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<document id="B201690016D3218FED7CAE5CB34845A6" ID-DOI="10.11646/zootaxa.5026.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300888" ID-ZooBank="A3AA106D-527A-4934-953C-C2EA746659FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039785980" checkinUser="plazi" docAuthor="Thandar, Ahmed S." docDate="2021" docId="03DC1543FF964A1CFF08F8F9FCB7FAB4" docLanguage="en" docName="zootaxa.5026.4.3.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyone Oken 1815" docType="treatment" docVersion="4" lastPageNumber="523" masterDocId="FFE56D3BFF9A4A0CFF9FFFD1FFF2FFB9" masterDocTitle="Nomenclatural changes in some sea cucumbers with the erection of a new genus and description of a Thyone? juvenile (? n. sp.) from the Gulf of California (Echinodermata: Holothuroidea: Dendrochirotida)" masterLastPageNumber="526" masterPageNumber="507" pageNumber="519" updateTime="1699139049944" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A57B93B46044FC040AFE245175ABCDD5">Nomenclatural changes in some sea cucumbers with the erection of a new genus and description of a Thyone? juvenile (? n. sp.) from the Gulf of California (Echinodermata: Holothuroidea: Dendrochirotida)</mods:title>
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<mods:namePart id="D0FB4DF8573D7F5232798E76F8D10F02">Thandar, Ahmed S.</mods:namePart>
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<mods:date id="A7271CAC90C778C6783AB2904EF76DA7">2021</mods:date>
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<treatment id="03DC1543FF964A1CFF08F8F9FCB7FAB4" ID-DOI="http://doi.org/10.5281/zenodo.5307843" ID-Zenodo-Dep="5307843" LSID="urn:lsid:plazi:treatment:03DC1543FF964A1CFF08F8F9FCB7FAB4" httpUri="http://treatment.plazi.org/id/03DC1543FF964A1CFF08F8F9FCB7FAB4" lastPageId="16" lastPageNumber="523" pageId="12" pageNumber="519">
<subSubSection id="C36FF7DEFF964A00FF08F8F9FEB9F8FA" box="[151,331,1832,1859]" pageId="12" pageNumber="519" type="nomenclature">
<paragraph id="8BCAA455FF964A00FF08F8F9FEB9F8FA" blockId="12.[151,331,1832,1895]" box="[151,331,1832,1859]" pageId="12" pageNumber="519">
<heading id="D0821339FF964A00FF08F8F9FEB9F8FA" bold="true" box="[151,331,1832,1859]" fontSize="11" level="1" pageId="12" pageNumber="519" reason="1">
<emphasis id="B9017847FF964A00FF08F8F9FEB9F8FA" bold="true" box="[151,331,1832,1859]" pageId="12" pageNumber="519">
<taxonomicName id="4C75DFD6FF964A00FF08F8F9FF00F8FA" ID-CoL="7Q4FF" authorityName="Oken" authorityYear="1815" box="[151,242,1832,1859]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="519" phylum="Echinodermata" rank="genus">
<emphasis id="B9017847FF964A00FF08F8F9FF00F8FA" bold="true" box="[151,242,1832,1859]" italics="true" pageId="12" pageNumber="519">Thyone</emphasis>
</taxonomicName>
?n. sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36FF7DEFF964A00FF08F89DFEF1F8DE" box="[151,259,1868,1895]" pageId="12" pageNumber="519" type="description">
<paragraph id="8BCAA455FF964A00FF08F89DFEF1F8DE" blockId="12.[151,331,1832,1895]" box="[151,259,1868,1895]" pageId="12" pageNumber="519">
<figureCitation id="134EB8D0FF964A00FF08F89DFEF1F8DE" box="[151,259,1868,1895]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="12" pageNumber="519">Figure 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36FF7DEFF964A00FF08F845FEB4F84E" pageId="12" pageNumber="519" type="materials_examined">
<paragraph id="8BCAA455FF964A00FF08F845FEB4F84E" blockId="12.[151,1436,1940,2039]" pageId="12" pageNumber="519">
<emphasis id="B9017847FF964A00FF08F845FE71F817" bold="true" box="[151,387,1940,1966]" pageId="12" pageNumber="519">Material examined:</emphasis>
