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<mods:title id="66CCD3A74043D5C75BF83E781E73137F">Taxonomic revision of the family Coleophoridae (Lepidoptera: Gelechioidea) in Korea, including seven newly recorded species of the genus Coleophora Hübner</mods:title>
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<mods:namePart id="BA7D0ADAF4E231721D387C50D19DAB19">Koo, Jun-Mo</mods:namePart>
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<mods:affiliation id="873BF4EF3DCAEB537AF552DA31BFD33D">Insect Systematic Bioinformatics Laboratory, Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea &amp; taran 9539 @ gmail. com; https: // orcid. org / 0000 - 0003 - 2639 - 6456</mods:affiliation>
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<mods:namePart id="FC22FC23C44AD90F512C531B1AB26156">Cho, Soowon</mods:namePart>
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<mods:affiliation id="82274107A263E5335A73946DF3F2BA51">Insect Systematic Bioinformatics Laboratory, Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea &amp; chosoowon @ gmail. com; https: // orcid. org / 0000 - 0001 - 6085 - 0064 * Corresponding author</mods:affiliation>
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23.
<taxonomicName id="4C574D19FFD1E04AFF5A0AB2FDCAFE5D" ID-CoL="5ZLVK" authority="Oku, 1974" authorityName="Oku" authorityYear="1974" box="[195,575,474,501]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD1E04AFF5A0AB2FE42FE5C" bold="true" box="[195,439,474,501]" italics="true" pageId="54" pageNumber="55">Coleophora kurokoi</emphasis>
<bibRefCitation id="EFC64B6BFFD1E04AFE270AB2FDCAFE5D" author="Oku, T." box="[446,575,474,500]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku, 1974</bibRefCitation>
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<paragraph id="8BE8369AFFD1E04AFF0E0A97FDBAFDB0" blockId="54.[151,591,474,537]" box="[151,591,511,537]" pageId="54" pageNumber="55">(Korean name: gug-hwa-tong-na-bang)</paragraph>
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<treatmentCitation id="0AF6108BFFD1E04AFF0E092BFDF2FDF2" author="Oku, T." box="[151,519,579,603]" page="256" pageId="54" pageNumber="55" year="1974">
<taxonomicName id="4C574D19FFD1E04AFF0E092BFDF2FDF2" ID-CoL="5ZLVK" authority="Oku, 1974: 256" authorityName="Oku" authorityPageNumber="256" authorityYear="1974" box="[151,519,579,603]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD1E04AFF0E092BFE97FDF2" box="[151,354,579,603]" italics="true" pageId="54" pageNumber="55">Coleophora kurokoi</emphasis>
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<bibRefCitation id="EFC64B6BFFD1E04AFEFE092BFDF2FDF2" author="Oku, T." box="[359,519,579,603]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku, 1974: 256</bibRefCitation>
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. Type locality:
<collectingCountry id="F340760AFFD1E04AFD06092CFD2EFDF2" box="[671,731,580,603]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
,
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,
<collectingMunicipality id="6B8CACE0FFD1E04AFCA6092BFC82FDF2" box="[831,887,579,603]" pageId="54" pageNumber="55">Sakai</collectingMunicipality>
// [depository unconfirmed here:
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, male, bred by
<collectorName id="26A2534CFFD1E04AFF7E090AFE47FDD3" box="[231,434,610,634]" pageId="54" pageNumber="55">Dr. Hiroshi Kuroko</collectorName>
].
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<emphasis id="B923EA88FFD1E04AFF0E09E8FE91FD31" box="[151,356,640,664]" italics="true" pageId="54" pageNumber="55">Coleophora kurokoi</emphasis>
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;
<treatmentCitation id="0AF6108BFFD1E04AFEEB09E8FDC9FD31" author="Moriuti, S." box="[370,572,640,665]" page="264" pageId="54" pageNumber="55" year="1982">
<bibRefCitation id="EFC64B6BFFD1E04AFEEB09E8FDC9FD31" author="Moriuti, S." box="[370,572,640,665]" pageId="54" pageNumber="55" pagination="260 - 265" refId="ref85594" refString="Moriuti, S. (1982 a) Coleophoridae. In: Inoue, H., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. &amp; Owada, M. (Eds.), Moths of Japan. Vol. I. Text. Kodansha, Tokyo, pp. 260 - 265." type="book chapter" year="1982">Moriuti 1982a: 264</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFDDE09E8FD10FD30" author="Vives Moreno, A." box="[583,741,640,665]" page="87" pageId="54" pageNumber="55" year="1988">
<bibRefCitation id="EFC64B6BFFD1E04AFDDE09E8FD10FD30" author="Vives Moreno, A." box="[583,741,640,665]" pageId="54" pageNumber="55" pagination="1 - 196" refId="ref89506" refString="Vives Moreno, A. (1988) Catalogo mundial sistematico y de distribucion de la Familia Coleophoridae Hubner, [1825] (Insecta: Lepidoptera). Boletin de Sanidad Vegetal, Fuera de Serie 12, 1 - 196." type="journal article" year="1988">Vives 1988: 87</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFD6809E8FC14FD30" author="Baldizzone, G." box="[753,993,640,665]" page="208" pageId="54" pageNumber="55" year="1989">
<bibRefCitation id="EFC64B6BFFD1E04AFD6809E8FC14FD30" author="Baldizzone, G." box="[753,993,640,665]" pageId="54" pageNumber="55" pagination="199 - 240" refId="ref77933" refString="Baldizzone, G. (1989 b) A taxonomic review of the Coleophoridae (Lepidoptera) of China. Contribution to the knowledge of the Coleophoridae, LIII. Tijdschrift voor Entomologie, 132, 199 - 240." type="journal article" year="1989">Baldizzone 1989b: 208</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFC7509E8FB11FD30" author="Li, H. H. &amp; Zheng, L. Y." box="[1004,1252,640,665]" page="191" pageId="54" pageNumber="55" year="2000">
<bibRefCitation id="EFC64B6BFFD1E04AFC7509E8FB11FD30" author="Li, H. H. &amp; Zheng, L. Y." box="[1004,1252,640,665]" pageId="54" pageNumber="55" pagination="188 - 192" refId="ref84716" refString="Li, H. H. &amp; Zheng, L. Y. (2000 b) Studies on the Chinese Coleophoridae (Lepidoptera): the Coleophora absinthii group, with description of one new species. Acta Entomologica Sinica, 43 (2), 188 - 192." type="journal article" year="2000">Li &amp; Zheng 2000b: 191</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFB7609E8FED6FD1E" author="Baldizzone, G. &amp; van der Wolf, H. &amp; Landry, J. F." page="72" pageId="54" pageNumber="55" year="2006">
<bibRefCitation id="EFC64B6BFFD1E04AFB7609E8FED6FD1E" author="Baldizzone, G. &amp; van der Wolf, H. &amp; Landry, J. F." pageId="54" pageNumber="55" refId="ref79088" refString="Baldizzone, G., van der Wolf, H. &amp; Landry, J. F. (2006) World catalogue of insects. Vol. 8. Coleophoridae, Coleophorinae (Lepidoptera). Apollo Books, Stenstrup, 215 pp. https: // doi. org / 10.1163 / 9789004475403" type="book" year="2006">
Baldizzone
<emphasis id="B923EA88FFD1E04AFAF009E9FA68FD31" box="[1385,1437,640,664]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2006: 72
</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFEB709F7FE3CFD1E" author="Oku, T." box="[302,457,671,695]" page="240" pageId="54" pageNumber="55" year="2013">
<bibRefCitation id="EFC64B6BFFD1E04AFEB709F7FE3CFD1E" author="Oku, T." box="[302,457,671,695]" pageId="54" pageNumber="55" pagination="231 - 245" refId="ref86438" refString="Oku, T. (2013) Coleophoridae. In: Hirowatari, T., Nasu, Y., Sakamaki, Y. &amp; Kishida, Y. (Eds.), The standard of moths in Japan III. Gakken Education Publishing, Tokyo, pp. 231 - 245." type="book chapter" year="2013">Oku 2013: 240</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFE4A09F7FD40FD1E" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[467,693,671,695]" page="324" pageId="54" pageNumber="55" year="2013">
<bibRefCitation id="EFC64B6BFFD1E04AFE4A09F7FD40FD1E" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[467,693,671,695]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFD9E09C8FDCFFD1E" box="[519,570,671,695]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2013b: 324
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;
<treatmentCitation id="0AF6108BFFD1E04AFD5909F7FC6AFD1E" author="Jinbo, U. &amp; Arita, Y. &amp; Nakajima, H. &amp; Kishida, Y. &amp; Yago, M. &amp; Owada, M." box="[704,927,671,695]" page="153" pageId="54" pageNumber="55" year="2014">
<bibRefCitation id="EFC64B6BFFD1E04AFD5909F7FC6AFD1E" author="Jinbo, U. &amp; Arita, Y. &amp; Nakajima, H. &amp; Kishida, Y. &amp; Yago, M. &amp; Owada, M." box="[704,927,671,695]" pageId="54" pageNumber="55" pagination="129 - 237" refId="ref83180" refString="Jinbo, U., Arita, Y., Nakajima, H., Kishida, Y., Yago, M. &amp; Owada, M. (2014) Moths of Gardens in the Imperial Palace, Tokyo (2009 - 2013). Memoirs of the National Museum of Nature and Science, Tokyo, 50, 129 - 237." type="journal article" year="2014">
Jinbo
<emphasis id="B923EA88FFD1E04AFD6409C8FCC5FD1E" box="[765,816,671,695]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2014: 153
</bibRefCitation>
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;
<treatmentCitation id="0AF6108BFFD1E04AFC3009F7FBCFFD1E" author="Park, K. T." box="[937,1082,671,695]" page="45" pageId="54" pageNumber="55" year="2014">
<bibRefCitation id="EFC64B6BFFD1E04AFC3009F7FBCFFD1E" author="Park, K. T." box="[937,1082,671,695]" pageId="54" pageNumber="55" pagination="2 - 73" refId="ref86592" refString="Park, K. T. (2014) Family Coleophoridae. In: Insect fauna of Korea. Vol. 16 (12). National Institute of Biological Resources, Incheon, pp. 2 - 73." type="book chapter" year="2014">Park 2014: 45</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C34D6511FFD1E04AFF0E098DFC63F9F6" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="8BE8369AFFD1E04AFF0E098DFE03FA96" blockId="54.[151,1437,741,2028]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF0E098DFEE7FD56" bold="true" box="[151,274,741,767]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
The species shares similarities in both external and genital characteristics with
<taxonomicName id="4C574D19FFD1E04AFBE7098EFA6DFD56" authority="Oku, 1974" authorityName="Oku" authorityYear="1974" box="[1150,1432,741,768]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFBE7098EFAEFFD56" box="[1150,1306,741,768]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
<bibRefCitation id="EFC64B6BFFD1E04AFAB9098DFA6DFD56" author="Oku, T." box="[1312,1432,741,767]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku, 1974</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C574D19FFD1E04AFF5E0862FD76FC8A" authority="Oku &amp; Kusunoki, 2018" authorityName="Oku &amp; Kusunoki" authorityYear="2018" box="[199,643,777,804]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="montaniella">
<emphasis id="B923EA88FFD1E04AFF5E0862FE9AFC8A" box="[199,367,777,804]" italics="true" pageId="54" pageNumber="55">C. montaniella</emphasis>
<bibRefCitation id="EFC64B6BFFD1E04AFEEF0861FD76FC8A" author="Oku, T. &amp; Kusunoki, Y." box="[374,643,777,803]" pageId="54" pageNumber="55" pagination="193 - 213" refId="ref86493" refString="Oku, T. &amp; Kusunoki, Y. (2018) New species of the genus Coleophora (Lepidoptera, Coleophoridae) from northern Japan. Tinea, 24 (3), 193 - 213." type="journal article" year="2018">Oku &amp; Kusunoki, 2018</bibRefCitation>
</taxonomicName>
. However, it can be distinguished by the following genital features: 1) in the male genitalia of
<taxonomicName id="4C574D19FFD1E04AFE3F0846FDE9FCEE" authorityName="Oku" authorityYear="1974" box="[422,540,813,840]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD1E04AFE3F0846FDE9FCEE" box="[422,540,813,840]" italics="true" pageId="54" pageNumber="55">C. kurokoi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC64B6BFFD1E04AFDC00845FD38FCEE" author="Oku, T." box="[601,717,813,839]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFD400845FCD4FCEE" box="[729,801,813,839]" captionStart="FIGURE 6" captionStartId="22.[151,250,1542,1566]" captionTargetBox="[162,1425,192,1507]" captionTargetId="figure-31@22.[151,1436,181,1518]" captionTargetPageId="22" captionText="FIGURE 6. Newly recorded Coleophora spp. Abdomen. A, female abdomen of Coleophora clypeiferella, slide no. KJM0228 (CBNUPM000333); B, male abdomen of C. kamchatica, slide no. KJM0157 (CBNUPM000212); C, female abdomen of C. lativittella, slide no. KJM0088 (OONNIN0000015631); D, male abdomen of C. levantis, slide no. KJM0091 (OONNIN0000015632); E, female abdomen of C. levantis, slide no. KJM0420 (in INU)." figureDoi="http://doi.org/10.5281/zenodo.14595197" httpUri="https://zenodo.org/record/14595197/files/figure.png" pageId="54" pageNumber="55">Figs 6</figureCitation>
,
<figureCitation id="136C2A1FFFD1E04AFCB60845FCCAFCEE" box="[815,831,813,839]" captionStart="FIGURE 7" captionStartId="26.[151,250,1584,1608]" captionTargetBox="[162,1425,192,1551]" captionTargetId="figure-31@26.[151,1436,181,1560]" captionTargetPageId="26" captionText="FIGURE 7. Newly recorded Coleophora spp. Abdomen. A, male abdomen of Coleophora parthenica, slide no. KJM0476 (CBNUPM000160); B, female abdomen of C. parthenica, slide no. KJM0477 (CBNUPM000161); C, male abdomen of C. uliginosella, slide no. KJM0461 (CBNUPM000214); D, female abdomen of C. uliginosella, slide no. KJM0462 (CBNUPM000215); E, male abdomen of C. vacciniella, slide no. KJM0366 (CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595199" httpUri="https://zenodo.org/record/14595199/files/figure.png" pageId="54" pageNumber="55">7</figureCitation>
), an elongated sacculus terminating with an upturned thorn-like process featuring a pointed apex and a triangular dorso-basal tooth, slightly curved and sclerotized two juxtal rods, and a cornutus with an irregular basal plate are present, while in
<taxonomicName id="4C574D19FFD1E04AFC6E081EFB61FC26" authorityName="Oku" authorityYear="1974" box="[1015,1172,885,912]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFC6E081EFB61FC26" box="[1015,1172,885,912]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC64B6BFFD1E04AFB49081DFAB1FC26" author="Oku, T." box="[1232,1348,885,911]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFAD6081DFA62FC26" box="[1359,1431,885,911]" captionStart="FIGURE 2" captionStartId="8.[151,250,1606,1630]" captionTargetBox="[161,1424,244,1572]" captionTargetId="figure-31@8.[151,1436,234,1582]" captionTargetPageId="8" captionText="FIGURE 2. Newly recorded Coleophora spp. Venation. A, Coleophora clypeiferella, with close-up of frenulum, slide no. KJM0369 (female, reversed left wings, CBNUPM000333); B, C. kamchatica, with close-up of frenulum, slide no. KJM0478 (male, CBNUPM000212); C, C.lativittella, frenulummissing, slideno.KJM0179(female,OONNIN0000015631);D,C.levantis, frenulum missing, slide no. KJM0178 (male, OONNIN0000015632); E, hindwing of C. levantis, with close-up of frenulum, slide no. KJM0479 (female); F, C. parthenica, with close-up of frenulum, slide no. KJM0484 (female, CBNUPM000161); G, C. uliginosella, with close-up of frenulum, slide no. KJM0482 (female, CBNUPM000215); H, C. vacciniella, with close-up of frenulum, slide no. KJM0483 (male, CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595189" httpUri="https://zenodo.org/record/14595189/files/figure.png" pageId="54" pageNumber="55">Figs 2</figureCitation>
,
<figureCitation id="136C2A1FFFD1E04AFF0E08F1FF53FC1A" box="[151,166,921,947]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="54" pageNumber="55">3</figureCitation>
), male genitalia possess the sacculus terminating in a relatively blunt (or rarely acute) thorn-like process featuring a triangular dorso-basal tooth, slightly curved and dorsally sclerotized two juxtal rods, and a simple (or rarely with a linear lateral lobe) cornutus, and in
<taxonomicName id="4C574D19FFD1E04AFDA1088AFD2AFC52" authorityName="Oku &amp; Kusunoki" authorityYear="2018" box="[568,735,993,1020]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="montaniella">
<emphasis id="B923EA88FFD1E04AFDA1088AFD2AFC52" box="[568,735,993,1020]" italics="true" pageId="54" pageNumber="55">C. montaniella</emphasis>
</taxonomicName>
(
<figureCitation id="136C2A1FFFD1E04AFD740889FC8CFC52" box="[749,889,993,1020]" captionStart="FIGURE 15" captionStartId="59.[151,250,1891,1915]" captionTargetBox="[215,1369,191,1857]" captionTargetId="figure-19@59.[207,1381,181,1867]" captionTargetPageId="59" captionText="FIGURE 15. Reconfirmed Coleophora spp. Male genitalia. A, B, Coleophora melanograpta, slide no. KJM0183 (CBNUPM000090); C, D, C. montaniella, slide no. KJM0354 (CBNUPM000126); E, F, C. obducta, slide no. KJM0299 (CBNUPM000203); G, H, C. quercicola, slide no. KJM0199 (CBNUPM000006); I, J, C. serinipennella, slide no. KJM0277 (CBNUPM000026)." figureDoi="http://doi.org/10.5281/zenodo.14595217" httpUri="https://zenodo.org/record/14595217/files/figure.png" pageId="54" pageNumber="55">Figs 15C, D</figureCitation>
), male genitalia possess the sacculus terminating in a rather pointed thorn-like process featuring two or three triangular teeth, robust one at the dorso-base, another at the ventral half, and the other (sometimes absent) at the ventro-base, slightly curved and sclerotized two juxtal rods, and a cornutus with a linear lateral lobe; 2) in the female genitalia of
<taxonomicName id="4C574D19FFD1E04AFC750F26FB97FBCE" authorityName="Oku" authorityYear="1974" box="[1004,1122,1101,1128]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD1E04AFC750F26FB97FBCE" box="[1004,1122,1101,1128]" italics="true" pageId="54" pageNumber="55">C. kurokoi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC64B6BFFD1E04AFB060F25FAE6FBCE" author="Oku, T." box="[1183,1299,1101,1127]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFA860F25FA96FBCE" box="[1311,1379,1101,1127]" captionStart="FIGURE 8" captionStartId="31.[151,250,1842,1866]" captionTargetBox="[162,1425,192,1807]" captionTargetId="figure-19@31.[151,1436,181,1818]" captionTargetPageId="31" captionText="FIGURE 8. Reconfirmed Coleophora spp. Adult. A, Coleophora adspersella, CBNUPM000378 (male); B, C. albicans, CBNUPM000000381 (male); C, C. cristata, CBNUPM000147 (male); D, C. elodella, CBNUPM000425 (male); E, C. falkovitshella, CBNUPM000229 (female); F, C. fasciella, CBNUPM000336 (male); G, C. flavovena, CBNUPM000097 (female); H, C. fuscocuprella, CBNUPM000023 (male)." figureDoi="http://doi.org/10.5281/zenodo.14595201" httpUri="https://zenodo.org/record/14595201/files/figure.png" pageId="54" pageNumber="55">Fig. 8</figureCitation>
), the ductus bursae with two sclerotized parts, one in the middle, and the other in the anterior 1/5 featuring a small narrow plate, and the corpus bursae with a small scobinate signum are present, while in
<taxonomicName id="4C574D19FFD1E04AFB830FFEFB43FB06" authorityName="Oku" authorityYear="1974" box="[1050,1206,1173,1200]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFB830FFEFB43FB06" box="[1050,1206,1173,1200]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC64B6BFFD1E04AFB6B0FFDFA90FB06" author="Oku, T." box="[1266,1381,1173,1199]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFAE90FFDFF52FB7A" captionStart="FIGURE 4" captionStartId="13.[151,250,1787,1811]" captionTargetBox="[242,1350,191,1754]" captionTargetId="figure-19@13.[228,1359,181,1763]" captionTargetPageId="13" captionText="FIGURE 4. Newly recorded Coleophora spp. Genitalia. A, male genitalia of Coleophora levantis, with close-up of sacculus, slide no. KJM0091 (OONNIN0000015632); B, male genitalia of C. levantis before slide preparation of A; C, phallus of male genitalia of C. levantis, with close-up of cornutus, slide no. KJM0091 (OONNIN0000015632); D, female genitalia of C. levantis, with close-up of sterigma+colliculum (left) and signum (right), slide no. KJM0420 (in INU); E, male genitalia of C. parthenica, slide no. KJM0476 (CBNUPM000160); F, phallus of male genitalia of C. parthenica, slide no. KJM0476 (CBNUPM000160); G, phallus of male genitalia of C. parthenica, slide no. KJM0121 (CBNUPM000018); H, female genitalia of C. parthenica, with close-up of sterigma+colliculum (top) and signum (bottom), slide no. KJM0477 (CBNUPM000161)." figureDoi="http://doi.org/10.5281/zenodo.14595193" httpUri="https://zenodo.org/record/14595193/files/figure.png" pageId="54" pageNumber="55">Fig. 4</figureCitation>
), female genitalia possess the ductus bursae with a sclerotized part in the anterior 2/3 bearing a dentate plate near the anterior end, and the corpus bursae with a small scobinate signum, and in
<taxonomicName id="4C574D19FFD1E04AFB8E0FB6FB4AFB5E" authorityName="Oku &amp; Kusunoki" authorityYear="2018" box="[1047,1215,1245,1272]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="montaniella">
<emphasis id="B923EA88FFD1E04AFB8E0FB6FB4AFB5E" box="[1047,1215,1245,1272]" italics="true" pageId="54" pageNumber="55">C. montaniella</emphasis>
</taxonomicName>
(
<figureCitation id="136C2A1FFFD1E04AFB490FB5FACFFB5E" box="[1232,1338,1245,1271]" captionStart="FIGURE 18" captionStartId="63.[151,250,1908,1932]" captionTargetBox="[206,1382,190,1874]" captionTargetId="figure-19@63.[196,1392,181,1884]" captionTargetPageId="63" captionText="FIGURE 18. Reconfirmed Coleophora spp. Female genitalia (all photographed in dorsal view). A, Coleophora honshuella, slide no. KJM0161 (CBNUPM000104); B, C. mayrella, slide no. KJM0474 (CBNUPM000326); C, C. melanograpta, slide no. KJM0184 (CBNUPM000072); D, C. montaniella, slide no. KJM0421 (CBNUPM000124); E, C. obducta, slide no. KJM0281 (CBNUPM000209)." figureDoi="http://doi.org/10.5281/zenodo.14595223" httpUri="https://zenodo.org/record/14595223/files/figure.png" pageId="54" pageNumber="55">Fig. 18D</figureCitation>
), female genitalia possess the ductus bursae featuring a sclerotized part in the anterior 2/5, and the corpus bursae with a short thorn-like or scobinate signum.
</paragraph>
<paragraph id="8BE8369AFFD1E04AFF5E0E21FBCBFA02" blockId="54.[151,1437,741,2028]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0E21FEF1FACA" box="[199,260,1353,1379]" italics="true" pageId="54" pageNumber="55">Adult</emphasis>
: (
<bibRefCitation id="EFC64B6BFFD1E04AFE820E21FE65FACA" author="Oku, T." box="[283,400,1353,1379]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
: wingspan 9.011.0 mm;
<bibRefCitation id="EFC64B6BFFD1E04AFD2B0E21FCADFACA" author="Moriuti, S." box="[690,856,1353,1379]" pageId="54" pageNumber="55" pagination="260 - 265" refId="ref85594" refString="Moriuti, S. (1982 a) Coleophoridae. In: Inoue, H., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. &amp; Owada, M. (Eds.), Moths of Japan. Vol. I. Text. Kodansha, Tokyo, pp. 260 - 265." type="book chapter" year="1982">Moriuti 1982a</bibRefCitation>
: wingspan 9.011.0 mm;
<bibRefCitation id="EFC64B6BFFD1E04AFBE00E21FB18FACA" author="Oku, T." box="[1145,1261,1353,1379]" pageId="54" pageNumber="55" pagination="231 - 245" refId="ref86438" refString="Oku, T. (2013) Coleophoridae. In: Hirowatari, T., Nasu, Y., Sakamaki, Y. &amp; Kishida, Y. (Eds.), The standard of moths in Japan III. Gakken Education Publishing, Tokyo, pp. 231 - 245." type="book chapter" year="2013">Oku 2013</bibRefCitation>
: wingspan 9.0 11.0 mm). See
<bibRefCitation id="EFC64B6BFFD1E04AFEDE0E05FE0CFA2E" author="Moriuti, S." box="[327,505,1389,1415]" pageId="54" pageNumber="55" refId="ref85660" refString="Moriuti, S. (1982 b) Coleophoridae. In: Inoue, H., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. &amp; Owada, M. (Eds.), Moths of Japan, Vol. II: Plates and Synonymic Catalogue. Kodansha, Tokyo, pls. 12 + 227 + 241 + 260." type="journal volume" year="1982">Moriuti (1982b</bibRefCitation>
: Pl. 12, Fig. 51:
<typeStatus id="54EC8838FFD1E04AFD220E05FCD4FA2E" box="[699,801,1389,1415]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
);
<bibRefCitation id="EFC64B6BFFD1E04AFCAF0E05FC41FA2E" author="Oku, T." box="[822,948,1389,1415]" pageId="54" pageNumber="55" pagination="231 - 245" refId="ref86438" refString="Oku, T. (2013) Coleophoridae. In: Hirowatari, T., Nasu, Y., Sakamaki, Y. &amp; Kishida, Y. (Eds.), The standard of moths in Japan III. Gakken Education Publishing, Tokyo, pp. 231 - 245." type="book chapter" year="2013">Oku (2013</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFC580E05FBFCFA2E" box="[961,1033,1389,1415]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="54" pageNumber="55">Figs 3</figureCitation>
-31-3, 4) [
<figureCitation id="136C2A1FFFD1E04AFBE40E05FB37FA2E" box="[1149,1218,1389,1415]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="54" pageNumber="55">Fig. 3</figureCitation>
-31-3 labeled as
<taxonomicName id="4C574D19FFD1E04AFA1A0E06FEE6FA02" authorityName="Oku" authorityYear="1974" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFA1A0E06FEE6FA02" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
</taxonomicName>
in error];
<bibRefCitation id="EFC64B6BFFD1E04AFE1F0EF9FDA4FA02" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[390,593,1425,1451]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFE260EFAFE02FA02" box="[447,503,1425,1451]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2013b
</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFDC50EF9FD6BFA02" box="[604,670,1425,1451]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="54" pageNumber="55">Fig. 3</figureCitation>
);
<bibRefCitation id="EFC64B6BFFD1E04AFD2C0EF9FCC3FA02" author="Park, K. T." box="[693,822,1425,1451]" pageId="54" pageNumber="55" pagination="2 - 73" refId="ref86592" refString="Park, K. T. (2014) Family Coleophoridae. In: Insect fauna of Korea. Vol. 16 (12). National Institute of Biological Resources, Incheon, pp. 2 - 73." type="book chapter" year="2014">Park (2014</bibRefCitation>
: Pl. for adults:
<figureCitation id="136C2A1FFFD1E04AFC470EF9FBC4FA02" box="[990,1073,1425,1451]" captionStart="FIGURE 18" captionStartId="63.[151,250,1908,1932]" captionTargetBox="[206,1382,190,1874]" captionTargetId="figure-19@63.[196,1392,181,1884]" captionTargetPageId="63" captionText="FIGURE 18. Reconfirmed Coleophora spp. Female genitalia (all photographed in dorsal view). A, Coleophora honshuella, slide no. KJM0161 (CBNUPM000104); B, C. mayrella, slide no. KJM0474 (CBNUPM000326); C, C. melanograpta, slide no. KJM0184 (CBNUPM000072); D, C. montaniella, slide no. KJM0421 (CBNUPM000124); E, C. obducta, slide no. KJM0281 (CBNUPM000209)." figureDoi="http://doi.org/10.5281/zenodo.14595223" httpUri="https://zenodo.org/record/14595223/files/figure.png" pageId="54" pageNumber="55">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BE8369AFFD1E04AFF5E0EDDFD57FA66" blockId="54.[151,1437,741,2028]" box="[199,674,1461,1488]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0EDDFE99FA66" box="[199,364,1461,1487]" italics="true" pageId="54" pageNumber="55">Male genitalia</emphasis>
: See
<bibRefCitation id="EFC64B6BFFD1E04AFE300EDDFDD3FA66" author="Oku, T." box="[425,550,1461,1487]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku (1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFDA80EDDFD8DFA66" box="[561,632,1461,1487]" captionStart="FIGURE 6" captionStartId="22.[151,250,1542,1566]" captionTargetBox="[162,1425,192,1507]" captionTargetId="figure-31@22.[151,1436,181,1518]" captionTargetPageId="22" captionText="FIGURE 6. Newly recorded Coleophora spp. Abdomen. A, female abdomen of Coleophora clypeiferella, slide no. KJM0228 (CBNUPM000333); B, male abdomen of C. kamchatica, slide no. KJM0157 (CBNUPM000212); C, female abdomen of C. lativittella, slide no. KJM0088 (OONNIN0000015631); D, male abdomen of C. levantis, slide no. KJM0091 (OONNIN0000015632); E, female abdomen of C. levantis, slide no. KJM0420 (in INU)." figureDoi="http://doi.org/10.5281/zenodo.14595197" httpUri="https://zenodo.org/record/14595197/files/figure.png" pageId="54" pageNumber="55">Figs 6</figureCitation>
,
<figureCitation id="136C2A1FFFD1E04AFD1D0EDDFD61FA66" box="[644,660,1461,1487]" captionStart="FIGURE 7" captionStartId="26.[151,250,1584,1608]" captionTargetBox="[162,1425,192,1551]" captionTargetId="figure-31@26.[151,1436,181,1560]" captionTargetPageId="26" captionText="FIGURE 7. Newly recorded Coleophora spp. Abdomen. A, male abdomen of Coleophora parthenica, slide no. KJM0476 (CBNUPM000160); B, female abdomen of C. parthenica, slide no. KJM0477 (CBNUPM000161); C, male abdomen of C. uliginosella, slide no. KJM0461 (CBNUPM000214); D, female abdomen of C. uliginosella, slide no. KJM0462 (CBNUPM000215); E, male abdomen of C. vacciniella, slide no. KJM0366 (CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595199" httpUri="https://zenodo.org/record/14595199/files/figure.png" pageId="54" pageNumber="55">7</figureCitation>
).
</paragraph>
<paragraph id="8BE8369AFFD1E04AFF5E0EB1FAD6FA5A" blockId="54.[151,1437,741,2028]" box="[199,1315,1497,1524]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0EB1FE73FA5A" box="[199,390,1497,1523]" italics="true" pageId="54" pageNumber="55">Female genitalia</emphasis>
: See
<bibRefCitation id="EFC64B6BFFD1E04AFE5D0EB1FDB4FA5A" author="Oku, T." box="[452,577,1497,1523]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku (1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFDD20EB1FD7BFA5A" box="[587,654,1497,1523]" captionStart="FIGURE 8" captionStartId="31.[151,250,1842,1866]" captionTargetBox="[162,1425,192,1807]" captionTargetId="figure-19@31.[151,1436,181,1818]" captionTargetPageId="31" captionText="FIGURE 8. Reconfirmed Coleophora spp. Adult. A, Coleophora adspersella, CBNUPM000378 (male); B, C. albicans, CBNUPM000000381 (male); C, C. cristata, CBNUPM000147 (male); D, C. elodella, CBNUPM000425 (male); E, C. falkovitshella, CBNUPM000229 (female); F, C. fasciella, CBNUPM000336 (male); G, C. flavovena, CBNUPM000097 (female); H, C. fuscocuprella, CBNUPM000023 (male)." figureDoi="http://doi.org/10.5281/zenodo.14595201" httpUri="https://zenodo.org/record/14595201/files/figure.png" pageId="54" pageNumber="55">Fig. 8</figureCitation>
);
<bibRefCitation id="EFC64B6BFFD1E04AFD3C0EB1FC85FA5A" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[677,880,1497,1523]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFD440EB2FCE3FA5A" box="[733,790,1497,1523]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2013b
</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFCE20EB1FC1EFA5A" box="[891,1003,1497,1524]" captionStart="FIGURE 7" captionStartId="26.[151,250,1584,1608]" captionTargetBox="[162,1425,192,1551]" captionTargetId="figure-31@26.[151,1436,181,1560]" captionTargetPageId="26" captionText="FIGURE 7. Newly recorded Coleophora spp. Abdomen. A, male abdomen of Coleophora parthenica, slide no. KJM0476 (CBNUPM000160); B, female abdomen of C. parthenica, slide no. KJM0477 (CBNUPM000161); C, male abdomen of C. uliginosella, slide no. KJM0461 (CBNUPM000214); D, female abdomen of C. uliginosella, slide no. KJM0462 (CBNUPM000215); E, male abdomen of C. vacciniella, slide no. KJM0366 (CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595199" httpUri="https://zenodo.org/record/14595199/files/figure.png" pageId="54" pageNumber="55">Figs 7, 7a</figureCitation>
);
<bibRefCitation id="EFC64B6BFFD1E04AFB990EB1FB74FA5A" author="Park, K. T." box="[1024,1153,1497,1523]" pageId="54" pageNumber="55" pagination="2 - 73" refId="ref86592" refString="Park, K. T. (2014) Family Coleophoridae. In: Insect fauna of Korea. Vol. 16 (12). National Institute of Biological Resources, Incheon, pp. 2 - 73." type="book chapter" year="2014">Park (2014</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFB120EB1FAE0FA5A" box="[1163,1301,1497,1523]" captionStart="FIGURE 18" captionStartId="63.[151,250,1908,1932]" captionTargetBox="[206,1382,190,1874]" captionTargetId="figure-19@63.[196,1392,181,1884]" captionTargetPageId="63" captionText="FIGURE 18. Reconfirmed Coleophora spp. Female genitalia (all photographed in dorsal view). A, Coleophora honshuella, slide no. KJM0161 (CBNUPM000104); B, C. mayrella, slide no. KJM0474 (CBNUPM000326); C, C. melanograpta, slide no. KJM0184 (CBNUPM000072); D, C. montaniella, slide no. KJM0421 (CBNUPM000124); E, C. obducta, slide no. KJM0281 (CBNUPM000209)." figureDoi="http://doi.org/10.5281/zenodo.14595223" httpUri="https://zenodo.org/record/14595223/files/figure.png" pageId="54" pageNumber="55">Figs 18A, B</figureCitation>
).
</paragraph>
<paragraph id="8BE8369AFFD1E04AFF5E0E95FC63F9F6" blockId="54.[151,1437,741,2028]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0E95FE2DF9B1" box="[199,472,1533,1560]" italics="true" pageId="54" pageNumber="55">Larval case (leaf miner)</emphasis>
: Tubular silken case (trivalved).
<bibRefCitation id="EFC64B6BFFD1E04AFCD70E95FC25F9BE" author="Oku, T." box="[846,976,1533,1559]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku (1974)</bibRefCitation>
mentioned, without any illustration, that the larval case of this species measures 6.07.0 mm in length while comparing it with that of
<taxonomicName id="4C574D19FFD1E04AFB260D4AFF3AF9F6" authority="Oku, 1974" authorityName="Oku" authorityYear="1974" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFB260D4AFAA8F992" box="[1215,1373,1569,1596]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
<bibRefCitation id="EFC64B6BFFD1E04AFAFC0D49FF3AF9F6" author="Oku, T." pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku, 1974</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFC64B6BFFD1E04AFE930D2DFE88F9F6" author="Oku, T." box="[266,381,1605,1631]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD1E04AFE110D2DFE39F9C9" box="[392,460,1605,1632]" captionStart="FIGURE 5" captionStartId="21.[151,250,1799,1823]" captionTargetBox="[175,1413,191,1765]" captionTargetId="figure-19@21.[164,1423,181,1775]" captionTargetPageId="21" captionText="FIGURE 5. Newly recorded Coleophora spp. Genitalia. A, male genitalia of Coleophora uliginosella, slide no. KJM0461 (CBNUPM000214); B, phallus of male genitalia of C. uliginosella, with close-up of cornutus, slide no. KJM0461 (CBNUPM000214); C, phallus of male genitalia of C. uliginosella, with close-up of cornutus, slide no. KJM0296 (PCQAIN0000012023); D, female genitalia of C. uliginosella, with close-up of sterigma (top) and signum (bottom), slide no. KJM0462 (CBNUPM000215); E, male genitalia of C. vacciniella, with close-up of distal portion of sacculus, slide no. KJM0366 (CBNUPM000385); F, phallus of male genitalia of C. vacciniella, with close-up of cornuti (left) and spines forming a row in distal 1/3 (right), slide no. KJM0366 (CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595195" httpUri="https://zenodo.org/record/14595195/files/figure.png" pageId="54" pageNumber="55">Fig. 5</figureCitation>
, where the case length is about
<quantity id="4CAF9B7FFFD1E04AFCB60D2DFC7CF9F6" box="[815,905,1605,1632]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="54" pageNumber="55" unit="mm" value="7.5">7.5 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D6511FFD1E04AFF5E0D01FB60F90E" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="8BE8369AFFD1E04AFF5E0D01FB60F90E" blockId="54.[151,1437,741,2028]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0D01FEAFF92A" bold="true" box="[199,346,1641,1667]" pageId="54" pageNumber="55">
<collectingRegion id="4993F878FFD1E04AFF5E0D01FF0AF92A" box="[199,255,1641,1667]" country="Afghanistan" name="Khost" pageId="54" pageNumber="55">Host</collectingRegion>
plants.
</emphasis>
[
<taxonomicName id="4C574D19FFD1E04AFEF40D01FE05F92A" box="[365,496,1641,1667]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="family">
<emphasis id="B923EA88FFD1E04AFEF40D01FE05F92A" bold="true" box="[365,496,1641,1667]" pageId="54" pageNumber="55">Asteraceae</emphasis>
</taxonomicName>
]
<taxonomicName id="4C574D19FFD1E04AFD9D0D02FCD8F92D" authority="Pamp." authorityName="Pamp." box="[516,813,1642,1668]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="princeps">
<emphasis id="B923EA88FFD1E04AFD9D0D02FD2EF92A" box="[516,731,1642,1668]" italics="true" pageId="54" pageNumber="55">Artemisia princeps</emphasis>
Pamp.
</taxonomicName>
and
<taxonomicName id="4C574D19FFD1E04AFCF30D01FA68F92A" authority="Ramat. ex Hemsl." authorityName="Hemsl." box="[874,1437,1641,1668]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" isHybrid="true" kingdom="Plantae" order="Asterales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="morifolium">
<emphasis id="B923EA88FFD1E04AFCF30D01FBD7F92A" box="[874,1058,1641,1667]" italics="true" pageId="54" pageNumber="55">Chrysanthemum</emphasis>
×
<emphasis id="B923EA88FFD1E04AFBDF0D01FB37F92A" box="[1094,1218,1641,1667]" italics="true" pageId="54" pageNumber="55">morifolium</emphasis>
Ramat. ex Hemsl.
</taxonomicName>
(recorded as
<taxonomicName id="4C574D19FFD1E04AFEB10DE5FCCCF90E" authority="(Sabine) Makino" authorityName="Makino" baseAuthorityName="Sabine" box="[296,825,1677,1704]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="variety" species="morifolium" variety="sinense">
<emphasis id="B923EA88FFD1E04AFEB10DE5FEA5F90E" box="[296,336,1677,1703]" italics="true" pageId="54" pageNumber="55">Ch.</emphasis>
×
<emphasis id="B923EA88FFD1E04AFEF40DE5FE1CF90E" box="[365,489,1677,1703]" italics="true" pageId="54" pageNumber="55">morifolium</emphasis>
var.
<emphasis id="B923EA88FFD1E04AFDB80DE6FD86F90E" box="[545,627,1678,1703]" italics="true" pageId="54" pageNumber="55">sinense</emphasis>
(Sabine) Makino
</taxonomicName>
) (
<bibRefCitation id="EFC64B6BFFD1E04AFCD60DE5FC36F90E" author="Oku, T." box="[847,963,1677,1703]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku 1974</bibRefCitation>
;
<bibRefCitation id="EFC64B6BFFD1E04AFC560DE5FB7CF90E" author="Falkovitsh, M. I." box="[975,1161,1677,1703]" pageId="54" pageNumber="55" pagination="264 - 286" refId="ref80895" refString="Falkovitsh, M. I. (2006) Host-plant Relationships of the Casebearers (Lepidoptera, Coleophoridae): Communication III. Entomological Review, 86 (3), 264 - 286. https: // doi. org / 10.1134 / S 0013873806030031" type="journal article" year="2006">Falkovitsh 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D6511FFD1E04AFF5E0DD9FA60F962" box="[199,1429,1713,1740]" pageId="54" pageNumber="55" type="distribution">
<paragraph id="8BE8369AFFD1E04AFF5E0DD9FA60F962" blockId="54.[151,1437,741,2028]" box="[199,1429,1713,1740]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0DD9FE94F962" bold="true" box="[199,353,1713,1739]" pageId="54" pageNumber="55">Distribution.</emphasis>
<collectingCountry id="F340760AFFD1E04AFEFE0DD9FE5FF962" box="[359,426,1713,1739]" name="China" pageId="54" pageNumber="55">China</collectingCountry>
(
<collectingRegion id="4993F878FFD1E04AFE200DD9FDE6F962" box="[441,531,1713,1739]" country="China" name="Yunnan" pageId="54" pageNumber="55">Yunnan</collectingRegion>
,
<collectingRegion id="4993F878FFD1E04AFD870DD9FD73F962" box="[542,646,1713,1739]" country="China" name="Zhejiang" pageId="54" pageNumber="55">Zhejiang</collectingRegion>
),
<collectingCountry id="F340760AFFD1E04AFD000DDAFD29F965" box="[665,732,1714,1740]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
,
<collectingCountry id="F340760AFFD1E04AFD7E0DDAFCDEF965" box="[743,811,1714,1740]" name="South Korea" pageId="54" pageNumber="55">Korea</collectingCountry>
(
<bibRefCitation id="EFC64B6BFFD1E04AFCA30DD9FBCFF962" author="Baldizzone, G. &amp; van der Wolf, H. &amp; Landry, J. F." box="[826,1082,1713,1739]" pageId="54" pageNumber="55" refId="ref79088" refString="Baldizzone, G., van der Wolf, H. &amp; Landry, J. F. (2006) World catalogue of insects. Vol. 8. Coleophoridae, Coleophorinae (Lepidoptera). Apollo Books, Stenstrup, 215 pp. https: // doi. org / 10.1163 / 9789004475403" type="book" year="2006">
Baldizzone
<emphasis id="B923EA88FFD1E04AFC590DDAFC0CF962" box="[960,1017,1713,1739]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2006
</bibRefCitation>
,
<bibRefCitation id="EFC64B6BFFD1E04AFBDC0DD9FAF3F962" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[1093,1286,1713,1739]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFBE70DDAFB42F962" box="[1150,1207,1713,1739]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2013b
</bibRefCitation>
;
<bibRefCitation id="EFC64B6BFFD1E04AFA8B0DD9FA7CF962" author="Park, K. T." box="[1298,1417,1713,1739]" pageId="54" pageNumber="55" pagination="2 - 73" refId="ref86592" refString="Park, K. T. (2014) Family Coleophoridae. In: Insect fauna of Korea. Vol. 16 (12). National Institute of Biological Resources, Incheon, pp. 2 - 73." type="book chapter" year="2014">Park 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D6511FFD1E04BFF5E0DBDFC52FCF4" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8BE8369AFFD1E04AFF5E0DBDFE06F8D6" blockId="54.[151,1437,741,2028]" pageId="54" pageNumber="55">
<emphasis id="B923EA88FFD1E04AFF5E0DBDFECEF946" bold="true" box="[199,315,1749,1775]" pageId="54" pageNumber="55">Remarks.</emphasis>
The species was first reported from
<collectingCountry id="F340760AFFD1E04AFD550DBEFCE5F959" box="[716,784,1750,1776]" name="South Korea" pageId="54" pageNumber="55">Korea</collectingCountry>
by
<bibRefCitation id="EFC64B6BFFD1E04AFCA10DBDFBF0F946" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[824,1029,1749,1775]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFCF60DBEFC52F946" box="[879,935,1749,1775]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2013b)
</bibRefCitation>
with
<specimenCount id="9D51FD13FFD1E04AFBDB0DBDFB22F946" box="[1090,1239,1749,1775]" count="3" pageId="54" pageNumber="55" type="female">three females</specimenCount>
[the same in
<bibRefCitation id="EFC64B6BFFD1E04AFAF00DBDFF12F8BA" author="Park, K. T." pageId="54" pageNumber="55" pagination="2 - 73" refId="ref86592" refString="Park, K. T. (2014) Family Coleophoridae. In: Insect fauna of Korea. Vol. 16 (12). National Institute of Biological Resources, Incheon, pp. 2 - 73." type="book chapter" year="2014">Park (2014)</bibRefCitation>
]. Additionally, we collected
<specimenCount id="9D51FD13FFD1E04AFDB40D91FD13F8BA" box="[557,742,1785,1811]" count="3" pageId="54" pageNumber="55" type="generic">three specimens</specimenCount>
,
<specimenCount id="9D51FD13FFD1E04AFD6A0D92FCAEF8BA" box="[755,859,1785,1812]" count="1" pageId="54" pageNumber="55" type="male">one male</specimenCount>
and
<specimenCount id="9D51FD13FFD1E04AFC0F0D92FBD6F8BA" box="[918,1059,1785,1812]" count="2" pageId="54" pageNumber="55" type="female">two females</specimenCount>
, which are well-matched in their barcode sequences with each other (p-distance ranging 0.0%), and the genitalia of
<specimenCount id="9D51FD13FFD1E04AFBD60C76FB2FF89E" box="[1103,1242,1821,1848]" count="2" pageId="54" pageNumber="55" type="female">two females</specimenCount>
morphologically align well with the illustration by
<bibRefCitation id="EFC64B6BFFD1E04AFD960C29FD15F8F2" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[527,736,1857,1883]" pageId="54" pageNumber="55" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD1E04AFDDE0C2AFD8BF8F2" box="[583,638,1857,1883]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2013b)
</bibRefCitation>
. However, the
<specimenCount id="9D51FD13FFD1E04AFC1F0C29FBCCF8F2" box="[902,1081,1857,1883]" count="3" pageId="54" pageNumber="55" type="generic">three specimens</specimenCount>
are not included here due to the confusion below that occurred.
</paragraph>
<paragraph id="8BE8369AFFD1E04BFF5E0CE1FBA1FECC" blockId="54.[151,1437,741,2028]" lastBlockId="55.[151,1437,151,861]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<bibRefCitation id="EFC64B6BFFD1E04AFF5E0CE1FEBCF80A" author="Oku, T." box="[199,329,1929,1955]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku (1974)</bibRefCitation>
described the species only with a diagnosis comparing it with
<taxonomicName id="4C574D19FFD1E04AFB9F0CE2FAD5F80A" authority="Oku, 1974" authorityName="Oku" authorityYear="1974" box="[1030,1312,1929,1956]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFB9F0CE2FB56F80A" box="[1030,1187,1929,1956]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
<bibRefCitation id="EFC64B6BFFD1E04AFB330CE1FAD5F80A" author="Oku, T." box="[1194,1312,1929,1955]" pageId="54" pageNumber="55" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku, 1974</bibRefCitation>
</taxonomicName>
(for which a description was provided) and provided illustrations of genitalia drawn by hand. In the description of the female genitalia of
<taxonomicName id="4C574D19FFD1E04AFEBB0CBAFE37F842" authorityName="Oku" authorityYear="1974" box="[290,450,2001,2028]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD1E04AFEBB0CBAFE37F842" box="[290,450,2001,2028]" italics="true" pageId="54" pageNumber="55">C. yomogiella</emphasis>
</taxonomicName>
, they lack the spinulate section in the ductus bursae, however there is no mention of the presence of the spinulate section in the diagnosis of
<taxonomicName id="4C574D19FFD0E04BFC8A0BFFFC7FFF18" authorityName="Oku" authorityYear="1974" box="[787,906,151,177]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD0E04BFC8A0BFFFC7FFF18" box="[787,906,151,177]" italics="true" pageId="55" pageNumber="56">C. kurokoi</emphasis>
</taxonomicName>
. Considering this, it is inferred that there is no spinulate section in those of
<taxonomicName id="4C574D19FFD0E04BFE4A0BD3FDBDFF7C" authorityName="Oku" authorityYear="1974" box="[467,584,187,213]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kurokoi">
<emphasis id="B923EA88FFD0E04BFE4A0BD3FDBDFF7C" box="[467,584,187,213]" italics="true" pageId="55" pageNumber="56">C. kurokoi</emphasis>
</taxonomicName>
, similar to those of
<taxonomicName id="4C574D19FFD0E04BFCBA0BD3FC4AFF7C" authorityName="Oku" authorityYear="1974" box="[803,959,187,213]" class="Insecta" family="Coleophoridae" genus="Coleophora" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="yomogiella">
<emphasis id="B923EA88FFD0E04BFCBA0BD3FC4AFF7C" box="[803,959,187,213]" italics="true" pageId="55" pageNumber="56">C. yomogiella</emphasis>
</taxonomicName>
. Therefore, upon comparing the illustrated female genitalia by
<bibRefCitation id="EFC64B6BFFD0E04BFEE50BB7FDA5FF50" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[380,592,223,249]" pageId="55" pageNumber="56" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD0E04BFE210BB7FE06FF50" box="[440,499,223,249]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
(2013b
</bibRefCitation>
:
<figureCitation id="136C2A1FFFD0E04BFDC40BB7FD25FF50" box="[605,720,223,249]" captionStart="FIGURE 7" captionStartId="26.[151,250,1584,1608]" captionTargetBox="[162,1425,192,1551]" captionTargetId="figure-31@26.[151,1436,181,1560]" captionTargetPageId="26" captionText="FIGURE 7. Newly recorded Coleophora spp. Abdomen. A, male abdomen of Coleophora parthenica, slide no. KJM0476 (CBNUPM000160); B, female abdomen of C. parthenica, slide no. KJM0477 (CBNUPM000161); C, male abdomen of C. uliginosella, slide no. KJM0461 (CBNUPM000214); D, female abdomen of C. uliginosella, slide no. KJM0462 (CBNUPM000215); E, male abdomen of C. vacciniella, slide no. KJM0366 (CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595199" httpUri="https://zenodo.org/record/14595199/files/figure.png" pageId="55" pageNumber="56">Figs 7, 7a</figureCitation>
) with the diagnosis and illustrations by
<bibRefCitation id="EFC64B6BFFD0E04BFB3B0BB7FAD4FF50" author="Oku, T." box="[1186,1313,223,249]" pageId="55" pageNumber="56" pagination="254 - 257" refId="ref86371" refString="Oku, T. (1974) Two new species of Coleophora (Lepidoptera, Coleophoridae) feeding on Artemisia in Japan. Kontyu, 42, 254 - 257." type="journal article" year="1974">Oku (1974</bibRefCitation>
:
<figureCitation id="136C2A1FFFD0E04BFAB60BB7FA81FF50" box="[1327,1396,223,249]" captionStart="FIGURE 8" captionStartId="31.[151,250,1842,1866]" captionTargetBox="[162,1425,192,1807]" captionTargetId="figure-19@31.[151,1436,181,1818]" captionTargetPageId="31" captionText="FIGURE 8. Reconfirmed Coleophora spp. Adult. A, Coleophora adspersella, CBNUPM000378 (male); B, C. albicans, CBNUPM000000381 (male); C, C. cristata, CBNUPM000147 (male); D, C. elodella, CBNUPM000425 (male); E, C. falkovitshella, CBNUPM000229 (female); F, C. fasciella, CBNUPM000336 (male); G, C. flavovena, CBNUPM000097 (female); H, C. fuscocuprella, CBNUPM000023 (male)." figureDoi="http://doi.org/10.5281/zenodo.14595201" httpUri="https://zenodo.org/record/14595201/files/figure.png" pageId="55" pageNumber="56">Fig. 8</figureCitation>
), it appears that the female genitalia illustrated by Kim
<emphasis id="B923EA88FFD0E04BFD6D0A6BFCDAFEB4" box="[756,815,259,285]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
have a much longer ductus bursae with the spinulate section in the posterior portion, while those illustrated by Oku lack a spinulate section. This suggests the need for further confirmation of the presence/absence of this character in subsequent research.
</paragraph>
<paragraph id="8BE8369AFFD0E04BFF5E0A07FC52FCF4" blockId="55.[151,1437,151,861]" pageId="55" pageNumber="56">
Additionally, the illustration of the female genitalia by
<bibRefCitation id="EFC64B6BFFD0E04BFCBD0A07FC06FE20" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[804,1011,367,393]" pageId="55" pageNumber="56" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD0E04BFCC50A07FC66FE20" box="[860,915,367,393]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
(2013b)
</bibRefCitation>
, including the
<specimenCount id="9D51FD13FFD0E04BFB0D0A07FAB2FE20" box="[1172,1351,367,393]" count="3" pageId="55" pageNumber="56" type="generic">three specimens</specimenCount>
that we collected, appears similar to those of
<taxonomicName id="4C574D19FFD0E04BFDA20AFBFC5BFE04" authority="Hofmann, 1869" authorityName="Hofmann" authorityYear="1869" box="[571,942,403,429]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFDA20AFBFD06FE04" box="[571,755,403,429]" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
<bibRefCitation id="EFC64B6BFFD0E04BFD620AFBFC5BFE04" author="Hofmann, O." box="[763,942,403,429]" pageId="55" pageNumber="56" pagination="107 - 122" refId="ref82642" refString="Hofmann, O. (1869) Beitrage zur Naturgeschichte der Coleophoren. Entomologische Zeitung, Stettin, 30, 107 - 122, 187 - 190." type="journal article" year="1869">Hofmann, 1869</bibRefCitation>
</taxonomicName>
. When comparing the barcode sequences of the three Korean specimens with those of two Finnish specimens of
<taxonomicName id="4C574D19FFD0E04BFC130ADFFBB4FE78" authorityName="Hofmann" authorityYear="1869" box="[906,1089,439,465]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFC130ADFFBB4FE78" box="[906,1089,439,465]" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
</taxonomicName>
from GenBank data (accession nos. HM873152.1, HM872896.1), the p-distance ranges 0.30.4%. However, there are clear morphological differences: 1) in the adults of the Korean specimens (
<bibRefCitation id="EFC64B6BFFD0E04BFD720A97FC59FDB0" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[747,940,511,537]" pageId="55" pageNumber="56" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD0E04BFCBC0A97FCA9FDB0" box="[805,860,511,537]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
2013b
</bibRefCitation>
:
<figureCitation id="136C2A1FFFD0E04BFC2C0A97FC02FDB0" box="[949,1015,511,537]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="55" pageNumber="56">Fig. 3</figureCitation>
), an evenly whitish flagellum without annulations, while in
<taxonomicName id="4C574D19FFD0E04BFE14094BFDB3FD94" authorityName="Hofmann" authorityYear="1869" box="[397,582,547,573]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFE14094BFDB3FD94" box="[397,582,547,573]" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC64B6BFFD0E04BFDCF094BFCEEFD94" author="Baldizzone, G." box="[598,795,547,573]" pageId="55" pageNumber="56" pagination="85 - 94" refId="ref77710" refString="Baldizzone, G. (1984) I Coleophoridae dei boschi di Valmanera (Asti). Contributi alla conoscenza dei Coleophoridae, XXXVIII (Lepidoptera). Rivista Piemontese di Storia Naturale, 5, 85 - 94." type="journal article" year="1984">Baldizzone 1984</bibRefCitation>
: Pl. II,
<figureCitation id="136C2A1FFFD0E04BFCE9094BFC22FD94" box="[880,983,547,573]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="8.[151,250,1606,1630]" captionStartId-1="12.[151,250,1787,1811]" captionStartId-2="13.[151,250,1787,1811]" captionTargetBox-0="[161,1424,244,1572]" captionTargetBox-1="[236,1351,191,1754]" captionTargetBox-2="[242,1350,191,1754]" captionTargetId-0="figure-31@8.[151,1436,234,1582]" captionTargetId-1="figure-31@12.[227,1360,181,1763]" captionTargetId-2="figure-19@13.[228,1359,181,1763]" captionTargetPageId-0="8" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 2. Newly recorded Coleophora spp. Venation. A, Coleophora clypeiferella, with close-up of frenulum, slide no. KJM0369 (female, reversed left wings, CBNUPM000333); B, C. kamchatica, with close-up of frenulum, slide no. KJM0478 (male, CBNUPM000212); C, C.lativittella, frenulummissing, slideno.KJM0179(female,OONNIN0000015631);D,C.levantis, frenulum missing, slide no. KJM0178 (male, OONNIN0000015632); E, hindwing of C. levantis, with close-up of frenulum, slide no. KJM0479 (female); F, C. parthenica, with close-up of frenulum, slide no. KJM0484 (female, CBNUPM000161); G, C. uliginosella, with close-up of frenulum, slide no. KJM0482 (female, CBNUPM000215); H, C. vacciniella, with close-up of frenulum, slide no. KJM0483 (male, CBNUPM000385)." captionText-1="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." captionText-2="FIGURE 4. Newly recorded Coleophora spp. Genitalia. A, male genitalia of Coleophora levantis, with close-up of sacculus, slide no. KJM0091 (OONNIN0000015632); B, male genitalia of C. levantis before slide preparation of A; C, phallus of male genitalia of C. levantis, with close-up of cornutus, slide no. KJM0091 (OONNIN0000015632); D, female genitalia of C. levantis, with close-up of sterigma+colliculum (left) and signum (right), slide no. KJM0420 (in INU); E, male genitalia of C. parthenica, slide no. KJM0476 (CBNUPM000160); F, phallus of male genitalia of C. parthenica, slide no. KJM0476 (CBNUPM000160); G, phallus of male genitalia of C. parthenica, slide no. KJM0121 (CBNUPM000018); H, female genitalia of C. parthenica, with close-up of sterigma+colliculum (top) and signum (bottom), slide no. KJM0477 (CBNUPM000161)." figureDoi-0="http://doi.org/10.5281/zenodo.14595189" figureDoi-1="http://doi.org/10.5281/zenodo.14595191" figureDoi-2="http://doi.org/10.5281/zenodo.14595193" httpUri-0="https://zenodo.org/record/14595189/files/figure.png" httpUri-1="https://zenodo.org/record/14595191/files/figure.png" httpUri-2="https://zenodo.org/record/14595193/files/figure.png" pageId="55" pageNumber="56">Figs 24</figureCitation>
), there are distinct whitish and blackish (or dark brown) annulations on the flagellum; 2) in the female genitalia of the Korean specimens (
<bibRefCitation id="EFC64B6BFFD0E04BFB4E092FFA6CFDC8" author="Kim, M. Y. &amp; Lee, B. W. &amp; Lee, H. S. &amp; Park, K. T." box="[1239,1433,583,609]" pageId="55" pageNumber="56" pagination="321 - 326" refId="ref83408" refString="Kim, M. Y., Lee, B. W., Lee, H. S. &amp; Park, K. T. (2013 b) Eleven species, including three unrecorded species, belonging to Coleophoridae (Lepidoptera) collected from Baengnyeong and Yeonpyeong Islands, Korea. Korean Journal of Applied Entomology, 52 (4), 321 - 326. https: // doi. org / 10.5656 / KSAE. 2013.08.0.036" type="journal article" year="2013">
Kim
<emphasis id="B923EA88FFD0E04BFA88092FFABDFDC8" box="[1297,1352,583,609]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
2013b
</bibRefCitation>
:
<figureCitation id="136C2A1FFFD0E04BFF0E0903FF2EFD2C" box="[151,219,619,645]" captionStart="FIGURE 3" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[236,1351,191,1754]" captionTargetId="figure-31@12.[227,1360,181,1763]" captionTargetPageId="12" captionText="FIGURE 3. Newly recorded Coleophora spp. Genitalia. A, female genitalia of Coleophora clypeiferella, with closeup of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0228 (CBNUPM000333); B, male genitalia of C. kamchatica, with close-up of transtilla bearing small spines (left) and distal portion of sacculus (right), slide no. KJM0157 (CBNUPM000212); C, phallus of male genitalia of C. kamchatica, with close-up of dorsal view of juxtal rods (left) and cornuti (right), slide no. KJM0157 (CBNUPM000212); D, female genitalia of C. lativittella, with close-up of sterigma+colliculum (top), posterior portion of ductus bursae (middle), and signum (bottom), slide no. KJM0088 (OONNIN0000015631)." figureDoi="http://doi.org/10.5281/zenodo.14595191" httpUri="https://zenodo.org/record/14595191/files/figure.png" pageId="55" pageNumber="56">Fig. 3</figureCitation>
), the ductus bursae without a dentate sclerotized plate at the anterior portion, and the corpus bursae with a small scobinate signum, while in
<taxonomicName id="4C574D19FFD0E04BFD8809E7FD3CFD00" authorityName="Hofmann" authorityYear="1869" box="[529,713,655,681]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFD8809E7FD3CFD00" box="[529,713,655,681]" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC64B6BFFD0E04BFD4109E7FC69FD00" author="Baldizzone, G." box="[728,924,655,681]" pageId="55" pageNumber="56" pagination="85 - 94" refId="ref77710" refString="Baldizzone, G. (1984) I Coleophoridae dei boschi di Valmanera (Asti). Contributi alla conoscenza dei Coleophoridae, XXXVIII (Lepidoptera). Rivista Piemontese di Storia Naturale, 5, 85 - 94." type="journal article" year="1984">Baldizzone 1984</bibRefCitation>
: Pl. IV,
<figureCitation id="136C2A1FFFD0E04BFC6C09E7FB83FD00" box="[1013,1142,655,681]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE12" captionStart-3="FIGURE13" captionStartId-0="33.[151,250,1555,1579]" captionStartId-1="40.[151,250,1787,1811]" captionStartId-2="41.[151,250,1666,1690]" captionStartId-3="44.[151,250,1926,1950]" captionTargetBox-0="[162,1425,192,1520]" captionTargetBox-1="[231,1357,190,1754]" captionTargetBox-2="[161,1426,191,1632]" captionTargetBox-3="[162,1425,191,1892]" captionTargetId-0="figure-19@33.[151,1436,181,1531]" captionTargetId-1="figure-31@40.[222,1366,181,1763]" captionTargetId-2="figure-19@41.[151,1436,181,1642]" captionTargetId-3="figure-31@44.[151,1436,181,1902]" captionTargetPageId-0="33" captionTargetPageId-1="40" captionTargetPageId-2="41" captionTargetPageId-3="44" captionText-0="FIGURE 10. Reconfirmed Coleophora spp. Adult. A, Coleophora serinipennella, CBNUPM000373 (male); B, C. silenella, CBNUPM000052 (male); C, C. sternipennella, CBNUPM000247 (female); D, C. therinella, CBNUPM000259 (male); E, C. versurella, CBNUPM000358 (male); F, C. vestianella, CBNUPM000396 (female)." captionText-1="FIGURE 11. Reconfirmed Coleophora spp.Venation.A, Coleophora adspersella, slide no. KJM0381 (male, CBNUPM000378); B, C. albicans, slide no. KJM0519 (male, CBNUPM000381); C, C. cristata, slide no. KJM0494 (female, CBNUPM000145); D, C. elodella, slide no. KJM0489 (male, CBNUPM000121); E, C. falkovitshella, slide no. KJM0508 (female, CBNUPM000229); F, C. fasciella, slide no. KJM0517 (male, holotype, CBNUPM000337); G, C. flavovena, slide no. KJM0487 (female, CBNUPM000098); H, C. fuscocuprella, slide no. KJM0209 (male, reversed left wings, CBNUPM000023); I, C. honshuella, slide no. KJM0165 (female, CBNUPM000104); J, C. kononenkoi, slide no. KJM0506 (male, CBNUPM000224); K, C. mayrella, slide no. KJM0516 (female, CBNUPM000290); L, C. melanograpta, slide no. KJM0486 (male, CBNUPM000090)." captionText-2="FIGURE12. Reconfirmed Coleophora spp.Venation.A, Coleophora montaniella, slideno.KJM0492 (male, CBNUPM000126); B, C. obducta, slide no. KJM0502 (male, CBNUPM000203); C, C. parki, slide no. KJM0541 (female, CBNUPM000395); D, C. quercicola, slide no. KJM0166 (female, CBNUPM000008); E, C. serinipennella, slide no. KJM0339 (female, CBNUPM000027); F, C. silenella, slide no. KJM0485 (male, CBNUPM000051); G, C. sternipennella, slide no. KJM0514 (female, CBNUPM000245); H, C. therinella, slide no. KJM0515 (male, CBNUPM000261); I, C. versurella, slide no. KJM0518 (female, CBNUPM000347); J, C. vestianella, slide no. KJM0507 (male, CBNUPM000228)." captionText-3="FIGURE13.ReconfirmedColeophoraspp.Malegenitalia.A,B,Coleophoraadspersella,slideno.KJM0044(CBNUPM000378); C, D, C. albicans, slide no. KJM0364 (CBNUPM000381); E, F, C. cristata, slide no. KJM0219 (CBNUPM000147); G, H, C. elodella, slide no. KJM0383 (CBNUPM000113); I, J, C. fasciella, slide no. KJM0284 (CBNUPM000334)." figureDoi-0="http://doi.org/10.5281/zenodo.14595205" figureDoi-1="http://doi.org/10.5281/zenodo.14595207" figureDoi-2="http://doi.org/10.5281/zenodo.14595211" figureDoi-3="http://doi.org/10.5281/zenodo.14595213" httpUri-0="https://zenodo.org/record/14595205/files/figure.png" httpUri-1="https://zenodo.org/record/14595207/files/figure.png" httpUri-2="https://zenodo.org/record/14595211/files/figure.png" httpUri-3="https://zenodo.org/record/14595213/files/figure.png" pageId="55" pageNumber="56">Figs 1013</figureCitation>
), the ductus bursae with a dentate sclerotized plate at the anterior portion, and the corpus bursae without any signa [most of the descriptions of the
<taxonomicName id="4C574D19FFD0E04BFF5809BFFE8CFD58" authorityName="Hofmann" authorityYear="1869" box="[193,377,727,753]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFF5809BFFE8CFD58" box="[193,377,727,753]" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
</taxonomicName>
document “signum absent in the corpus bursae”, however, there is a paper by
<bibRefCitation id="EFC64B6BFFD0E04BFB7709BFFEB1FCBC" author="Rynarzewski, T. &amp; Walczak, U." pageId="55" pageNumber="56" pagination="218 - 223" refId="ref87454" refString="Rynarzewski, T. &amp; Walczak, U. (2013) Studies on the Coleophoridae (Lepidoptera) of Poland. IV. Coleophora chrysanthemi HOFMANN, 1869 - a new species to the Polish fauna. Wiadomosci Entomologiczne, 32 (3), 218 - 223." type="journal article" year="2013">Rynarzewski &amp; Walczak (2013</bibRefCitation>
:
<figureCitation id="136C2A1FFFD0E04BFED60993FE67FCBC" box="[335,402,763,789]" captionStart="FIGURE 2" captionStartId="8.[151,250,1606,1630]" captionTargetBox="[161,1424,244,1572]" captionTargetId="figure-31@8.[151,1436,234,1582]" captionTargetPageId="8" captionText="FIGURE 2. Newly recorded Coleophora spp. Venation. A, Coleophora clypeiferella, with close-up of frenulum, slide no. KJM0369 (female, reversed left wings, CBNUPM000333); B, C. kamchatica, with close-up of frenulum, slide no. KJM0478 (male, CBNUPM000212); C, C.lativittella, frenulummissing, slideno.KJM0179(female,OONNIN0000015631);D,C.levantis, frenulum missing, slide no. KJM0178 (male, OONNIN0000015632); E, hindwing of C. levantis, with close-up of frenulum, slide no. KJM0479 (female); F, C. parthenica, with close-up of frenulum, slide no. KJM0484 (female, CBNUPM000161); G, C. uliginosella, with close-up of frenulum, slide no. KJM0482 (female, CBNUPM000215); H, C. vacciniella, with close-up of frenulum, slide no. KJM0483 (male, CBNUPM000385)." figureDoi="http://doi.org/10.5281/zenodo.14595189" httpUri="https://zenodo.org/record/14595189/files/figure.png" pageId="55" pageNumber="56">Fig. 2</figureCitation>
) which shows a small signum in the corpus bursae]. Therefore, it is necessary to include
<taxonomicName id="4C574D19FFD0E04BFA1A0993FEDBFC90" authorityName="Hofmann" authorityYear="1869" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="chrysanthemi">
<emphasis id="B923EA88FFD0E04BFA1A0993FEDBFC90" italics="true" pageId="55" pageNumber="56">C. chrysanthemi</emphasis>
</taxonomicName>
and its allies in further research for clarification, and until this problem is resolved, we have decided to keep the record of this species on the list of Korean
<taxonomicName id="4C574D19FFD0E04BFD63082BFC56FCF4" authorityName="Hubner" authorityYear="1825" box="[762,931,835,861]" class="Insecta" family="Coleophoridae" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="family">Coleophoridae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>