215 lines
28 KiB
XML
215 lines
28 KiB
XML
<document id="63A84B60460C8DF195B24BDB2C1C4D75" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFA93C4DFE93F9A8FBEB6A33" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada inusitata Köhler 2011, n.sp." docType="treatment" docUuid="5E55C8D9-5E35-4584-8FF0-EC0DE7DB76B4" docUuidSource="ZooBank" docVersion="3" lastPageNumber="186" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="185" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="668C1580B4FB174904F8FF839CF9688D">
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<mods:title id="04010E377CB611EACD695C5BF40C4EDC">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
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<mods:name id="61879DE6C9F229ECB49DCBD9BD02EBF3" type="personal">
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<mods:role id="60CAB3E6A43E843A00BB02C22DB0D8B2">
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<mods:roleTerm id="0B85500F8C29F1F12D5EED7188AD002B">Author</mods:roleTerm>
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<mods:namePart id="91AF461358B0D58C4D69226F4D03FB30">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="731D347877AB50E768C19BE92E35D11D">text</mods:typeOfResource>
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<mods:titleInfo id="4DE003FDFEBB7C0026CFFB10C70150FB">
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<mods:title id="CBB5C976B92F7C703487726C02CC7338">Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part id="F35B0ED78019D08E76D0F5FF7F24D1B5">
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<mods:date id="2486B4437C4B1432D61C0065B6593F3E">2011</mods:date>
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<mods:detail id="58C828468F16F7C117F2450AD7EE1DF0" type="pubDate">
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<mods:number id="0D7DB24F51DFAD1099EBAF250A233D0B">2011-11-30</mods:number>
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</mods:detail>
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<mods:detail id="A27807D8AA7DB2A496E41FA3F292FA7E" type="volume">
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<mods:number id="13427F3518581D312EE92B17CD63B337">63</mods:number>
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</mods:detail>
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<mods:detail id="1FEED15ABF4DE7634D6B5B110FDF395C" type="issue">
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<mods:number id="D1D48211FEBB004594B7C085B03FFBE1">2</mods:number>
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<mods:extent id="C7B3B4681451C8D4A8E3F71AB210C732" unit="page">
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<mods:start id="DC11674AE37971968F7B138D838A8AD8">167</mods:start>
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<mods:end id="87D5D4A37B7B31435CBE59840DAC5C5C">202</mods:end>
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<mods:url id="0023EB8FE625898565EDB986B67CBA70">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
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</mods:location>
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<mods:classification id="1604A0503B6CA5A6248E3E743C556947">journal article</mods:classification>
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<mods:identifier id="7B195509B91AD54604407C11B5949F4F" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
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<mods:identifier id="E485D52248FA8A040902BAC1D053AE3F" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="3A7E77727D78BDAA0EC382077575D6B3" type="Zenodo-Dep">5239547</mods:identifier>
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<mods:identifier id="AAC6CADAE2991B4FA8A3AA27B53631E8" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
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</mods:mods>
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<treatment id="0384503AFFA93C4DFE93F9A8FBEB6A33" LSID="urn:lsid:zoobank.org:act:5E55C8D9-5E35-4584-8FF0-EC0DE7DB76B4" httpUri="http://treatment.plazi.org/id/0384503AFFA93C4DFE93F9A8FBEB6A33" lastPageId="19" lastPageNumber="186" pageId="18" pageNumber="185">
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<subSubSection id="C337B2A7FFA93C4CFE93F9A8FD7868DE" box="[299,661,1658,1684]" pageId="18" pageNumber="185" type="nomenclature">
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<paragraph id="8B92E12CFFA93C4CFE93F9A8FD7868DE" blockId="18.[299,661,1658,1684]" box="[299,661,1658,1684]" pageId="18" pageNumber="185">
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<heading id="D0DA5640FFA93C4CFE93F9A8FD7868DE" box="[299,661,1658,1684]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="185" reason="2">
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<emphasis id="B9593D3EFFA93C4CFE93F9A8FD7868DE" bold="true" box="[299,661,1658,1684]" pageId="18" pageNumber="185">
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<taxonomicName id="4C2D9AAFFFA93C4CFE93F9A8FDBF68DE" ID-CoL="D7RB" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[299,594,1658,1684]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="185" phylum="Mollusca" rank="species" species="inusitata" status="sp. nov.">
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<emphasis id="B9593D3EFFA93C4CFE93F9A8FDBF68DE" bold="true" box="[299,594,1658,1684]" italics="true" pageId="18" pageNumber="185">Amplirhagada inusitata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26A8045FFA93C4CFDE1F9A8FD7868DE" box="[601,661,1658,1684]" pageId="18" pageNumber="185" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C337B2A7FFA93C4CFF1EF961FDB16999" pageId="18" pageNumber="185" type="materials_examined">
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||
<paragraph id="8B92E12CFFA93C4CFF1EF961FD6C6969" blockId="18.[166,794,1715,1827]" pageId="18" pageNumber="185">
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<materialsCitation id="3B45EB71FFA93C4CFF1EF961FD966969" collectingDate="1987-06-14" collectionCode="V" collectorName="NW Kimberley, N" latitude="-14.952778" location="Prince Frederick Harbour" longLatPrecision="21" longitude="125.275" pageId="18" pageNumber="185" specimenCode="RFS 14-1" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B9593D3EFFA93C4CFF1EF961FED16881" bold="true" box="[166,316,1715,1739]" pageId="18" pageNumber="185">
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||
<typeStatus id="54965F8EFFA93C4CFF1EF961FF326881" box="[166,223,1715,1739]" pageId="18" pageNumber="185">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingRegion id="49E92FCEFFA93C4CFEF3F966FDE26881" box="[331,527,1716,1739]" country="Australia" name="Western Australia" pageId="18" pageNumber="185">Western Australia</collectingRegion>
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,
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<collectorName id="26D884FAFFA93C4CFDA4F966FD086881" box="[540,741,1716,1739]" pageId="18" pageNumber="185">NW Kimberley, N</collectorName>
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side of
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<location id="8EF2B7F7FFA93C4CFF7CF903FE3568A2" LSID="urn:lsid:plazi:treatment:0384503AFFA93C4DFE93F9A8FBEB6A33:8EF2B7F7FFA93C4CFF7CF903FE3568A2" box="[196,472,1745,1768]" latitude="-14.952778" longLatPrecision="21" longitude="125.275" name="Prince Frederick Harbour" pageId="18" pageNumber="185" stateProvince="Western Australia">Prince Frederick Harbour</location>
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, near mouth,
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<locationDeviation id="9A20F80EFFA93C4CFDD3F903FEFC694F" location="Mt. Anderdon" pageId="18" pageNumber="185">
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<quantity id="4CD54CC9FFA93C4CFDD3F903FD4268A2" box="[619,687,1745,1768]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="185" unit="km" value="16.0">16 km</quantity>
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W of Mt. Anderdon
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</locationDeviation>
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,
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<quantity id="4CD54CC9FFA93C4CFEA2F93CFE8D694F" box="[282,352,1774,1797]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="185" unit="km" value="0.3">0.3 km</quantity>
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from coast;
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<geoCoordinate id="EE1987EBFFA93C4CFE66F93CFDB9694F" box="[478,596,1774,1797]" degrees="14" direction="south" minutes="57" orientation="latitude" pageId="18" pageNumber="185" precision="15" seconds="10" value="-14.952778">14°57'10"S</geoCoordinate>
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<geoCoordinate id="EE1987EBFFA93C4CFDE1F93CFD33694F" box="[601,734,1774,1797]" degrees="125" direction="east" minutes="16" orientation="longitude" pageId="18" pageNumber="185" precision="15" seconds="30" value="125.275">125°16'30"E</geoCoordinate>
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||
(
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<specimenCode id="DB8B4957FFA93C4CFD51F93CFF346969" collectionCode="RFS" pageId="18" pageNumber="185">RFS 14-1</specimenCode>
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; coll.
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<collectionCode id="ED3C79E9FFA93C4CFEA1F8DEFEC16969" box="[281,300,1804,1827]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="185" type="Museum">V</collectionCode>
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||
. Kessner,
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<date id="FF93C7ECFFA93C4CFE20F8DEFDF16969" box="[408,540,1804,1827]" pageId="18" pageNumber="185" value="1987-06-14">
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<collectingDate id="EFD73E04FFA93C4CFE20F8DEFDF16969" box="[408,540,1804,1827]" pageId="18" pageNumber="185" value="1987-06-14">14 Jun 1987</collectingDate>
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</date>
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) (
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<figureCitation id="1316FDA9FFA93C4CFD8AF8DEFD9C6969" box="[562,625,1804,1827]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="18" pageNumber="185">Fig. 1</figureCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8B92E12CFFA93C4CFF1EF894FDB16999" blockId="18.[166,794,1862,2003]" pageId="18" pageNumber="185">
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||
<emphasis id="B9593D3EFFA93C4CFF1EF894FEB76914" bold="true" box="[166,346,1862,1886]" pageId="18" pageNumber="185">Type material</emphasis>
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||
.
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||
<materialsCitation id="3B45EB71FFA93C4CFED7F894FE646931" accessionNumber="S34728" collectionCode="WAM" pageId="18" pageNumber="185" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54965F8EFFA93C4CFED7F894FE306917" box="[367,477,1862,1885]" pageId="18" pageNumber="185" type="holotype">Holotype</typeStatus>
|
||
<collectionCode id="ED3C79E9FFA93C4CFE50F894FDC06917" box="[488,557,1862,1885]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="185" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFA93C4CFD80F894FD7F6917" box="[568,658,1862,1885]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34728" pageId="18" pageNumber="185" type="EnaNcbi">S34728</accessionNumber>
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||
(preserved specimen, dissected)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B45EB71FFA93C4CFE2DF8B6FE9469D2" accessionNumber="S34729" collectionCode="WAM" pageId="18" pageNumber="185" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54965F8EFFA93C4CFE2DF8B6FE116931" box="[405,508,1892,1915]" pageId="18" pageNumber="185" type="paratype">Paratypes</typeStatus>
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||
<collectionCode id="ED3C79E9FFA93C4CFDBDF8B6FDAB6931" box="[517,582,1892,1915]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="185" type="Museum">WAM</collectionCode>
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||
<accessionNumber id="947E7CCFFFA93C4CFDF7F8B6FD4C6931" box="[591,673,1892,1915]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34729" pageId="18" pageNumber="185" type="EnaNcbi">S34729</accessionNumber>
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(preserved specimen, juvenile)
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</materialsCitation>
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,
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<materialsCitation id="3B45EB71FFA93C4CFE3CF853FCFB69D2" box="[388,790,1921,1944]" collectionCode="FMNH" pageId="18" pageNumber="185" specimenCode="FMNH 220537" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DB8B4957FFA93C4CFE3CF853FDC569D2" box="[388,552,1921,1944]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="185" type="Museum">FMNH 220537</specimenCode>
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(preserved specimen)
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</materialsCitation>
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,
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<materialsCitation id="3B45EB71FFA93C4CFF1EF84CFF126999" accessionNumber="S34730" collectionCode="AM, WAM" pageId="18" pageNumber="185" specimenCount="1" typeStatus="paratype">
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||
<collectionCode id="ED3C79E9FFA93C4CFF1EF84CFF3C69FF" box="[166,209,1950,1973]" country="Australia" name="Australian Museum" pageId="18" pageNumber="185" type="Museum">AM</collectionCode>
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C.472929 (2 dried shells),
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<collectionCode id="ED3C79E9FFA93C4CFE5EF84CFDCB69FF" box="[486,550,1950,1973]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="185" type="Museum">WAM</collectionCode>
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||
<accessionNumber id="947E7CCFFFA93C4CFD93F84CFD9669FF" box="[555,635,1950,1973]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34730" pageId="18" pageNumber="185" type="EnaNcbi">S34730</accessionNumber>
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(3 dried shells) (Pl. 1.9)
|
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</materialsCitation>
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,
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<materialsCitation id="3B45EB71FFA93C4CFEB2F86EFDB56999" accessionNumber="S34730" box="[266,600,1980,2003]" collectionCode="FMNH" isTailEnumeration="true" pageId="18" pageNumber="185" specimenCode="FMNH 220540" specimenCount="1" typeStatus="paratype">
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||
<specimenCode id="DB8B4957FFA93C4CFEB2F86EFE436999" box="[266,430,1980,2003]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="185" type="Museum">FMNH 220540</specimenCode>
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(6 dried shells)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C337B2A7FFA93C4CFCFFF9ABFA7C6886" pageId="18" pageNumber="185" type="etymology">
|
||
<paragraph id="8B92E12CFFA93C4CFCFFF9ABFA7C6886" blockId="18.[839,1466,1657,1740]" pageId="18" pageNumber="185">
|
||
<emphasis id="B9593D3EFFA93C4CFCFFF9ABFC2F68DB" bold="true" box="[839,962,1657,1681]" pageId="18" pageNumber="185">Etymology</emphasis>
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||
. Species epithet derived from “inusitata” (Latin, unusual, uncommon, extraordinary; of female gender) in reference to its remarkably distinct penial morphology.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C337B2A7FFA93C4DFB8EF922FB7E6D7D" lastPageId="19" lastPageNumber="186" pageId="18" pageNumber="185" type="description">
|
||
<paragraph id="8B92E12CFFA93C4CFB8EF922FB276940" blockId="18.[1078,1226,1776,1802]" box="[1078,1226,1776,1802]" pageId="18" pageNumber="185">
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||
<emphasis id="B9593D3EFFA93C4CFB8EF922FB276940" bold="true" box="[1078,1226,1776,1802]" pageId="18" pageNumber="185">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA93C4DFCFFF8F5FE1C699C" blockId="18.[839,1466,1831,2001]" lastBlockId="19.[121,749,1749,2006]" lastPageId="19" lastPageNumber="186" pageId="18" pageNumber="185">
|
||
<emphasis id="B9593D3EFFA93C4CFCFFF8F5FC936974" box="[839,894,1831,1854]" italics="true" pageId="18" pageNumber="185">Shell</emphasis>
|
||
(Pl. 1.9;
|
||
<figureCitation id="1316FDA9FFA93C4CFC57F8F5FB6E6974" box="[1007,1155,1831,1854]" captionStart="Figure 26" captionStartId="18.[166,229,1475,1496]" captionTargetBox="[166,1466,151,1466]" captionTargetId="figure-96@18.[166,1466,151,1466]" captionTargetPageId="18" captionText="Figure 26. Amplirhagada inusitata n.sp. SEM of shell (AM C.472929), radula and jaw (WAM S34728). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth. (F) Close-up of central and lateral teeth viewed from above. (G) Transition from outer lateral to inner marginal teeth viewed from above. Scale bars: A–B, 200 µm; C, 1 mm; E–G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239600" httpUri="https://zenodo.org/record/5239600/files/figure.png" pageId="18" pageNumber="185">Fig. 26 A–C</figureCitation>
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||
). Broadly conical with low spire. Thin, translucent. Periphery sharply angulate; upper sector of whorls flattened. Umbilicus 30–50% concealed by columellar reflection. Background colour pale yellowish brown; peripheral band diffuse to well developed, moderately broad, brown, visible on visible on last whorl only; subsutural band diffuse to well developed, brown, visible on last whorl only; outer and inner colour like shell. Protoconch
|
||
<quantity id="4CD54CC9FFA83C4DFFC1F8DDFF24696C" box="[121,201,1807,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="186" unit="mm" value="2.6">2.6 mm</quantity>
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||
in diameter, comprising about one whorl, with very indistinct radially elongated pustulations. Teleoconch with faint axial growth lines, evenly distributed across shell surface. Angle of aperture 45–60 degrees; outer lip thin, well rounded, slightly expanded; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 16.8±1.7 × 21.4±
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||
<quantity id="4CD54CC9FFA83C4DFE8AF86DFE6E699C" box="[306,387,1983,2006]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="186" unit="mm" value="0.7">0.7 mm</quantity>
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||
(
|
||
<tableCitation id="C6AFD497FFA83C4DFE29F86DFE0C699C" box="[401,481,1983,2006]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximum–minimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="19" pageNumber="186">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA83C4DFC80FF49FC5A6EA6" blockId="19.[794,1422,154,823]" pageId="19" pageNumber="186">
|
||
<emphasis id="B9593D3EFFA83C4DFC80FF49FB666EF8" box="[824,1163,155,178]" italics="true" pageId="19" pageNumber="186">Radular and jaw morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFA83C4DFB26FF48FADA6EFB" box="[1182,1335,154,177]" captionStart="Figure 26" captionStartId="18.[166,229,1475,1496]" captionTargetBox="[166,1466,151,1466]" captionTargetId="figure-96@18.[166,1466,151,1466]" captionTargetPageId="18" captionText="Figure 26. Amplirhagada inusitata n.sp. SEM of shell (AM C.472929), radula and jaw (WAM S34728). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth. (F) Close-up of central and lateral teeth viewed from above. (G) Transition from outer lateral to inner marginal teeth viewed from above. Scale bars: A–B, 200 µm; C, 1 mm; E–G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239600" httpUri="https://zenodo.org/record/5239600/files/figure.png" pageId="19" pageNumber="186">Fig. 26 D–G</figureCitation>
|
||
). Tooth formula C + 14–19 + 4 +?; 125 rows of teeth (n = 1). Jaw with 10 plates.
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA83C4DFC80FF21FC536D53" blockId="19.[794,1422,154,823]" pageId="19" pageNumber="186">
|
||
<emphasis id="B9593D3EFFA83C4DFC80FF21FBF86F40" box="[824,1045,243,266]" italics="true" pageId="19" pageNumber="186">Genital morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFA83C4DFB9FFF20FB446F43" box="[1063,1193,242,265]" captionStart-0="Figure 27" captionStart-1="Figure 28" captionStartId-0="19.[121,187,643,664]" captionStartId-1="19.[121,184,1633,1654]" captionTargetBox-0="[133,734,201,623]" captionTargetBox-1="[132,744,821,1615]" captionTargetId-0="figure-86@19.[121,748,187,640]" captionTargetId-1="figure-131@19.[121,748,813,1621]" captionTargetPageId-0="19" captionTargetPageId-1="19" captionText-0="Figure 27. Genitalia of Amplirhagada inusitata n.sp. (WAM S34728). For labelling of structures see Fig. 3. Scale bar = 10 mm." captionText-1="Figure 28. Penial anatomy of Amplirhagada inusitata n.sp. (WAM S34728). For labelling of structures see Fig. 4. Scale bar = 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239602" figureDoi-1="http://doi.org/10.5281/zenodo.5239604" httpUri-0="https://zenodo.org/record/5239602/files/figure.png" httpUri-1="https://zenodo.org/record/5239604/files/figure.png" pageId="19" pageNumber="186">Figs 27–28</figureCitation>
|
||
). Penis long, curved to bending, tubular, clearly longer than anterior part of oviduct. Penial sheath very delicate. Penial retractor muscle very short. Penial verge extremely elongated, extending almost entire length of penis, with pointed tip, proximally the lumen of the penial verge forms an extended duct that is connected to vas deferens, possibly functioning as epiphallus. Penial chamber extending only distal half of entire length of penial complex, inner penial wall almost entirely smooth, in proximal third corrugated longitudinal pilasters are present consisting of fused pustules. No main stimulatory pilaster differentiated. Vas deferens rather thin, undulating, entering penial sheath close to penial apex. Vagina moderately long, tubular to slightly inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix reaching base of spermoviduct. Free oviduct long, bending, tubular, comprising more than half of length of anterior part of oviduct. Spermoviduct as long as anterior part of oviduct.
|
||
</paragraph>
|
||
<caption id="DF52B1A4FFA83C4DFFC1FD51FD006CF9" ID-DOI="http://doi.org/10.5281/zenodo.5239602" ID-Zenodo-Dep="5239602" httpUri="https://zenodo.org/record/5239602/files/figure.png" pageId="19" pageNumber="186" startId="19.[121,187,643,664]" targetBox="[133,734,201,623]" targetPageId="19">
|
||
<paragraph id="8B92E12CFFA83C4DFFC1FD51FD006CF9" blockId="19.[121,749,643,691]" pageId="19" pageNumber="186">
|
||
Figure 27. Genitalia of
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFECAFD51FD8B6CD2" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[370,614,643,664]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="species" species="inusitata" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA83C4DFECAFD51FD8B6CD2" box="[370,614,643,664]" italics="true" pageId="19" pageNumber="186">Amplirhagada inusitata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA83C4DFDD7FD51FD726CD2" box="[623,671,643,664]" pageId="19" pageNumber="186" rank="species">n.sp.</taxonomicNameLabel>
|
||
(WAM
|
||
<accessionNumber id="947E7CCFFFA83C4DFFC1FD4CFF2B6CF9" box="[121,198,670,691]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34728" pageId="19" pageNumber="186" type="EnaNcbi">S34728</accessionNumber>
|
||
). For labelling of structures see Fig. 3. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B92E12CFFA83C4DFC80FCF2FB7E6D7D" blockId="19.[794,1422,154,823]" box="[824,1171,800,823]" pageId="19" pageNumber="186">
|
||
<emphasis id="B9593D3EFFA83C4DFC80FCF2FBE66D7D" box="[824,1035,800,823]" italics="true" pageId="19" pageNumber="186">Aestivation strategy</emphasis>
|
||
. Free sealer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337B2A7FFA83C4DFCA2FC88FBEB6A33" pageId="19" pageNumber="186" type="discussion">
|
||
<paragraph id="8B92E12CFFA83C4DFCA2FC88FBEB6A33" blockId="19.[794,1421,858,1145]" pageId="19" pageNumber="186">
|
||
<emphasis id="B9593D3EFFA83C4DFCA2FC88FC6F6D38" bold="true" box="[794,898,858,882]" pageId="19" pageNumber="186">Remarks</emphasis>
|
||
. Anatomical description based on dissection of the
|
||
<typeStatus id="54965F8EFFA83C4DFCFDFCAAFC4A6DC5" box="[837,935,888,911]" pageId="19" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
. Listed by Solem (1991) as “
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFB4AFCAAFCA26DE6" authority="NSP" authorityName="NSP" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9593D3EFFA83C4DFB4AFCAAFA606DC5" box="[1266,1421,888,911]" italics="true" pageId="19" pageNumber="186">Amplirhagada</emphasis>
|
||
NSP
|
||
</taxonomicName>
|
||
26”. Shell similar to other relatively large and broadly conical species with spiral bands, such as
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFB44FC61FC476DAD" authority="Iredale, 1938" authorityName="Iredale" authorityYear="1938" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="species" species="combeana">
|
||
<emphasis id="B9593D3EFFA83C4DFB44FC61FA606D80" box="[1276,1421,947,970]" italics="true" pageId="19" pageNumber="186">A. combeana</emphasis>
|
||
<bibRefCitation id="EFBC9CDDFFA83C4DFCA2FC02FC476DAD" author="Iredale, T" box="[794,938,976,999]" pageId="19" pageNumber="186" pagination="83 - 124" refId="ref19941" refString="Iredale, T. 1938. A basic list of the land Mollusca of Australia - Part III. Australian Zoologist 9: 83 - 124." type="journal article" year="1938">Iredale, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFC5CFC02FACF6DAD" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[996,1314,976,999]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="species" species="mitchelliana">
|
||
<emphasis id="B9593D3EFFA83C4DFC5CFC02FB676DAD" box="[996,1162,976,999]" italics="true" pageId="19" pageNumber="186">A. mitchelliana</emphasis>
|
||
<bibRefCitation id="EFBC9CDDFFA83C4DFB2DFC02FACF6DAD" author="Solem, A" box="[1173,1314,976,999]" pageId="19" pageNumber="186" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, differing from all other congeners by extremely long and tubular penial verge, narrowly elongated penis, smooth penial wall with only short corrugated pilasters at its base. Found in sympatry with
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFC07FB97FB566A16" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[959,1211,1093,1116]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="species" species="pusilla">
|
||
<emphasis id="B9593D3EFFA83C4DFC07FB97FBC46A16" box="[959,1065,1093,1116]" italics="true" pageId="19" pageNumber="186">A. pusilla</emphasis>
|
||
<bibRefCitation id="EFBC9CDDFFA83C4DFB8AFB97FB566A16" author="Solem, A" box="[1074,1211,1093,1116]" pageId="19" pageNumber="186" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2D9AAFFFA83C4DFB4DFB97FA606A16" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[1269,1421,1093,1116]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="186" phylum="Mollusca" rank="species" species="epiphallica" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA83C4DFB4DFB97FA606A16" box="[1269,1421,1093,1116]" italics="true" pageId="19" pageNumber="186">A. epiphallica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA83C4DFCA2FBB0FCA16A33" box="[794,844,1122,1145]" pageId="19" pageNumber="186" rank="species">n.sp.</taxonomicNameLabel>
|
||
described below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |