136 lines
17 KiB
XML
136 lines
17 KiB
XML
<document id="0D8E76BFA3523DC30382956908C90759" ID-DOI="10.11646/zootaxa.5026.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300888" ID-ZooBank="A3AA106D-527A-4934-953C-C2EA746659FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039785980" checkinUser="plazi" docAuthor="Thandar, Ahmed S." docDate="2021" docId="03DC1543FF9E4A08FF08F9CCFADEF87D" docLanguage="en" docName="zootaxa.5026.4.3.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyoninae Panning 1949" docType="treatment" docVersion="4" lastPageNumber="511" masterDocId="FFE56D3BFF9A4A0CFF9FFFD1FFF2FFB9" masterDocTitle="Nomenclatural changes in some sea cucumbers with the erection of a new genus and description of a Thyone? juvenile (? n. sp.) from the Gulf of California (Echinodermata: Holothuroidea: Dendrochirotida)" masterLastPageNumber="526" masterPageNumber="507" pageNumber="511" updateTime="1699139049944" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CABB4F34330B57C25E3E2517F007E5C9">Nomenclatural changes in some sea cucumbers with the erection of a new genus and description of a Thyone? juvenile (? n. sp.) from the Gulf of California (Echinodermata: Holothuroidea: Dendrochirotida)</mods:title>
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<mods:namePart id="E84B9C8ECB79687D0E5750D49EC064A4">Thandar, Ahmed S.</mods:namePart>
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<mods:title id="95FE11FA0F1753FDB591795C95C560E1">Zootaxa</mods:title>
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<mods:date id="217C0BE92FC9DABF26F9CBEC928B3625">2021</mods:date>
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<mods:number id="511E87A86B82E36C2B0EA4E17734C1F4">2021-08-27</mods:number>
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<treatment id="03DC1543FF9E4A08FF08F9CCFADEF87D" ID-DOI="http://doi.org/10.5281/zenodo.5307831" ID-Zenodo-Dep="5307831" LSID="urn:lsid:plazi:treatment:03DC1543FF9E4A08FF08F9CCFADEF87D" httpUri="http://treatment.plazi.org/id/03DC1543FF9E4A08FF08F9CCFADEF87D" lastPageNumber="511" pageId="4" pageNumber="511">
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<subSubSection id="C36FF7DEFF9E4A08FF08F9CCFDACF981" box="[151,606,1565,1592]" pageId="4" pageNumber="511" type="nomenclature">
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<paragraph id="8BCAA455FF9E4A08FF08F9CCFDACF981" blockId="4.[151,606,1565,1592]" box="[151,606,1565,1592]" pageId="4" pageNumber="511">
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<heading id="D0821339FF9E4A08FF08F9CCFDACF981" bold="true" box="[151,606,1565,1592]" fontSize="11" level="1" pageId="4" pageNumber="511" reason="1">
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<emphasis id="B9017847FF9E4A08FF08F9CCFDACF981" bold="true" box="[151,606,1565,1592]" pageId="4" pageNumber="511">
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Subfamily
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<taxonomicName id="4C75DFD6FF9E4A08FE80F9CFFDACF981" authority="Panning, 1949" authorityName="Panning" authorityYear="1949" box="[287,606,1566,1592]" class="Holothuroidea" family="Thyonidae" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="subFamily" subFamily="Thyoninae">
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Thyoninae
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<bibRefCitation id="EFE4D9A4FF9E4A08FE33F9CFFDACF981" author="Panning, A." box="[428,606,1566,1592]" pageId="4" pageNumber="511" pagination="404 - 470" refId="ref9653" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning, 1949</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36FF7DEFF9E4A08FF08F9B7FADEF87D" pageId="4" pageNumber="511" type="discussion">
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<paragraph id="8BCAA455FF9E4A08FF08F9B7FADEF87D" blockId="4.[151,1437,1638,1988]" pageId="4" pageNumber="511">
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<emphasis id="B9017847FF9E4A08FF08F9B7FEF9F939" bold="true" box="[151,267,1638,1664]" pageId="4" pageNumber="511">Remarks.</emphasis>
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<bibRefCitation id="EFE4D9A4FF9E4A08FE8BF9B7FE3AF939" author="Smirnov, A. V." box="[276,456,1638,1664]" pageId="4" pageNumber="511" pagination="793832" refId="ref10054" refString="Smirnov, A. V. (2012) System of Class Holothuroidea. Paleontological Journal, 46 (8), 793832. https: // doi. org / 10.1134 / S 0031030112080126" type="journal article" year="2012">Smirnov (2012)</bibRefCitation>
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raised
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<bibRefCitation id="EFE4D9A4FF9E4A08FD83F9B7FD2CF939" author="Panning, A." box="[540,734,1638,1664]" pageId="4" pageNumber="511" pagination="404 - 470" refId="ref9653" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning’s (1949)</bibRefCitation>
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subfamily
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<taxonomicName id="4C75DFD6FF9E4A08FCC0F9B7FC25F939" authorityName="Panning" authorityYear="1949" box="[863,983,1638,1664]" class="Holothuroidea" family="Thyonidae" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="subFamily" subFamily="Thyoninae">Thyoninae</taxonomicName>
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to family rank and this classification is here utilised. Within the family three species currently classified in the genus
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<taxonomicName id="4C75DFD6FF9E4A08FB94F95BFBBDF91D" authorityName="Selenka" authorityYear="1867" box="[1035,1103,1674,1700]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="genus">
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<emphasis id="B9017847FF9E4A08FB94F95BFBBDF91D" box="[1035,1103,1674,1700]" italics="true" pageId="4" pageNumber="511">Stolus</emphasis>
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</taxonomicName>
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(sensu
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<bibRefCitation id="EFE4D9A4FF9E4A08FB38F95BFAB6F91D" author="Panning, A." box="[1191,1348,1674,1700]" pageId="4" pageNumber="511" pagination="404 - 470" refId="ref9653" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning 1949</bibRefCitation>
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) do not strictly belong in it. These include the Caribbean
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<taxonomicName id="4C75DFD6FF9E4A08FD57F97FFBC8F970" authority="(Lampert, 1885)" baseAuthorityName="Lampert" baseAuthorityYear="1885" box="[712,1082,1710,1737]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="cognatus">
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<emphasis id="B9017847FF9E4A08FD57F97FFC8BF971" box="[712,889,1710,1736]" italics="true" pageId="4" pageNumber="511">Stolus cognatus</emphasis>
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(
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<bibRefCitation id="EFE4D9A4FF9E4A08FC16F97FFBC0F970" author="Lampert, K." box="[905,1074,1710,1737]" pageId="4" pageNumber="511" refId="ref8958" refString="Lampert, K. (1885) Die Seewalzen, eine Systematische Monographie mit Bestimmungs und Verbreitungs Tabellen. In: Semper, C. (Ed.), Zweiter teil. Wissenschaftliche Resultate, Wiesbaden, 4 (3), 311 pp., 1 pl." type="book" year="1885">Lampert, 1885</bibRefCitation>
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)
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</taxonomicName>
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with usually equal or subequal tentacles and simple, smooth plates in the body wall, and
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<taxonomicName id="4C75DFD6FF9E4A08FC8AF903FB62F955" authority="(Koehler & Vaney, 1908)" baseAuthorityName="Koehler & Vaney" baseAuthorityYear="1908" box="[789,1168,1746,1772]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="rapax">
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<emphasis id="B9017847FF9E4A08FC8AF903FC80F955" box="[789,882,1746,1772]" italics="true" pageId="4" pageNumber="511">S. rapax</emphasis>
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(
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<bibRefCitation id="EFE4D9A4FF9E4A08FC1FF903FB7AF955" author="Koehler, R. & Vaney, C." box="[896,1160,1746,1772]" pageId="4" pageNumber="511" refId="ref8875" refString="Koehler, R. & Vaney, C. (1908) Holothuries recuielles par l'INVESTIGATOR dans l'Ocean Indien. II. Les Holothuries Littorales. Trustees Indian Museum, Calcutta, 54 pp., 3 pls." type="book" year="1908">Koehler & Vaney, 1908</bibRefCitation>
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)
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</taxonomicName>
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from the Bay of Bengal and