<materialsCitation id="3B1DAE08FF964A00FE14F844FEB4F84E" accessionNumber="E51306" collectingDate="1967-10-21" collectionCode="USNM" collectorName="Col &amp; Burch, B. &amp; Burch T. &amp; Acc. &amp; C. Gust Ahearn" country="Mexico" county="North Pacific Ocean" elevation="20" latitude="31.316668" location="Choya" longLatPrecision="1216" longitude="-113.683334" municipality="Gulf of California" pageId="12" pageNumber="519" specimenCount="1" stateProvince="Sonora">
<collectionCode id="ED643C90FF964A00FE14F844FE2EF816" box="[395,476,1941,1967]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="519" type="Museum">USNM</collectionCode>
<accessionNumber id="942639B6FF964A00FE7CF845FDCFF816" box="[483,573,1940,1967]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E51306" pageId="12" pageNumber="519" type="EnaNcbi">E51306</accessionNumber>
,
<collectingCounty id="62ABDCD9FF964A00FDD7F845FCC2F816" box="[584,816,1940,1967]" pageId="12" pageNumber="519">North Pacific Ocean</collectingCounty>
,
<collectingMunicipality id="6BAE3E2FFF964A00FCA3F845FBFBF817" box="[828,1033,1940,1966]" pageId="12" pageNumber="519">Gulf of California</collectingMunicipality>
,
<collectingCountry id="F362E4C5FF964A00FB8AF845FB9FF817" box="[1045,1133,1940,1966]" name="Mexico" pageId="12" pageNumber="519">Mexico</collectingCountry>
,
<collectingRegion id="49B16AB7FF964A00FBE6F844FB3EF816" box="[1145,1228,1941,1967]" country="Mexico" name="Sonora" pageId="12" pageNumber="519">Sonora</collectingRegion>
;
<location id="8EAAF28EFF964A00FB47F845FAD7F817" LSID="urn:lsid:plazi:treatment:03DC1543FF964A1CFF08F8F9FCB7FAB4:8EAAF28EFF964A00FB47F845FAD7F817" box="[1240,1317,1940,1966]" country="Mexico" county="North Pacific Ocean" latitude="31.316668" longLatPrecision="1216" longitude="-113.683334" municipality="Gulf of California" name="Choya" pageId="12" pageNumber="519" stateProvince="Sonora">Choya</location>
; sta 67149 (
<geoCoordinate id="EE41C292FF964A00FF3FF868FEFDF86A" box="[160,271,1977,2003]" degrees="31" direction="north" minutes="19" orientation="latitude" pageId="12" pageNumber="519" precision="925" value="31.316668">31° 19 N</geoCoordinate>
,
<geoCoordinate id="EE41C292FF964A00FE85F868FE6FF86A" box="[282,413,1977,2003]" degrees="113" direction="west" minutes="41" orientation="longitude" pageId="12" pageNumber="519" precision="925" value="-113.683334">113° 41 W</geoCoordinate>
),
<quantity id="4C8D09B0FF964A00FE2FF869FE15F86B" box="[432,487,1976,2003]" metricMagnitude="1" metricUnit="m" metricValue="1.9812" pageId="12" pageNumber="519" unit="ft" value="65.0">
<elevation id="00584366FF964A00FE2FF869FE15F86B" box="[432,487,1976,2003]" metricMagnitude="1" metricUnit="m" metricValue="1.9812" pageId="12" pageNumber="519" unit="ft" value="65.0">65 ft</elevation>
</quantity>
,
<date id="FFCB8295FF964A00FE6DF868FD8DF86B" box="[498,639,1976,2003]" pageId="12" pageNumber="519" value="1967-10-21">
<collectingDate id="EF8F7B7DFF964A00FE6DF868FD8DF86B" box="[498,639,1976,2003]" pageId="12" pageNumber="519" value="1967-10-21">21 Oct 1967</collectingDate>
</date>
,
<collectorName id="2680C183FF964A00FD15F869FD45F86B" box="[650,695,1976,2002]" pageId="12" pageNumber="519">Col</collectorName>
:
<collectorName id="2680C183FF964A00FD5EF869FCDEF86A" box="[705,812,1976,2003]" pageId="12" pageNumber="519">Burch, B.</collectorName>
&amp;
<collectorName id="2680C183FF964A00FCCFF869FC41F86A" box="[848,947,1976,2003]" pageId="12" pageNumber="519">Burch T.</collectorName>
,
<collectorName id="2680C183FF964A00FC22F868FC03F86A" box="[957,1009,1977,2003]" pageId="12" pageNumber="519">Acc.</collectorName>
no: 300062, det.
<taxonomicName id="4C75DFD6FF964A00FB2BF869FADCF86A" box="[1204,1326,1976,2003]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="519" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B9017847FF964A00FB2BF869FAF4F86B" box="[1204,1286,1976,2002]" italics="true" pageId="12" pageNumber="519">Thyone</emphasis>
sp.
</taxonomicName>
?,
<collectorName id="2680C183FF964A00FAD8F868FF1EF84F" pageId="12" pageNumber="519">C. Gust Ahearn</collectorName>
,
<specimenCount id="9D736FDCFF964A00FF67F80CFEB4F84E" box="[248,326,2013,2039]" pageId="12" pageNumber="519" type="generic">1 spec.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF0AF4DDFF974A01FF08F917FB2BF89F" ID-DOI="http://doi.org/10.5281/zenodo.5300904" ID-Zenodo-Dep="5300904" httpUri="https://zenodo.org/record/5300904/files/figure.png" pageId="13" pageNumber="520" startId="13.[151,250,1734,1758]" targetBox="[268,1316,205,1702]" targetPageId="13">
<paragraph id="8BCAA455FF974A01FF08F917FB2BF89F" blockId="13.[151,1437,1734,1831]" pageId="13" pageNumber="520">
<emphasis id="B9017847FF974A01FF08F917FEE6F967" bold="true" box="[151,276,1734,1758]" pageId="13" pageNumber="520">FIGURE 8.</emphasis>
<taxonomicName id="4C75DFD6FF974A01FE83F917FDD4F966" authority="Sluiter, 1910" authorityName="Sluiter" authorityYear="1910" box="[284,550,1734,1759]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="520" phylum="Echinodermata" rank="species" species="vilis">
<emphasis id="B9017847FF974A01FE83F917FE6AF967" box="[284,408,1734,1758]" italics="true" pageId="13" pageNumber="520">Thyone vilis</emphasis>
Sluiter, 1910
</taxonomicName>
. Lectotype, ZMUA, H1114. A. Calcareous ring (middorsal view); B. Table from body wall; C. Rod from tube foot.; D. Table from introvert; E. Rosette from introvert; G. Plate of?Molo Strait specimen. (AG copied from Sluiter 1910); F. Plate from Molo Strait specimen (authors examination). (All ossicles to same scale).
</paragraph>
</caption>
<caption id="DF0AF4DDFF944A02FF08F971FEECF964" ID-DOI="http://doi.org/10.5281/zenodo.5300906" ID-Zenodo-Dep="5300906" httpUri="https://zenodo.org/record/5300906/files/figure.png" pageId="14" pageNumber="521" startId="14.[151,250,1696,1720]" targetBox="[207,1379,189,1665]" targetPageId="14">
<paragraph id="8BCAA455FF944A02FF08F971FEECF964" blockId="14.[151,1437,1696,1757]" pageId="14" pageNumber="521">
<emphasis id="B9017847FF944A02FF08F971FEE6F900" bold="true" box="[151,276,1696,1721]" pageId="14" pageNumber="521">FIGURE 9.</emphasis>
<taxonomicName id="4C75DFD6FF944A02FE82F970FDDAF900" authority="Sluiter, 1910" authorityName="Sluiter" authorityYear="1910" box="[285,552,1697,1721]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="521" phylum="Echinodermata" rank="species" species="vilis">
<emphasis id="B9017847FF944A02FE82F970FE6BF900" box="[285,409,1697,1721]" italics="true" pageId="14" pageNumber="521">Thyone vilis</emphasis>
Sluiter, 1910
</taxonomicName>
. USNM
<accessionNumber id="942639B6FF944A02FD1AF970FD26F900" box="[645,724,1697,1721]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E48738" pageId="14" pageNumber="521" type="EnaNcbi">E48738</accessionNumber>
A, B &amp; C. Regular, irregular and round tables of body wall; D.Tube feet deposits.
</paragraph>
</caption>
<caption id="DF0AF4DDFF954A03FF08F85DFD0EF871" ID-DOI="http://doi.org/10.5281/zenodo.5300908" ID-Zenodo-Dep="5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="15" pageNumber="522" startId="15.[151,250,1932,1956]" targetBox="[162,1427,190,1899]" targetPageId="15">
<paragraph id="8BCAA455FF954A03FF08F85DFD0EF871" blockId="15.[151,1436,1932,1992]" pageId="15" pageNumber="522">
<emphasis id="B9017847FF954A03FF08F85DFED0F81D" bold="true" box="[151,290,1932,1956]" pageId="15" pageNumber="522">FIGURE 10.</emphasis>
<taxonomicName id="4C75DFD6FF954A03FEB5F85DFE87F81D" authorityName="Oken" authorityYear="1815" box="[298,373,1932,1956]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="522" phylum="Echinodermata" rank="genus">
<emphasis id="B9017847FF954A03FEB5F85DFE87F81D" box="[298,373,1932,1956]" italics="true" pageId="15" pageNumber="522">Thyone</emphasis>
</taxonomicName>
<emphasis id="B9017847FF954A03FEE2F85DFE4EF81D" bold="true" box="[381,444,1932,1956]" pageId="15" pageNumber="522">?n.sp.</emphasis>
A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale).
</paragraph>
</caption>
<subSubSection id="C36FF7DEFF8A4A1CFF58FF47FC75FCAC" pageId="16" pageNumber="523" type="description">
<paragraph id="8BCAA455FF8A4A1CFF58FF47FDB2FE4C" blockId="16.[151,1437,150,1293]" pageId="16" pageNumber="523">
<emphasis id="B9017847FF8A4A1CFF58FF47FEABFF09" bold="true" box="[199,345,150,176]" pageId="16" pageNumber="523">Description.</emphasis>
Specimen small, perhaps juvenile, previously dissected, with introvert and tentacles expelled but still attached to body. Length
<quantity id="4C8D09B0FF8A4A1CFE7AFF6AFDC4FF6C" box="[485,566,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="523" unit="mm" value="10.0">10 mm</quantity>
, breadth in mid-body about
<quantity id="4C8D09B0FF8A4A1CFCEBFF6BFC45FF6C" box="[884,951,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="523" unit="mm" value="5.0">5 mm</quantity>
, preserved colour whitish. Body somewhat barrel-shaped with mouth and anus at opposite ends with the mid-ventral ambulacrum appearing as a sole lacking tube feet. Tube feet elsewhere well-developed, mostly confined to the remaining four ambulacra with a few also scattered in the inter-ambulacra, short, suckers more or less of the same diameter as tube feet. Anal teeth present. Calcareous ring (
<figureCitation id="134EB8D0FF8A4A1CFEC4FE9AFE13FEDC" box="[347,481,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10A</figureCitation>
) well calcified with deeply cleft radial plates and both radial and interradial plates anteriorly bifid with the latter slightly fused to radial plates. Tube of calcareous ring short, fragmentation of the ring and the processes not discernible perhaps because of the presumed juvenility of the specimen. Polian vesicle single, sac-like; stone canal short, straight, free; madreporite kidney-shaped. Other internal organs lost, perhaps due to evisceration or previous dissection.
</paragraph>
<paragraph id="8BCAA455FF8A4A1CFF58FE2EFC75FCAC" blockId="16.[151,1437,150,1293]" pageId="16" pageNumber="523">
Ossicles of body wall comprise tables (
<figureCitation id="134EB8D0FF8A4A1CFD17FE2EFCF9FDA0" box="[648,779,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10B</figureCitation>
) with disc of usually four holes, spire short or elongated, ending in two teeth, some with a spire diverging into two denticulate arms (note by first examiner Gust Ahearn). Tube feet tables (
<figureCitation id="134EB8D0FF8A4A1CFEC3FD96FE12FDD8" box="[348,480,583,609]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10C</figureCitation>
) typically of the
<taxonomicName id="4C75DFD6FF8A4A1CFD39FD96FD0AFDD8" authorityName="Oken" authorityYear="1815" box="[678,760,583,609]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="523" phylum="Echinodermata" rank="genus">
<emphasis id="B9017847FF8A4A1CFD39FD96FD0AFDD8" box="[678,760,583,609]" italics="true" pageId="16" pageNumber="523">Thyone</emphasis>
</taxonomicName>
<typeStatus id="54CE1AF7FF8A4A1CFC9EFD96FCC3FDD8" box="[769,817,583,609]" pageId="16" pageNumber="523">type</typeStatus>
with the disc straight or slightly curved and the spire, when present, reduced or elongated, terminating in two diverging teeth; disc with four central holes and one at each extremity. End plates present (
<figureCitation id="134EB8D0FF8A4A1CFE7AFD5EFD97FD10" box="[485,613,655,681]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10F</figureCitation>
), well-formed with few minute central holes and several concentric circles of larger holes around these. Introvert deposits (
<figureCitation id="134EB8D0FF8A4A1CFD32FD62FCC2FD74" box="[685,816,691,717]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10D</figureCitation>
) comprise well-developed tables with multilocular disc and short bidentate spire, spire sometimes absent, then tables resembling multilocular plates. Tentacle deposits as minute plates with several holes and a jagged margin (
<figureCitation id="134EB8D0FF8A4A1CFD68FD2AFC88FCAC" box="[759,890,763,789]" captionStart="FIGURE 10" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1427,190,1899]" captionTargetId="figure-16@15.[151,1436,181,1908]" captionTargetPageId="15" captionText="FIGURE 10. Thyone?n.sp. A. Calcareous ring; B. Body wall tables; C. Tube feet deposits; D. Introvert tables; E. Tentacle ossicles; F. Tube foot end plates.(All ossicles to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5300908" httpUri="https://zenodo.org/record/5300908/files/figure.png" pageId="16" pageNumber="523">Figure 10E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36FF7DEFF8A4A1CFF58FCCEFCB7FAB4" pageId="16" pageNumber="523" type="discussion">
<paragraph id="8BCAA455FF8A4A1CFF58FCCEFCB7FAB4" blockId="16.[151,1437,150,1293]" pageId="16" pageNumber="523">
<emphasis id="B9017847FF8A4A1CFF58FCCEFEC9FC80" bold="true" box="[199,315,799,825]" pageId="16" pageNumber="523">Remarks.</emphasis>
Because of the singularity and presumed juvenility of the specimen it was not possible to determine its exact identity. The calcareous ring is clearly like all East Pacific species of
<taxonomicName id="4C75DFD6FF8A4A1CFB98FC92FBABFCE4" authorityName="Oken" authorityYear="1815" box="[1031,1113,835,861]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="523" phylum="Echinodermata" rank="genus">
<emphasis id="B9017847FF8A4A1CFB98FC92FBABFCE4" box="[1031,1113,835,861]" italics="true" pageId="16" pageNumber="523">Thyone</emphasis>
</taxonomicName>
in being deeply cleft but the tentacle and introvert deposits indicate that the current specimen perhaps belong to a species not yet described. Since many holothuroids can lose or modify their deposits with age (see
<bibRefCitation id="EFE4D9A4FF8A4A1CFC24FC5AFBA8FC1C" author="Thandar, A. S." box="[955,1114,907,933]" pageId="16" pageNumber="523" pagination="278 - 286" refId="ref10089" refString="Thandar, A. S. (1987) The southern African stichopodid holothurians, with notes on the changes in spicule composition with age in the endemic Neostichopus grammatus (H. L. Clark). South African Journal of Zoology, 22, 278 - 286. https: // doi. org / 10.1080 / 02541858.1987.11448058" type="journal article" year="1987">Thandar 1987</bibRefCitation>
, 1991,
<bibRefCitation id="EFE4D9A4FF8A4A1CFB36FC5AFAC9FC1C" author="Massin, C. L." box="[1193,1339,907,933]" pageId="16" pageNumber="523" pagination="179 - 191" refId="ref9379" refString="Massin, C. L. (1992) Three new species of Dendrochirotida (Holothuroidea: Echinodermata) from the Weddell Sea (Antarctica). Bulletin de L'Institut Royal Des Sciences Naturelles de Belgique, Biologie, 62, 179 - 191." type="journal article" year="1992">Massin 1992</bibRefCitation>
, Martins 2019), describing the single, perhaps juvenile individual as a new species is not here exercised. Although the body wall table discs may match those of
<taxonomicName id="4C75DFD6FF8A4A1CFDA4FC02FC21FC54" authority="Deichmann, 1941" authorityName="Deichmann" authorityYear="1941" box="[571,979,979,1005]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="523" phylum="Echinodermata" rank="species" species="bidentata">
<emphasis id="B9017847FF8A4A1CFDA4FC02FCF2FC54" box="[571,768,979,1005]" italics="true" pageId="16" pageNumber="523">Thyone bidentata</emphasis>
<bibRefCitation id="EFE4D9A4FF8A4A1CFC97FC02FC21FC54" author="Deichmann, E." box="[776,979,979,1005]" pageId="16" pageNumber="523" pagination="61 - 195" refId="ref8529" refString="Deichmann, E. (1941) The Holothurioidea collected by the VELERO III during the years 1932 - 1938. Part I. Dendrochirotida. Allan Hancock Pacific Expeditions, 8 (3), 61 - 195., pls. 10 - 30." type="journal article" year="1941">Deichmann, 1941</bibRefCitation>
</taxonomicName>
, known from
<collectingRegion id="49B16AB7FF8A4A1CFBEAFC02FB15FC54" box="[1141,1255,979,1005]" country="United States of America" name="California" pageId="16" pageNumber="523">California</collectingRegion>
to Columbia, it differs in the spire of the tables and the absence of rosettes in the tentacles, unless by some stretch of imagination the minute plates were considered to be rosettes by
<bibRefCitation id="EFE4D9A4FF8A4A1CFD4BFBCAFC57FB8C" author="Deichmann, E." box="[724,933,1051,1077]" pageId="16" pageNumber="523" pagination="61 - 195" refId="ref8529" refString="Deichmann, E. (1941) The Holothurioidea collected by the VELERO III during the years 1932 - 1938. Part I. Dendrochirotida. Allan Hancock Pacific Expeditions, 8 (3), 61 - 195., pls. 10 - 30." type="journal article" year="1941">Deichmann (1941)</bibRefCitation>
but regrettably they were not illustrated. The tube feet table discs are also less curved in the current material and their spire short and not bidentate. The specimen also comes quite close to
<taxonomicName id="4C75DFD6FF8A4A1CFE5EFBB2FD1BFBC4" authority="Deichmann, 1937" authorityName="Deichmann" authorityYear="1937" box="[449,745,1123,1149]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="523" phylum="Echinodermata" rank="species" species="benti">
<emphasis id="B9017847FF8A4A1CFE5EFBB2FDE5FBC4" box="[449,535,1123,1149]" italics="true" pageId="16" pageNumber="523">T. benti</emphasis>
<bibRefCitation id="EFE4D9A4FF8A4A1CFDBFFBB2FD1BFBC4" author="Deichmann, E." box="[544,745,1123,1149]" pageId="16" pageNumber="523" pagination="161 - 176" refId="ref8406" refString="Deichmann, E. (1937) The Templeton Crocker Expedition. IX. Holothurians from the Gulf of California, the West Coast of Lower California, and Clarion Island. Zoologica (New York), 22, 161 - 176." type="journal article" year="1937">Deichmann, 1937</bibRefCitation>
</taxonomicName>
from the Californian coast but differs in the absence of any bifurcations to the anterior tips of the plates of the calcareous ring, the presence of a taller spire to the body wall tables and the absence of rods in the tentacles. One anomaly in the current specimen is the occurrence of the solelike ventral ambulacrum. Whether this is a consistent feature or an abnormality will have to be determined from new material. It does not seem likely that it is due to preservation.
</paragraph>
</subSubSection>
</treatment>
</document>