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<document id="B654AB8396C15A608C7E230A112589C9" ID-DOI="10.11646/zootaxa.3681.4.1" ID-GBIF-Dataset="17afa4e8-b105-41f8-bf86-e856fc41a3a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="222798" ID-ZooBank="ABBE4175-C2E8-4BB5-9B61-599BD3D0F632" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460640219132" checkinUser="plazi" docAuthor="Colloff, Matthew J. &amp; Cameron, Stephen L." docDate="2013" docId="03EF6064E46FFFE76AEA0364FD07721E" docLanguage="en" docName="zt03681p346.pdf" docOrigin="Zootaxa 3681 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus Knulle 1957" docType="treatment" docVersion="10" lastPageNumber="306" masterDocId="FFD6181CE46BFFE26A7D0410FF8B764E" masterDocTitle="A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia" masterLastPageNumber="346" masterPageNumber="301" pageNumber="305" updateTime="1698335724857" updateUser="plazi">
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<mods:title id="DE1CF706B980E3BF8050A4BE623E16DD">A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia</mods:title>
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<mods:namePart id="83E5CA359E5F64C9DB4D5219A9ECA12F">Colloff, Matthew J.</mods:namePart>
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<mods:namePart id="24E6D2DC189F36744A48F6C6F5488EAF">Cameron, Stephen L.</mods:namePart>
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<mods:date id="2E6B986AD182A528AB1132B384BCE1C9">2013</mods:date>
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<treatment id="03EF6064E46FFFE76AEA0364FD07721E" ID-DOI="http://doi.org/10.5281/zenodo.6163944" ID-GBIF-Taxon="119571169" ID-Zenodo-Dep="6163944" LSID="urn:lsid:plazi:treatment:03EF6064E46FFFE76AEA0364FD07721E" httpUri="http://treatment.plazi.org/id/03EF6064E46FFFE76AEA0364FD07721E" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
<subSubSection id="C35C82F9E46FFFE66AEA0364FE7A71C0" box="[151,497,1908,1934]" pageId="4" pageNumber="305" type="nomenclature">
<paragraph id="8BF9D172E46FFFE66AEA0364FE7A71C0" blockId="4.[151,497,1908,1934]" box="[151,497,1908,1934]" pageId="4" pageNumber="305">
<heading id="D0B1661EE46FFFE66AEA0364FE7A71C0" bold="true" box="[151,497,1908,1934]" fontSize="11" level="1" pageId="4" pageNumber="305" reason="1">
<taxonomicName id="4C46AAF1E46FFFE66AEA0364FE7A71C0" ID-CoL="84DV" authority="Knulle, 1957" authorityName="Knulle" authorityYear="1957" box="[151,497,1908,1934]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46FFFE66AEA0364FE7A71C0" bold="true" box="[151,497,1908,1934]" pageId="4" pageNumber="305">
<emphasis id="B9320D60E46FFFE66AEA0364FEC771C0" bold="true" box="[151,332,1908,1934]" italics="true" pageId="4" pageNumber="305">Tyrphonothrus</emphasis>
<bibRefCitation id="EFD7AC83E46FFFE66B2E0364FE7A71C0" author="Knulle" box="[339,497,1908,1934]" pageId="4" pageNumber="305" refString="Knulle, W. (1957) Morphologische und entwicklungsgeschichtliche Untersuchungen zum phylogenetischen System der Acari. Acariformes Zachv. I. Oribatei, Malaconothridae. Mitteilungen aus dem zoologischen Museum in Berlin, 33, 97 - 213. http: // dx. doi. org / 10.1002 / mmnz. 19570330103" type="journal article" year="1957">Knülle, 1957</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BF9D172E46FFFE66AEA03ABFD5E719F" blockId="4.[151,1297,1979,2032]" box="[151,725,1979,2001]" pageId="4" pageNumber="305">
<taxonomicName id="4C46AAF1E46FFFE66AEA03ABFD0E719F" authority="Knulle, 1957" authorityName="Knulle" authorityYear="1957" box="[151,645,1979,2001]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="subGenus" subGenus="Tyrphonothrus">
<emphasis id="B9320D60E46FFFE66AEA03ABFEC7719F" box="[151,332,1979,2001]" italics="true" pageId="4" pageNumber="305">Trimalaconothrus</emphasis>
(
<emphasis id="B9320D60E46FFFE66B2603ABFE64719F" box="[347,495,1979,2001]" italics="true" pageId="4" pageNumber="305">Tyrphonothrus</emphasis>
)
<bibRefCitation id="EFD7AC83E46FFFE66B8303ABFD0E719F" author="Knulle" box="[510,645,1979,2001]" pageId="4" pageNumber="305" refString="Knulle, W. (1957) Morphologische und entwicklungsgeschichtliche Untersuchungen zum phylogenetischen System der Acari. Acariformes Zachv. I. Oribatei, Malaconothridae. Mitteilungen aus dem zoologischen Museum in Berlin, 33, 97 - 213. http: // dx. doi. org / 10.1002 / mmnz. 19570330103" type="journal article" year="1957">Knülle, 1957</bibRefCitation>
</taxonomicName>
, p. 160.
</paragraph>
<paragraph id="8BF9D172E46FFFE66AEA03C9FA9A71A1" blockId="4.[151,1297,1979,2032]" box="[151,1297,2009,2032]" pageId="4" pageNumber="305">
Type-species:
<emphasis id="B9320D60E46FFFE66B5503C9FD9471BE" box="[296,543,2009,2032]" italics="true" pageId="4" pageNumber="305">
<taxonomicName id="4C46AAF1E46FFFE66B5503C9FE5671A1" ID-CoL="b8aa36c7-6696-441a-8b43-7105eee47948" box="[296,477,2009,2031]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">Trimalaconothrus</taxonomicName>
novus
</emphasis>
<bibRefCitation id="EFD7AC83E46FFFE6685803C9FD3171A1" author="Sellnick" box="[549,698,2009,2031]" pageId="4" pageNumber="305" refString="Sellnick, M. (1921) Oribatiden von zwergbirkenmoor bei Neulinum, Kr. Kulm, vom moor am kleinen heidsee bei heubude unweit Danzig. Schriften der Naturforschenden Gesellschaft in Danzig (New Series), 15, 69 - 77." type="journal article" year="1921">Sellnick, 1921</bibRefCitation>
, p. 76 (=
<taxonomicName id="4C46AAF1E46FFFE6696703CBFCE671A1" ID-CoL="7X3ZX" box="[794,877,2010,2032]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="maior">
<emphasis id="B9320D60E46FFFE6696703CBFCE671A1" box="[794,877,2010,2032]" italics="true" pageId="4" pageNumber="305">T. maior</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFD7AC83E46FFFE6690603C9FB8771A1" author="Berlese" box="[891,1036,2009,2031]" pageId="4" pageNumber="305" refString="Berlese, A. (1910) Acari nuovi. Manipulus V, VI. Redia, 6, 199 - 234." type="journal article" year="1910">Berlese, 1910</bibRefCitation>
]) by original designation.
</paragraph>
<paragraph id="8BF9D172E46EFFE76AEA0488FC8876E0" blockId="5.[151,961,152,205]" box="[151,771,152,174]" pageId="5" pageNumber="306">
<taxonomicName id="4C46AAF1E46EFFE76AEA0488FE4A76E0" authority="Hammer, 1962" authorityName="Hammer" authorityYear="1962" box="[151,449,152,174]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76AEA0488FEAA76E0" box="[151,289,152,174]" italics="true" pageId="5" pageNumber="306">Fossonothrus</emphasis>
<bibRefCitation id="EFD7AC83E46EFFE76B5A0489FE4A76E0" author="Hammer" box="[295,449,152,174]" pageId="5" pageNumber="306" refString="Hammer, M. (1962) Investigations on the oribatid fauna of the Andes Mountains. III. Chile. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 13 (2), 1 - 96." type="journal article" year="1962">Hammer, 1962</bibRefCitation>
</taxonomicName>
, p. 20; syn.
<bibRefCitation id="EFD7AC83E46EFFE768460488FD4B76E3" author="Subias" box="[571,704,152,174]" pageId="5" pageNumber="306" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (supplement), 3 - 305. http: // dx. doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías, 2004</bibRefCitation>
, p. 62.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E46EFFE76AEA04A7FB91747A" pageId="5" pageNumber="306" type="materials_examined">
<paragraph id="8BF9D172E46EFFE76AEA04A7FC4A7682" blockId="5.[151,961,152,205]" box="[151,961,182,205]" pageId="5" pageNumber="306">
<typeStatus id="54FD6FD0E46EFFE76AEA04A7FF437682" box="[151,200,183,204]" pageId="5" pageNumber="306">Type</typeStatus>
species:
<taxonomicName id="4C46AAF1E46EFFE76B5B04A7FD0C7682" authority="Hammer, 1962" authorityName="Hammer" authorityYear="1962" box="[294,647,182,205]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9320D60E46EFFE76B5B04A7FE6C7683" box="[294,487,183,205]" italics="true" pageId="5" pageNumber="306">Fossonothrus latus</emphasis>
<bibRefCitation id="EFD7AC83E46EFFE76B9004A7FD0C7682" author="Hammer" box="[493,647,182,204]" pageId="5" pageNumber="306" refString="Hammer, M. (1962) Investigations on the oribatid fauna of the Andes Mountains. III. Chile. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 13 (2), 1 - 96." type="journal article" year="1962">Hammer, 1962</bibRefCitation>
</taxonomicName>
, p. 20, by original designation.
</paragraph>
<paragraph id="8BF9D172E46EFFE76AEA04E8FB91747A" blockId="5.[151,1437,248,1104]" pageId="5" pageNumber="306">
<emphasis id="B9320D60E46EFFE76AEA04E8FE43775F" bold="true" box="[151,456,248,273]" pageId="5" pageNumber="306">Definition and diagnosis.</emphasis>
Malaconothrid mites with the following combination of characters. Carina strongly incurved anteriorly, S-shaped, broad, enclosing seta
<emphasis id="B9320D60E46EFFE7689B050DFD71777B" box="[742,762,285,309]" italics="true" pageId="5" pageNumber="306">le</emphasis>
. Seta
<emphasis id="B9320D60E46EFFE7693F050EFCD3777B" box="[834,856,286,309]" italics="true" pageId="5" pageNumber="306">in</emphasis>
typically very long, flagelliform, at least 3–5 × the length of seta
<emphasis id="B9320D60E46EFFE76B430552FEDC7717" box="[318,343,322,345]" italics="true" pageId="5" pageNumber="306">ex</emphasis>
; never short. Caudal region of hysterosoma typically much broader than anterior region. With centrodorsal ridges and M-shaped posterior ridges, or ridges secondarily lost, as indicated by vestigial structures. Anteriolateral margin of epimeral plates I triangular or rounded. Apodeme IV broad, straight, more or less transverse. Genital setae 5–12 pairs. Tarsi, particularly Tarsus I, elongated, at least 2.5 × longer than maximum width. Setae
<emphasis id="B9320D60E46EFFE76B4505C2FECF77A7" box="[312,324,466,489]" italics="true" pageId="5" pageNumber="306">e</emphasis>
2 and
<emphasis id="B9320D60E46EFFE76BEF05C1FE2B77A7" box="[402,416,465,489]" italics="true" pageId="5" pageNumber="306">h</emphasis>
2, and occasionally
<emphasis id="B9320D60E46EFFE768F205C1FD1677A7" box="[655,669,465,489]" italics="true" pageId="5" pageNumber="306">h</emphasis>
1, longer than other notogastral setae in the
<emphasis id="B9320D60E46EFFE76EC305C2FB6977A7" box="[1214,1250,465,489]" italics="true" pageId="5" pageNumber="306">c -h</emphasis>
series. Various combinations of prodorsal, notogastral, epimeral, genital and adanal setae barbed or setae smooth. Setae of tibiae, genuae and femora barbed or smooth; tarsal setae smooth. Pre-tarsi tridactylous.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E46EFFE76ABA0650FD07721E" pageId="5" pageNumber="306" type="discussion">
<paragraph id="8BF9D172E46EFFE76ABA0650FB6874A6" blockId="5.[151,1437,248,1104]" pageId="5" pageNumber="306">
<emphasis id="B9320D60E46EFFE76ABA0650FEB07416" bold="true" box="[199,315,576,600]" pageId="5" pageNumber="306">Remarks.</emphasis>
<bibRefCitation id="EFD7AC83E46EFFE76B3E0651FE737416" author="Hammer" box="[323,504,576,600]" pageId="5" pageNumber="306" refString="Hammer, M. (1962) Investigations on the oribatid fauna of the Andes Mountains. III. Chile. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 13 (2), 1 - 96." type="journal article" year="1962">Hammer (1962)</bibRefCitation>
did not provide a diagnosis for
<taxonomicName id="4C46AAF1E46EFFE769150650FB8B7416" box="[872,1024,576,600]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE769150650FB8B7416" box="[872,1024,576,600]" italics="true" pageId="5" pageNumber="306">Fossonothrus</emphasis>
</taxonomicName>
, having conflated the description of the genus with that of
<taxonomicName id="4C46AAF1E46EFFE76BE90675FE627432" box="[404,489,612,636]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9320D60E46EFFE76BE90675FE627432" box="[404,489,612,636]" italics="true" pageId="5" pageNumber="306">F. latus</emphasis>
</taxonomicName>
, though clearly her intent was to include within
<taxonomicName id="4C46AAF1E46EFFE76E730674FB2D7432" box="[1038,1190,612,636]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76E730674FB2D7432" box="[1038,1190,612,636]" italics="true" pageId="5" pageNumber="306">Fossonothrus</emphasis>
</taxonomicName>
those species with Sshaped carinae, longitudinal ridges on the central notogaster and diagonal ridges on the posterior notogaster, a convex posteriomedian margin to epimere IV and hexagonal genital plates. These are characters shared with
<taxonomicName id="4C46AAF1E46EFFE76AEA06C0FE5D74A6" authority="Knulle, 1957" authorityName="Knulle" authorityYear="1957" box="[151,470,720,744]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76AEA06C0FEB674A6" box="[151,317,720,744]" italics="true" pageId="5" pageNumber="306">Tyrphonothrus</emphasis>
<bibRefCitation id="EFD7AC83E46EFFE76B3906C0FE5D74A6" author="Knulle" box="[324,470,720,744]" pageId="5" pageNumber="306" refString="Knulle, W. (1957) Morphologische und entwicklungsgeschichtliche Untersuchungen zum phylogenetischen System der Acari. Acariformes Zachv. I. Oribatei, Malaconothridae. Mitteilungen aus dem zoologischen Museum in Berlin, 33, 97 - 213. http: // dx. doi. org / 10.1002 / mmnz. 19570330103" type="journal article" year="1957">Knülle, 1957</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C46AAF1E46EFFE7687006C0FD2D74A6" box="[525,678,720,744]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE7687006C0FD2D74A6" box="[525,678,720,744]" italics="true" pageId="5" pageNumber="306">Fossonothrus</emphasis>
</taxonomicName>
was considered its junior synonym (
<bibRefCitation id="EFD7AC83E46EFFE76E3D06C0FB5C74A6" author="Subias" box="[1088,1239,720,744]" pageId="5" pageNumber="306" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (supplement), 3 - 305. http: // dx. doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8BF9D172E46EFFE76ABA06E4FD07721E" blockId="5.[151,1437,248,1104]" pageId="5" pageNumber="306">
<taxonomicName id="4C46AAF1E46EFFE76ABA06E4FEE67542" box="[199,365,756,780]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76ABA06E4FEE67542" box="[199,365,756,780]" italics="true" pageId="5" pageNumber="306">Tyrphonothrus</emphasis>
</taxonomicName>
consists of two clades, with
<taxonomicName id="4C46AAF1E46EFFE768D106E5FC937542" box="[684,792,757,780]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="maior">
<emphasis id="B9320D60E46EFFE768D106E5FC937542" box="[684,792,757,780]" italics="true" pageId="5" pageNumber="306">Ty. maior</emphasis>
</taxonomicName>
(the
<typeStatus id="54FD6FD0E46EFFE7692D06E5FC0B7543" box="[848,896,757,781]" pageId="5" pageNumber="306">type</typeStatus>
of
<taxonomicName id="4C46AAF1E46EFFE769D906E4FBC17542" box="[932,1098,756,780]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE769D906E4FBC17542" box="[932,1098,756,780]" italics="true" pageId="5" pageNumber="306">Tyrphonothrus</emphasis>
</taxonomicName>
) in one and
<taxonomicName id="4C46AAF1E46EFFE76EA906E5FAB87542" box="[1236,1331,756,780]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9320D60E46EFFE76EA906E5FAB87542" box="[1236,1331,756,780]" italics="true" pageId="5" pageNumber="306">Ty. latus</emphasis>
</taxonomicName>
(the
<typeStatus id="54FD6FD0E46EFFE76F1106E5FA177543" box="[1388,1436,757,781]" pageId="5" pageNumber="306">type</typeStatus>
of
<taxonomicName id="4C46AAF1E46EFFE76ACB0708FEC5757E" box="[182,334,792,816]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76ACB0708FEC5757E" box="[182,334,792,816]" italics="true" pageId="5" pageNumber="306">Fossonothrus</emphasis>
</taxonomicName>
) in the other (
<figureCitation id="137DCDF7E46EFFE76B8E0708FDB1757E" box="[499,570,792,816]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1909]" captionTargetBox="[268,1324,208,1836]" captionTargetId="figure@3.[247,1338,200,1846]" captionTargetPageId="3" captionText="FIGURE 2. Cladogram of selected Malaconothridae showing two major clades, Tyrphonothrus and Malaconothrus. Designations of species to nominate subgenera, Trimalaconothrus (Tyrphonothrus) and Malaconothrus (Cristonothrus) follows Subías (2004). Bold type = species with complete notogastral ridges; * = species with partial ridges; M (M), M (C) = monodactylous; T (T), T (Ty) = tridactylous. Percentage probabilities of each node are indicated if other than 100." httpUri="https://zenodo.org/record/222800/files/figure.png" pageId="5" pageNumber="306">Fig. 2</figureCitation>
). It might be seem justifiable to formally recognise the clades as subgenera. However, the characters that separate them relate to the epimeral setal formula and its variations, including the presence or absence of seta
<emphasis id="B9320D60E46EFFE76BA70770FE7D7536" box="[474,502,864,888]" italics="true" pageId="5" pageNumber="306">3a</emphasis>
(absent in the
<taxonomicName id="4C46AAF1E46EFFE768DE0771FC997536" box="[675,786,865,888]" class="Arachnida" family="Malaconothridae" genus="Trimalaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="maior">
<emphasis id="B9320D60E46EFFE768DE0771FC997536" box="[675,786,865,888]" italics="true" pageId="5" pageNumber="306">Ty. maior</emphasis>
</taxonomicName>
clade except in
<taxonomicName id="4C46AAF1E46EFFE769AC0771FBB47536" box="[977,1087,864,888]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B9320D60E46EFFE769AC0771FBB47536" box="[977,1087,864,888]" italics="true" pageId="5" pageNumber="306">Ty glaber</emphasis>
</taxonomicName>
; present in the
<taxonomicName id="4C46AAF1E46EFFE76E880771FADC7536" box="[1269,1367,864,888]" class="Arachnida" family="Malaconothridae" genus="Fossonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9320D60E46EFFE76E880771FADC7536" box="[1269,1367,864,888]" italics="true" pageId="5" pageNumber="306">Ty. latus</emphasis>
</taxonomicName>
clade except in
<taxonomicName id="4C46AAF1E46EFFE76B7E0795FE0B75D2" box="[259,384,900,924]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="sacculus">
<emphasis id="B9320D60E46EFFE76B7E0795FE0B75D2" box="[259,384,900,924]" italics="true" pageId="5" pageNumber="306">T. sacculus</emphasis>
</taxonomicName>
). The epimeral setal formula is a complex character and the absence of any one particular seta may be independent of, or coupled with, the presence or absence of another. Further complicating the issue, the loss of
<emphasis id="B9320D60E46EFFE76A9207DCFE8075AA" box="[239,267,972,996]" italics="true" pageId="5" pageNumber="306">3a</emphasis>
shows a mosaic distribution within the Malaconothroidea, occurring in
<taxonomicName id="4C46AAF1E46EFFE76E2407DCFB7275AA" box="[1113,1273,972,996]" class="Arachnida" family="Trhypochthoniidae" genus="Mucronothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76E2407DCFB7275AA" box="[1113,1273,972,996]" italics="true" pageId="5" pageNumber="306">Mucronothrus</emphasis>
</taxonomicName>
, one clade of
<taxonomicName id="4C46AAF1E46EFFE76AEA07E0FEB67246" box="[151,317,1008,1032]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE76AEA07E0FEB67246" box="[151,317,1008,1032]" italics="true" pageId="5" pageNumber="306">Tyrphonothrus</emphasis>
</taxonomicName>
and in at least eight
<taxonomicName id="4C46AAF1E46EFFE7685B07E0FD5A7246" box="[550,721,1008,1032]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE7685B07E0FD5A7246" box="[550,721,1008,1032]" italics="true" pageId="5" pageNumber="306">Malaconothrus</emphasis>
</taxonomicName>
spp. belonging to different minor clades. Accordingly, it seems wiser not to create subgenera within
<taxonomicName id="4C46AAF1E46EFFE768420004FD6D7262" box="[575,742,1044,1068]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B9320D60E46EFFE768420004FD6D7262" box="[575,742,1044,1068]" italics="true" pageId="5" pageNumber="306">Tyrphonothrus</emphasis>
</taxonomicName>
until a clearer and more detailed analysis of character states separating the two clades can be undertaken.
</paragraph>
</subSubSection>
</treatment>
</document>
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