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<taxonomicName id="4C75DFD6FF9E4A08FF5AF927FDB2F8A9" authority="Rajpal & Thandar, 1999" authorityName="Rajpal & Thandar" authorityYear="1999" box="[197,576,1782,1808]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="kilberti">
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<emphasis id="B9017847FF9E4A08FF5AF927FEC3F8A9" box="[197,305,1782,1808]" italics="true" pageId="4" pageNumber="511">S. kilberti</emphasis>
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<bibRefCitation id="EFE4D9A4FF9E4A08FEA9F927FDB2F8A9" author="Rajpal, V. & Thandar, AS" box="[310,576,1782,1808]" pageId="4" pageNumber="511" pagination="130 - 134" refId="ref9867" refString="Rajpal, V. & Thandar, AS. (1999) Stolus kilberti, a new species from the east coast of South Africa (Echinodermata: Holothuroidea: Dendrochirotida) with a key to the genus Stolus Selenka. South African Journal Zoology, 34, 130 - 134. https: // doi. org / 10.1080 / 02541858.1999.11448498" type="journal article" year="1999">Rajpal & Thandar, 1999</bibRefCitation>
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</taxonomicName>
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from the east coast of
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<collectingCountry id="F362E4C5FF9E4A08FCA9F927FC37F8A9" box="[822,965,1782,1808]" name="South Africa" pageId="4" pageNumber="511">South Africa</collectingCountry>
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, both with smooth, reticulated plates in the body wall. While the writer was tempted to revive
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<bibRefCitation id="EFE4D9A4FF9E4A08FD4DF8CBFC4AF88C" author="Deichmann, E." box="[722,952,1818,1845]" pageId="4" pageNumber="511" pagination="381 - 410" refId="ref8576" refString="Deichmann, E. (1954) The Holothurians of the Gulf of Mexico. In: Galtsoff, P. S. (Coord.), The Gulf of Mexico, its origin, waters, and marine life, Bulletin US Fisheries Commission, 55. pp. 381 - 410." type="journal article" year="1954">Deichmann’s (1954)</bibRefCitation>
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genus
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<taxonomicName id="4C75DFD6FF9E4A08FB97F8CBFB89F88D" authorityName="Deichmann" authorityYear="1954" box="[1032,1147,1818,1844]" class="Holothuroidea" family="Phyllophoridae" genus="Thyoneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="genus">
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<emphasis id="B9017847FF9E4A08FB97F8CBFB89F88D" box="[1032,1147,1818,1844]" italics="true" pageId="4" pageNumber="511">Thyoneria</emphasis>
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</taxonomicName>
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(unaccepted by WoRMS, 732056, accessed
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<date id="FFCB8295FF9E4A08FEFDF8EFFE0AF8E1" box="[354,504,1854,1880]" pageId="4" pageNumber="511" value="2021-06-16">16 June 2021</date>
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) for
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<taxonomicName id="4C75DFD6FF9E4A08FDB0F8EFFD42F8E1" baseAuthorityName="Lampert" baseAuthorityYear="1885" box="[559,688,1854,1880]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="cognatus">
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<emphasis id="B9017847FF9E4A08FDB0F8EFFD42F8E1" box="[559,688,1854,1880]" italics="true" pageId="4" pageNumber="511">S. cognatus</emphasis>
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</taxonomicName>
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this step is here not taken (see Remarks under this species) but the latter two species are here assigned to a new genus
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<taxonomicName id="4C75DFD6FF9E4A08FD43F8B3FC82F8C5" authority="In" authorityName="Thandar" authorityYear="2021" box="[732,880,1890,1916]" class="Holothuroidea" family="Phyllophoridae" genus="Pseudostolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="genus" status="gen. nov.">
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<emphasis id="B9017847FF9E4A08FD43F8B3FC82F8C5" box="[732,880,1890,1916]" italics="true" pageId="4" pageNumber="511">Pseudostolus</emphasis>
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</taxonomicName>
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<emphasis id="B9017847FF9E4A08FCE8F8B3FC34F8C5" bold="true" box="[887,966,1890,1916]" pageId="4" pageNumber="511">
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<taxonomicNameLabel id="A232C53CFF9E4A08FCE8F8B3FC34F8C5" box="[887,966,1890,1916]" pageId="4" pageNumber="511" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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In addition, three other species, currently standing in
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<taxonomicName id="4C75DFD6FF9E4A08FE82F857FE9DF819" authorityName="Oken" authorityYear="1815" box="[285,367,1926,1952]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="genus">
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<emphasis id="B9017847FF9E4A08FE82F857FE9DF819" box="[285,367,1926,1952]" italics="true" pageId="4" pageNumber="511">Thyone</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C75DFD6FF9E4A08FE1EF856FD09F819" authority="Deichmann, 1930" authorityName="Deichmann" authorityYear="1930" box="[385,763,1926,1952]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="pseudofusus">
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<emphasis id="B9017847FF9E4A08FE1EF856FDD5F819" box="[385,551,1926,1952]" italics="true" pageId="4" pageNumber="511">T. pseudofusus</emphasis>
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<bibRefCitation id="EFE4D9A4FF9E4A08FDAFF857FD09F819" author="Deichmann, E." box="[560,763,1926,1952]" pageId="4" pageNumber="511" pagination="41 - 226" refId="ref8369" refString="Deichmann, E. (1930) The holothurians of the western part of the Atlantic Ocean. Bulletin Museum Comparative Zoology, Harvard, 71 (3), 41 - 226, 24 pls." type="journal article" year="1930">Deichmann, 1930</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="4C75DFD6FF9E4A08FC96F856FB97F819" authority="Deichmann, 1941" authorityName="Deichmann" authorityYear="1941" box="[777,1125,1926,1952]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="parafusus">
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<emphasis id="B9017847FF9E4A08FC96F856FC66F819" box="[777,916,1926,1952]" italics="true" pageId="4" pageNumber="511">T. parafusus</emphasis>
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<bibRefCitation id="EFE4D9A4FF9E4A08FC02F857FB97F819" author="Deichmann, E." box="[925,1125,1926,1952]" pageId="4" pageNumber="511" pagination="61 - 195" refId="ref8529" refString="Deichmann, E. (1941) The Holothurioidea collected by the VELERO III during the years 1932 - 1938. Part I. Dendrochirotida. Allan Hancock Pacific Expeditions, 8 (3), 61 - 195., pls. 10 - 30." type="journal article" year="1941">Deichmann, 1941</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C75DFD6FF9E4A08FB00F856FF22F87D" authority="Sluiter, 1914" authorityName="Sluiter" authorityYear="1914" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="species" species="uniannulata">
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<emphasis id="B9017847FF9E4A08FB00F856FAB1F819" box="[1183,1347,1926,1952]" italics="true" pageId="4" pageNumber="511">T. uniannulata</emphasis>
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<bibRefCitation id="EFE4D9A4FF9E4A08FAD3F857FF22F87D" author="Sluiter, C. P." pageId="4" pageNumber="511" pagination="1 - 28" refId="ref10004" refString="Sluiter, C. P. (1914) Die von Dr. P. N. van Kampen warend seiner Fahrt mit Regierungsdampfer GIER 1906 - 1909, im Indischen Archipel gesammelten Holothurien. Buitensorg Contrib. Faune Indes Neerlandaiises 1, 1 - 28 and 1 plate." type="book chapter" year="1914">Sluiter, 1914</bibRefCitation>
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</taxonomicName>
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) are here transferred to the genus
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<emphasis id="B9017847FF9E4A08FDD1F87BFD6BF87D" box="[590,665,1962,1988]" italics="true" pageId="4" pageNumber="511">
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<taxonomicName id="4C75DFD6FF9E4A08FDD1F87BFD67F87D" authorityName="Selenka" authorityYear="1867" box="[590,661,1962,1988]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="511" phylum="Echinodermata" rank="genus">Stolus</taxonomicName>
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,
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</emphasis>
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whose diagnosis is herein amended to take in these forms.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |