refString="De Grave, S., Anker, A., Pindar, C., Sandell, J. & Johnson, M. L. (2017) Eleven new records of caridean shrimps from Cuban waters (Decapoda, Caridea). Crustaceana, 90, 617 - 624. https: // doi. org / 10.1163 / 15685403 - 00003675" type="journal article" year="2017">
|
||
De Grave
|
||
<emphasis id="B955EAE9FFEDFFDAFC7B1F620362FAE2" box="[996,1052,1328,1354]" italics="true" pageId="16" pageNumber="466">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, it is difficult to make more firm conclusions about the the above observed morphological differences. In the present study, they are considered as part of intraspecific variation within
|
||
<taxonomicName id="4C214D78FFEDFFDAFDC41F2A0599FA3A" box="[603,743,1400,1426]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFEDFFDAFDC41F2A0599FA3A" box="[603,743,1400,1426]" italics="true" pageId="16" pageNumber="466">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFEDFFDAFCBE1F2A0412FA3A" authorityName="in Anker" authorityYear="2010" box="[801,876,1401,1426]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFEDFFDAFCBE1F2A0412FA3A" box="[801,876,1401,1426]" italics="true" pageId="16" pageNumber="466">T. toro</emphasis>
|
||
</taxonomicName>
|
||
, awaiting a thorough morphological + molecular analysis of this material and eventually, collection of new material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |