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<document id="7CBCCDE63694899857B2A971F89E1725" ID-DOI="10.5281/zenodo.207054" ID-GBIF-Dataset="16f22b99-413e-4c26-96b7-db8ff1159d65" ID-ISSN="1175-5326" ID-Zenodo-Dep="207054" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460343357970" checkinUser="plazi" docAuthor="Liu, Yang, Qin, Dao-Zheng, Fletcher, Murray J. &amp; Zhang, Ya-Lin" docDate="2011" docId="038217388C72FFF9B4A4FF560AE6C79E" docLanguage="en" docName="zt03003p042.pdf" docOrigin="Zootaxa 3003" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empoasca (Matsumurasca) schima Thapa 1985" docType="treatment" docVersion="13" lastPageNumber="36" masterDocId="FFBB6F408C7FFFF7B433FFC10F77C571" masterDocTitle="Review of Empoasca (Matsumurasca) Anufriev (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini), with description of three new species from China" masterLastPageNumber="42" masterPageNumber="22" pageNumber="35" updateTime="1698261599315" updateUser="plazi">
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<mods:title id="F16656F5E10D1E85C99913BE26FF237F">Review of Empoasca (Matsumurasca) Anufriev (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini), with description of three new species from China</mods:title>
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<mods:namePart id="FC2D4D5EDFB878709C792F07C0A2D7C3">Liu, Yang</mods:namePart>
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<mods:namePart id="3A52D04B34F162B48C56428194B86E2B">Qin, Dao-Zheng</mods:namePart>
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<mods:namePart id="05AC8B7F01B775B34815C0BBA001C286">Fletcher, Murray J.</mods:namePart>
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<mods:namePart id="C0A865542A24BD2114CCEA52DC572922">Zhang, Ya-Lin</mods:namePart>
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<mods:date id="AD2E5883860BC9AA13051564A8152FC1">2011</mods:date>
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<treatment id="038217388C72FFF9B4A4FF560AE6C79E" ID-DOI="http://doi.org/10.5281/zenodo.6182028" ID-GBIF-Taxon="119406824" ID-Zenodo-Dep="6182028" LSID="urn:lsid:plazi:treatment:038217388C72FFF9B4A4FF560AE6C79E" httpUri="http://treatment.plazi.org/id/038217388C72FFF9B4A4FF560AE6C79E" lastPageId="14" lastPageNumber="36" pageId="13" pageNumber="35">
<subSubSection id="C331F5A58C72FFFAB4A4FF560D95C5C0" box="[151,738,151,177]" pageId="13" pageNumber="35" type="nomenclature">
<paragraph id="8B94A62E8C72FFFAB4A4FF560D95C5C0" blockId="13.[151,738,151,177]" box="[151,738,151,177]" pageId="13" pageNumber="35">
<heading id="D0DC11428C72FFFAB4A4FF560D95C5C0" bold="true" box="[151,738,151,177]" fontSize="11" level="1" pageId="13" pageNumber="35" reason="1">
<taxonomicName id="4C2BDDAD8C72FFFAB4A4FF560D95C5C0" ID-CoL="8J3T5" authority="Thapa, 1985" authorityName="Thapa" authorityYear="1985" box="[151,738,151,177]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="species" species="schima" subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C72FFFAB4A4FF560D95C5C0" bold="true" box="[151,738,151,177]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4A4FF560E63C5C0" bold="true" box="[151,276,151,177]" italics="true" pageId="13" pageNumber="35">Empoasca</emphasis>
(
<emphasis id="B95F7A3C8C72FFFAB516FF560EAEC5C0" bold="true" box="[293,473,151,177]" italics="true" pageId="13" pageNumber="35">Matsumurasca</emphasis>
)
<emphasis id="B95F7A3C8C72FFFAB5D8FF560D36C5C0" bold="true" box="[491,577,151,177]" italics="true" pageId="13" pageNumber="35">schima</emphasis>
<bibRefCitation id="EFBADBDF8C72FFFAB67BFF560D95C5C0" author="Thapa" box="[584,738,151,177]" pageId="13" pageNumber="35" refString="Thapa, V. K. (1985) Some empoascan leafhoppers (Homoptera, Cicadellidae, Typhlocybinae) from the Kathmandu Valley, Nepal, Journal of Entomological Research, 9, 65 - 74." type="journal article" year="1985">Thapa, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331F5A58C72FFFAB4A4FF210C25C463" pageId="13" pageNumber="35" type="reference_group">
<paragraph id="8B94A62E8C72FFFAB4A4FF210D32C585" blockId="13.[151,850,222,275]" box="[151,581,222,245]" pageId="13" pageNumber="35">
<treatmentCitationGroup id="AB3B81008C72FFFAB4A4FF210D32C585" box="[151,581,222,245]" pageId="13" pageNumber="35">
<taxonomicName id="4C2BDDAD8C72FFFAB4A4FF210E76C584" ID-CoL="8HG44" box="[151,257,224,245]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B95F7A3C8C72FFFAB4A4FF210E76C584" box="[151,257,224,245]" italics="true" pageId="13" pageNumber="35">Empoasca</emphasis>
</taxonomicName>
(s. str.)
<taxonomicName id="4C2BDDAD8C72FFFAB560FF1E0D32C585" ID-CoL="8J3T5" authority="Thapa, 1985: 69" authorityName="Thapa" authorityPageNumber="69" authorityYear="1985" box="[339,581,222,245]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="species" species="schima">
<emphasis id="B95F7A3C8C72FFFAB560FF1E0EECC584" box="[339,411,223,245]" italics="true" pageId="13" pageNumber="35">schima</emphasis>
<treatmentCitation id="0A8A803F8C72FFFAB592FF1F0D32C585" author="Thapa" box="[417,581,222,244]" page="69" pageId="13" pageNumber="35" year="1985">
<bibRefCitation id="EFBADBDF8C72FFFAB592FF1F0D55C585" author="Thapa" box="[417,546,222,244]" pageId="13" pageNumber="35" refString="Thapa, V. K. (1985) Some empoascan leafhoppers (Homoptera, Cicadellidae, Typhlocybinae) from the Kathmandu Valley, Nepal, Journal of Entomological Research, 9, 65 - 74." type="journal article" year="1985">Thapa, 1985</bibRefCitation>
: 69
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B94A62E8C72FFFAB4A4FF3F0C25C463" blockId="13.[151,850,222,275]" box="[151,850,253,275]" pageId="13" pageNumber="35">
<treatmentCitationGroup id="AB3B81008C72FFFAB4A4FF3F0C25C463" box="[151,850,253,275]" pageId="13" pageNumber="35">
<taxonomicName id="4C2BDDAD8C72FFFAB4A4FF3F0C25C463" ID-CoL="8J3T5" authority="Thapa, Dworakowska, 1994: 104" authorityName="Thapa, Dworakowska" authorityPageNumber="104" authorityYear="1994" box="[151,850,253,275]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="species" species="schima" subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C72FFFAB4A4FF3F0E76C462" box="[151,257,254,275]" italics="true" pageId="13" pageNumber="35">Empoasca</emphasis>
(
<emphasis id="B95F7A3C8C72FFFAB53CFF3F0ED2C462" box="[271,421,254,275]" italics="true" pageId="13" pageNumber="35">Matsumurasca</emphasis>
)
<emphasis id="B95F7A3C8C72FFFAB587FF3C0E8BC462" box="[436,508,253,275]" italics="true" pageId="13" pageNumber="35">schima</emphasis>
Thapa,
<treatmentCitation id="0A8A803F8C72FFFAB67DFF3C0C25C463" author="Dworakowska" box="[590,850,253,275]" page="104" pageId="13" pageNumber="35" year="1994">
<bibRefCitation id="EFBADBDF8C72FFFAB67DFF3C0C54C463" author="Dworakowska" box="[590,803,253,275]" pageId="13" pageNumber="35" refString="Dworakowska, I. (1994) Typhlocybinae (Auchenorrhyncha: Cicadellidae) of Sikkim, a preliminary survey. Folia Entomologica Hungarica, 55, 93 - 215." type="journal article" year="1994">Dworakowska, 1994</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C331F5A58C72FFF9B4A4FE810CD8C74D" lastPageId="14" lastPageNumber="36" pageId="13" pageNumber="35" type="materials_examined">
<paragraph id="8B94A62E8C72FFFAB4A4FE810D13C40D" blockId="13.[151,1212,319,380]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4A4FE810E42C429" bold="true" box="[151,309,319,344]" pageId="13" pageNumber="35">
<typeStatus id="5490188C8C72FFFAB4A4FE810FA5C429" box="[151,210,320,344]" pageId="13" pageNumber="35">Type</typeStatus>
locality.
</emphasis>
Bhandarkhal, Kathmandu,
<collectingCountry id="F33CE6BE8C72FFFAB659FEFE0DC6C426" box="[618,689,319,343]" name="Nepal" pageId="13" pageNumber="35">Nepal</collectingCountry>
. (Forest Research Institute, Dehra Dun,
<collectingCountry id="F33CE6BE8C72FFFAB041FEFE0BC6C426" box="[1138,1201,319,343]" name="India" pageId="13" pageNumber="35">India</collectingCountry>
).
<emphasis id="B95F7A3C8C72FFFAB4F4FEA20E17C40D" bold="true" box="[199,352,355,380]" pageId="13" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="F33CE6BE8C72FFFAB554FEA50EDAC40D" box="[359,429,356,380]" name="Nepal" pageId="13" pageNumber="35">Nepal</collectingCountry>
,
<collectingCountry id="F33CE6BE8C72FFFAB58BFEA50E86C40D" box="[440,497,356,380]" name="India" pageId="13" pageNumber="35">India</collectingCountry>
(Sikkim).
</paragraph>
<paragraph id="8B94A62E8C72FFFAB4A4FE0E0D56C77B" blockId="13.[151,848,463,522]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4A4FE0E0C27C498" bold="true" box="[151,848,463,489]" pageId="13" pageNumber="35">
<taxonomicName id="4C2BDDAD8C72FFFAB4A4FE0E0D9DC498" authority="Qin and Liu" authorityName="Qin and Liu" box="[151,746,463,489]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="species" species="spinalis" status="sp. nov." subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C72FFFAB4A4FE0E0E63C498" bold="true" box="[151,276,463,489]" italics="true" pageId="13" pageNumber="35">Empoasca</emphasis>
(
<emphasis id="B95F7A3C8C72FFFAB516FE0E0EAEC498" bold="true" box="[293,473,463,489]" italics="true" pageId="13" pageNumber="35">Matsumurasca</emphasis>
)
<emphasis id="B95F7A3C8C72FFFAB5D8FE0E0D3FC498" bold="true" box="[491,584,463,489]" italics="true" pageId="13" pageNumber="35">spinalis</emphasis>
Qin and Liu
</taxonomicName>
,
<taxonomicNameLabel id="A26CC7478C72FFFAB6C5FE100C27C498" box="[758,848,465,489]" pageId="13" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1310BAAB8C72FFFAB4ACFE330E4CC77B" box="[159,315,498,522]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="13" pageNumber="35">Figs 119127</figureCitation>
,
<figureCitation id="1310BAAB8C72FFFAB575FE330E05C77B" box="[326,370,498,522]" captionStart="FIGURES 138 146" captionStartId="16.[151,264,1708,1731]" captionTargetBox="[188,1388,211,1671]" captionTargetId="figure@16.[151,1436,195,1687]" captionTargetPageId="16" captionText="FIGURES 138 146. Faces. 138, E. (M.) biloba Qin &amp; Zhang, 2008; 139, E. (M.) biprocessa Qin &amp; Liu, sp. nov.; 140, E. (M.) clypealata Qin &amp; Zhang, 2008; 141, E. (M.) dentalis Qin &amp; Liu, sp. nov.; 142, E. (M.) diversa Vilbaste, 1968; 143, E. (M.) onukii Matsuda, 1952; 144, E. (M.) parvifacia Dworakowska, 1994; 145, E. (M.) quadrialata Qin &amp; Zhang, 2008; 146, E. (M.) spinalis Qin &amp; Liu, sp. nov." httpUri="https://zenodo.org/record/207069/files/figure.png" pageId="13" pageNumber="35">146</figureCitation>
,
<figureCitation id="1310BAAB8C72FFFAB54DFE330EDDC77B" box="[382,426,498,522]" captionStart="FIGURES 147 155" captionStartId="17.[151,264,1726,1749]" captionTargetBox="[195,1385,199,1694]" captionTargetId="figure@17.[151,1436,193,1705]" captionTargetPageId="17" captionText="FIGURES 147 155. Heads and thoraxes, dorsal view. 147, E. (M.) biloba Qin &amp; Zhang, 2008; 148, E. (M.) biprocessa Qin &amp; Liu, sp. nov.; 149, E. (M.) clypealata Qin &amp; Zhang, 2008; 150, E. (M.) dentalis Qin &amp; Liu, sp. nov.; 151, E. (M.) diversa Vilbaste, 1968; 152, E. (M.) onukii Matsuda, 1952; 153, E. (M.) parvifacia Dworakowska, 1994; 154, E. (M.) quadrialata Qin &amp; Zhang, 2008; 155, E. (M.) spinalis Qin &amp; Liu, sp. nov." httpUri="https://zenodo.org/record/207070/files/figure.png" pageId="13" pageNumber="35">155</figureCitation>
,
<figureCitation id="1310BAAB8C72FFFAB585FE330E95C77B" box="[438,482,498,522]" captionStart="FIGURES 156 164" captionStartId="18.[151,264,1488,1511]" captionTargetBox="[175,1394,195,1449]" captionTargetId="figure@18.[151,1436,193,1467]" captionTargetPageId="18" captionText="FIGURES 156 164. Whole body, dorsal view. 156, E. (M.) biloba Qin &amp; Zhang, 2008; 157, E. (M.) biprocessa Qin &amp; Liu, sp. nov.; 158, E. (M.) clypealata Qin &amp; Zhang, 2008; 159, E. (M.) dentalis Qin &amp; Liu, sp. nov.; 160, E. (M.) diversa Vilbaste, 1968; 161, E. (M.) onukii Matsuda, 1952; 162, E. (M.) parvifacia Dworakowska, 1994; 163, E. (M.) quadrialata Qin &amp; Zhang, 2008; 164, E. (M.) spinalis Qin &amp; Liu, sp. nov." httpUri="https://zenodo.org/record/207071/files/figure.png" pageId="13" pageNumber="35">164</figureCitation>
,
<figureCitation id="1310BAAB8C72FFFAB5DDFE330D6EC77B" box="[494,537,498,522]" captionStart="FIGURES 165 173" captionStartId="19.[151,264,1452,1475]" captionTargetBox="[151,1399,213,1424]" captionTargetId="figure@19.[151,1436,193,1431]" captionTargetPageId="19" captionText="FIGURES 165 173. Whole body, lateral view. 165, E. (M.) biloba Qin &amp; Zhang, 2008; 166, E. (M.) biprocessa Qin &amp; Liu, sp. nov.; 167, E. (M.) clypealata Qin &amp; Zhang, 2008; 168, E. (M.) dentalis Qin &amp; Liu, sp. nov.; 169, E. (M.) diversa Vilbaste, 1968; 170, E. (M.) onukii Matsuda, 1952; 171, E. (M.) parvifacia Dworakowska, 1994; 172, E. (M.) quadrialata Qin &amp; Zhang, 2008; 173, E. (M.) spinalis Qin &amp; Liu, sp. nov." httpUri="https://zenodo.org/record/207072/files/figure.png" pageId="13" pageNumber="35">173</figureCitation>
)
</paragraph>
<paragraph id="8B94A62E8C72FFFAB4A4FDFA0EECC707" blockId="13.[151,1279,570,631]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4A4FDFA0E32C722" bold="true" box="[151,325,570,595]" pageId="13" pageNumber="35">
<typeStatus id="5490188C8C72FFFAB4A4FDFA0FA5C722" box="[151,210,571,595]" pageId="13" pageNumber="35">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5490188C8C72FFFAB57FFDFB0EC0C723" box="[332,439,570,594]" pageId="13" pageNumber="35" type="holotype">Holotype</typeStatus>
, 3 (
<collectionCode id="ED3A3EEB8C72FFFAB5D6FDFB0D38C723" box="[485,591,570,594]" pageId="13" pageNumber="35">NWAFU</collectionCode>
),
<collectingCountry id="F33CE6BE8C72FFFAB651FDFB0DDEC723" box="[610,681,570,594]" name="China" pageId="13" pageNumber="35">China</collectingCountry>
: Mt. Jigong, Henan Prov.,
<date id="FF9580EE8C72FFFAB7E0FDFB0B29C723" box="[979,1118,570,594]" pageId="13" pageNumber="35" value="1997-07-11">11 Jul. 1997</date>
, coll. Jian Hu. Length. 3 3.9 mm.
</paragraph>
<caption id="DF54F6A68C72FFFAB4A4F9B20C00C3B6" httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="13" pageNumber="35" targetBox="[294,1301,705,1611]" targetPageId="13">
<paragraph id="8B94A62E8C72FFFAB4A4F9B20C00C3B6" blockId="13.[151,1436,1651,1735]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4A4F9B20E1AC3F9" bold="true" box="[151,365,1651,1674]" pageId="13" pageNumber="35">FIGURES 119127.</emphasis>
<taxonomicName id="4C2BDDAD8C72FFFAB547F9B50C1EC3F8" authority="Qin and Liu" authorityName="Qin and Liu" box="[372,873,1651,1673]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="35" phylum="Arthropoda" rank="species" species="spinalis" status="sp. nov." subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C72FFFAB547F9B50EAAC3F8" box="[372,477,1652,1673]" italics="true" pageId="13" pageNumber="35">Empoasca</emphasis>
(
<emphasis id="B95F7A3C8C72FFFAB5DEF9B50DF4C3F8" box="[493,643,1652,1673]" italics="true" pageId="13" pageNumber="35">Matsumurasca</emphasis>
)
<emphasis id="B95F7A3C8C72FFFAB6A7F9B20D94C3F8" box="[660,739,1651,1673]" italics="true" pageId="13" pageNumber="35">spinalis</emphasis>
Qin and Liu
</taxonomicName>
,
<emphasis id="B95F7A3C8C72FFFAB747F9B50CB2C3F8" bold="true" box="[884,965,1652,1673]" pageId="13" pageNumber="35">
<taxonomicNameLabel id="A26CC7478C72FFFAB747F9B50CB2C3F8" box="[884,965,1652,1673]" pageId="13" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes.
</paragraph>
</caption>
<paragraph id="8B94A62E8C72FFFAB4F6F92C0FA4C2AE" blockId="13.[151,1437,1773,2015]" pageId="13" pageNumber="35">
<emphasis id="B95F7A3C8C72FFFAB4F6F92C0E79C277" bold="true" box="[197,270,1773,1798]" pageId="13" pageNumber="35">Color.</emphasis>
Vertex cyan, with longitudinal creamy patch along coronal suture not reaching anterior margin, between eyes with creamy streak sublaterally at each side of vertex. Eyes brown, ocelli surrounded by creamy patch mesocaudad. Frontoclypeal area yellowish green, centrally with longitudinal creamy patch in basal half; anteclypeus and genae green. Pronotum bearing irregular creamy patches at anterior margin and arcuate area behind eyes; scutellum with quadrate creamy patch antero-mesally and apicad of scutoscutellar sulcus with an irregular creamy patch. Forewing yellowish green, fore- and hindwing subhyaline. Legs yellowish except tarsi and apical half of hind tibia cyan.
</paragraph>
<paragraph id="8B94A62E8C71FFF9B4F6FF560CD8C74D" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="36">
Abdominal apodemes parallel-sided, rounded apically, surpassing midlength of segment V (
<figureCitation id="1310BAAB8C71FFF9B0D4FF560A3BC5DE" box="[1255,1356,151,175]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Fig. 127</figureCitation>
). Male pygofer slightly elongated and rounded apically, with about 11 stout setae on each side; ventral pygofer appendage nearly straight, directed dorsocaudad and exceeding caudal margin of pygofer lobe, tapering in apical 1/4 to pointed apex (
<figureCitation id="1310BAAB8C71FFF9B50BFEC50EA0C46D" box="[312,471,260,284]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Figs 119, 120</figureCitation>
). Subgenital plate far exceeding pygofer, prominently broadened at base and triangularly protruded basolaterad, apical 1/4 curved dorsocaudad, basal group with 4 long, thick setae, about 23 short marginal microsetae, 2223 long lateral macrosetae and fine microsetae arranged in 2 irregular rows (
<figureCitation id="1310BAAB8C71FFF9B118FE8D0AE7C415" box="[1323,1424,332,356]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Fig. 122</figureCitation>
). Paramere with 5 apical teeth and 2 relatively big teeth subapically preceeded by about 45 setae and few sensory pits (
<figureCitation id="1310BAAB8C71FFF9B4FDFE550E47C4DD" box="[206,304,404,428]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Fig. 126</figureCitation>
). Aedeagal shaft straight in lateral view, nearly as long as preatrium, broad basally and narrowing terminally, with short spinose process baso-ventrally and 2 teeth on dorsal side, dorsal surface in basal 1/3 and ventral surface mostly concave, dorsal apodeme of aedeagus short, in ventral aspect expanded laterad, gonopore ventrad near apex (
<figureCitation id="1310BAAB8C71FFF9B52BFE3E0EC9C766" box="[280,446,511,535]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Figs 123, 124</figureCitation>
). Connective lamellate, caudal margin incised medially (
<figureCitation id="1310BAAB8C71FFF9B062FE3E0BC1C766" box="[1105,1206,511,535]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Fig. 125</figureCitation>
). Anal tube process curved anteroventrad and gradually tapered to pointed apex (
<figureCitation id="1310BAAB8C71FFF9B773FDE50CD5C74D" box="[832,930,548,572]" captionStart="FIGURES 119 127" captionStartId="13.[151,264,1651,1674]" captionTargetBox="[294,1301,705,1611]" captionTargetId="figure@13.[256,1331,684,1630]" captionTargetPageId="13" captionText="FIGURES 119 127. Empoasca (Matsumurasca) spinalis Qin and Liu, sp. nov., 119, male pygofer, lateral view; 120, ventral pygofer appendage; 121, anal tube and anal styli; 122, subgenital plate, lateral view; 123, aedeagus, ventral view; 124, same, lateral view; 125, connective; 126, paramere; 127, abdominal apodemes." httpUri="https://zenodo.org/record/207067/files/figure.png" pageId="14" pageNumber="36">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C331F5A58C71FFF9B4F6FD890ED7C72E" box="[197,416,583,608]" pageId="14" pageNumber="36" type="description">
<paragraph id="8B94A62E8C71FFF9B4F6FD890ED7C72E" blockId="14.[151,1437,151,752]" box="[197,416,583,608]" pageId="14" pageNumber="36">
<emphasis id="B95F7A3C8C71FFF9B4F6FD890E54C711" bold="true" box="[197,291,584,608]" pageId="14" pageNumber="36">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C331F5A58C71FFF9B4F6FDAD0BECC7F5" box="[197,1179,620,644]" pageId="14" pageNumber="36" type="etymology">
<paragraph id="8B94A62E8C71FFF9B4F6FDAD0BECC7F5" blockId="14.[151,1437,151,752]" box="[197,1179,620,644]" pageId="14" pageNumber="36">
<emphasis id="B95F7A3C8C71FFF9B4F6FDAD0E3CC7F5" bold="true" box="[197,331,620,644]" pageId="14" pageNumber="36">Etymology.</emphasis>
The new species name alludes to the spinose process on the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="C331F5A58C71FFF9B4F6FD4E0AE6C79E" pageId="14" pageNumber="36" type="discussion">
<paragraph id="8B94A62E8C71FFF9B4F6FD4E0AE6C79E" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="36">
<emphasis id="B95F7A3C8C71FFF9B4F6FD4E0E3DC7D9" bold="true" box="[197,330,655,680]" pageId="14" pageNumber="36">Discussion.</emphasis>
<taxonomicName id="4C2BDDAD8C71FFF9B561FD500D17C7D9" box="[338,608,655,680]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="36" phylum="Arthropoda" rank="species" species="spinalis" status="sp. nov." subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C71FFF9B561FD500EB0C7D9" box="[338,455,657,680]" italics="true" pageId="14" pageNumber="36">Empoasca</emphasis>
(
<emphasis id="B95F7A3C8C71FFF9B5EAFD500E8FC7D9" box="[473,504,657,680]" italics="true" pageId="14" pageNumber="36">M.</emphasis>
)
<emphasis id="B95F7A3C8C71FFF9B63AFD510D17C7D9" box="[521,608,656,680]" italics="true" pageId="14" pageNumber="36">spinalis</emphasis>
</taxonomicName>
<emphasis id="B95F7A3C8C71FFF9B65AFD510DB3C7D9" bold="true" box="[617,708,656,680]" pageId="14" pageNumber="36">
<taxonomicNameLabel id="A26CC7478C71FFF9B65AFD510DB3C7D9" box="[617,708,656,680]" pageId="14" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C2BDDAD8C71FFF9B76FFD500BC7C7D6" authority="Matsuda, 1952" authorityName="Matsuda" authorityYear="1952" box="[860,1200,655,680]" class="Insecta" family="Cicadellidae" genus="Empoacsa" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="36" phylum="Arthropoda" rank="species" species="onukii" subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C71FFF9B76FFD500C02C7D9" box="[860,885,657,680]" italics="true" pageId="14" pageNumber="36">E.</emphasis>
(
<emphasis id="B95F7A3C8C71FFF9B7B4FD500CD2C7D9" box="[903,933,657,680]" italics="true" pageId="14" pageNumber="36">M.</emphasis>
)
<emphasis id="B95F7A3C8C71FFF9B784FD510C8AC7D9" box="[951,1021,656,680]" italics="true" pageId="14" pageNumber="36">onukii</emphasis>
<bibRefCitation id="EFBADBDF8C71FFF9B036FD4E0BC7C7D6" author="Matsuda" box="[1029,1200,655,679]" pageId="14" pageNumber="36" refString="Matsuda, R. (1952) A new green Empoasca leafhopper injurious to the tea shrub in Japan. Oyo-Kontyu, 8, 19 - 21." type="journal article" year="1952">Matsuda, 1952</bibRefCitation>
</taxonomicName>
, but differs from the latter by the ventral pygofer appendage being straight (sinuate in
<taxonomicName id="4C2BDDAD8C71FFF9B74AFD740B6FC7BD" box="[889,1048,692,716]" class="Insecta" family="Cicadellidae" genus="Empoacsa" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="36" phylum="Arthropoda" rank="species" species="onukii" subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C71FFF9B74AFD740CE5C7BD" box="[889,914,693,716]" italics="true" pageId="14" pageNumber="36">E.</emphasis>
(
<emphasis id="B95F7A3C8C71FFF9B790FD740CB5C7BD" box="[931,962,693,716]" italics="true" pageId="14" pageNumber="36">M.</emphasis>
)
<emphasis id="B95F7A3C8C71FFF9B7E1FD750B6FC7BD" box="[978,1048,692,716]" italics="true" pageId="14" pageNumber="36">onukii</emphasis>
</taxonomicName>
), and by the aedeagal shaft armed with a short spinose process baso-ventrally and 2 dorsal teeth (spinose process and teeth absent in
<taxonomicName id="4C2BDDAD8C71FFF9B0D7FD180AF6C781" box="[1252,1409,727,752]" class="Insecta" family="Cicadellidae" genus="Empoacsa" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="36" phylum="Arthropoda" rank="species" species="onukii" subGenus="Matsumurasca">
<emphasis id="B95F7A3C8C71FFF9B0D7FD180B8BC781" box="[1252,1276,729,752]" italics="true" pageId="14" pageNumber="36">E.</emphasis>
(
<emphasis id="B95F7A3C8C71FFF9B13FFD180A5CC781" box="[1292,1323,729,752]" italics="true" pageId="14" pageNumber="36">M.</emphasis>
)
<emphasis id="B95F7A3C8C71FFF9B108FD190AF6C781" box="[1339,1409,728,752]" italics="true" pageId="14" pageNumber="36">onukii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>