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<document id="14990EABC9DF8EC308A773B91AD8B6E9" ID-DOI="10.5852/ejt.2024.921.2429" ID-ISSN="2118-9773" ID-Zenodo-Dep="10663874" ID-ZooBank="02633F29-4CDF-4027-BEBF-07AD2F925B42" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1707990651991" checkinUser="plazi" docAuthor="Bellvert, Adrià, Dimitrov, Dragomir, Zamani, Alireza &amp; Arnedo, Miquel A." docDate="2024" docId="03DA7943B73ED85972738EB1451CFE97" docLanguage="en" docName="EJT.2024.921.210-235.pdf" docOrigin="European Journal of Taxonomy 921 (210)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2429/10759" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dysdera caspica Dunin 1990" docType="treatment" docVersion="3" lastPageNumber="218" masterDocId="FFE3013BB738D85170598D5B445AFFC5" masterDocTitle="Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia" masterLastPageNumber="235" masterPageNumber="210" pageNumber="216" updateTime="1708090743251" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="801CF2F254848DA9E37BFE59A39E0D29">Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia</mods:title>
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<mods:affiliation id="09652729CDEFF4FE300D4815EC874507">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
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<mods:namePart id="E81861E1DB869250D75B9E1F239C4968">Zamani, Alireza</mods:namePart>
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<mods:namePart id="04EA0C5E990DA5EF51A4B2996CEAE82A">Arnedo, Miquel A.</mods:namePart>
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<paragraph id="8BCCC855B73ED85772738EB14050FBC0" blockId="6.[554,1034,1002,1065]" box="[554,1034,1002,1029]" pageId="6" pageNumber="216">
<heading id="D0847F39B73ED85772738EB14050FBC0" box="[554,1034,1002,1029]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="216" reason="2">
<taxonomicName id="4C73B3D6B73ED85772738EB147D2FBC0" authority="Dunin, 1990" authorityName="Dunin" authorityYear="1990" box="[554,904,1002,1029]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="216" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B9071447B73ED85772738EB146B4FBC1" bold="true" box="[554,750,1002,1029]" italics="true" pageId="6" pageNumber="216">Dysdera caspica</emphasis>
<bibRefCitation id="EFE2B5A4B73ED85772AC8EB047D2FBC0" author="Dunin" box="[757,904,1003,1029]" firstAuthor="Dunin" pageId="6" pageNumber="216" pagination="141 - 148" refId="ref11789" refString="Dunin P. M. 1990. Spiders of the genus Dysdera (Aranei, Haplogynae, Dysderidae) from Azerbaijan. Z oologicheskii Zhurnal 69 (6): 141 - 148." type="journal article" year="1990">Dunin, 1990</bibRefCitation>
</taxonomicName>
comb. rev.
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</paragraph>
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<figureCitation id="1348D4D0B73ED85772BE89554717FBEC" box="[743,845,1038,1065]" captionStart="Figs 35" captionStartId="7.[189,241,1963,1989]" captionTargetBox="[189,1398,1092,1925]" captionTargetId="figure-299@7.[189,1398,1092,1925]" captionTargetPageId="7" captionText="Figs 35. Male Dysdera caspica Dunin, 1990 comb. rev. (crba000530 CRBA), left palp (F broken on 3 and 4). 3. Prolateral view. 4. Anterior view. 5. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663882" httpUri="https://zenodo.org/record/10663882/files/figure.png" pageId="6" pageNumber="216">Figs 35</figureCitation>
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<paragraph id="8BCCC855B73ED85770E4890B4687FBAF" blockId="6.[189,733,1104,1130]" box="[189,733,1104,1130]" pageId="6" pageNumber="216">
<treatmentCitationGroup id="AB63EF7BB73ED85770E4890B4687FBAF" box="[189,733,1104,1130]" pageId="6" pageNumber="216">
<treatmentCitation id="0AD2EE44B73ED85770E4890B4603FBAF" author="Dunin P. M." box="[189,601,1104,1130]" page="143" pageId="6" pageNumber="216" year="1990">
<taxonomicName id="4C73B3D6B73ED85770E4890B4603FBAF" authority="Dunin, 1990: 143" authorityName="Dunin" authorityPageNumber="143" authorityYear="1990" box="[189,601,1104,1130]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="216" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B9071447B73ED85770E4890B45D8FBAF" box="[189,386,1104,1130]" italics="true" pageId="6" pageNumber="216">Dysdera caspica</emphasis>
<bibRefCitation id="EFE2B5A4B73ED85771D0890B4603FBAF" author="Dunin" box="[393,601,1104,1130]" firstAuthor="Dunin" page="143" pageId="6" pageNumber="216" pagination="141 - 148" refId="ref11789" refString="Dunin P. M. 1990. Spiders of the genus Dysdera (Aranei, Haplogynae, Dysderidae) from Azerbaijan. Z oologicheskii Zhurnal 69 (6): 141 - 148." type="journal article" year="1990">Dunin, 1990: 143</bibRefCitation>
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, fig. 4.14.
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</paragraph>
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<treatmentCitationGroup id="AB63EF7BB73ED85770E489CA4696FB69" box="[189,716,1169,1196]" pageId="6" pageNumber="216">
<taxonomicName id="4C73B3D6B73ED85770E489CA45C4FB6E" ID-CoL="3899C" baseAuthorityName="Dunin" baseAuthorityYear="1990" box="[189,414,1169,1195]" class="Arachnida" family="Dysderidae" genus="Dysderella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="216" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B9071447B73ED85770E489CA45C4FB6E" box="[189,414,1169,1195]" italics="true" pageId="6" pageNumber="216">Dysderella caspica</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2EE44B73ED85771E289CA462CFB6E" author="Dunin P. M." box="[443,630,1169,1196]" page="67" pageId="6" pageNumber="216" year="1992">
<bibRefCitation id="EFE2B5A4B73ED85771E289CA462CFB6E" author="Dunin" box="[443,630,1169,1196]" firstAuthor="Dunin" page="67" pageId="6" pageNumber="216" pagination="35 - 76" refId="ref11823" refString="Dunin P. M. 1992. The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta 1 (3): 35 - 76." type="journal article" year="1992">Dunin 1992: 67</bibRefCitation>
</treatmentCitation>
, fig. 12
</treatmentCitationGroup>
(transfer from
<taxonomicName id="4C73B3D6B73ED857732689CA47B8FB6E" ID-CoL="494G" baseAuthorityName="sensu Deeleman-Reinhold &amp; Deeleman" baseAuthorityYear="1988" box="[895,994,1169,1195]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B9071447B73ED857732689CA47B8FB6E" box="[895,994,1169,1195]" italics="true" pageId="6" pageNumber="216">Dysdera</emphasis>
</taxonomicName>
).
</paragraph>
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<heading id="D0847F39B73ED85770E48988456CFB28" bold="true" box="[189,310,1235,1261]" fontSize="11" level="3" pageId="6" pageNumber="216" reason="3">
<emphasis id="B9071447B73ED85770E48988456CFB28" bold="true" box="[189,310,1235,1261]" pageId="6" pageNumber="216">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B73ED85770E48859458AFAD9" blockId="6.[189,464,1282,1308]" box="[189,464,1282,1308]" pageId="6" pageNumber="216">
See
<bibRefCitation id="EFE2B5A4B73ED85770B688594596FAD9" author="Dunin" box="[239,460,1282,1308]" firstAuthor="Dunin" page="143" pageId="6" pageNumber="216" pagination="141 - 148" refId="ref11789" refString="Dunin P. M. 1990. Spiders of the genus Dysdera (Aranei, Haplogynae, Dysderidae) from Azerbaijan. Z oologicheskii Zhurnal 69 (6): 141 - 148." type="journal article" year="1990">Dunin (1990: 143)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB73ED85770E488184689FA74" pageId="6" pageNumber="216" type="materials_examined">
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<emphasis id="B9071447B73ED85770E4881845F0FA98" bold="true" box="[189,426,1347,1373]" pageId="6" pageNumber="216">Material examined</emphasis>
</heading>
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<paragraph id="8BCCC855B73ED85770E488284689FA74" blockId="6.[189,1399,1394,1457]" pageId="6" pageNumber="216">
<materialsCitation id="3B1BC208B73ED85770E488284695FA74" collectingDate="2003-06-18" collectionCode="CRBA" collectorName="Y. M. Marusik" country="Azerbaijan" latitude="40.490555" location="Peninsula" longLatPrecision="1" longitude="50.15167" pageId="6" pageNumber="216" specimenCount="1" specimenCount-male="1" stateProvince="Baku">
<collectingCountry id="F36488C5B73ED85770E488284535FA48" box="[189,367,1395,1421]" name="Azerbaijan" pageId="6" pageNumber="216">AZERBAIJAN</collectingCountry>
<specimenCount id="9D7503DCB73ED85771DC882845EBFA48" box="[389,433,1395,1421]" count="1" pageId="6" pageNumber="216" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49B706B7B73ED85771E588294609FA49" box="[444,595,1394,1420]" country="Azerbaijan" name="Baku" pageId="6" pageNumber="216">Baku Region</collectingRegion>
Absheron
<location id="8EAC9E8EB73ED857729588294765FA49" LSID="urn:lsid:plazi:treatment:03DA7943B73ED85972738EB1451CFE97:8EAC9E8EB73ED857729588294765FA49" box="[716,831,1394,1420]" country="Azerbaijan" latitude="40.490555" longLatPrecision="1" longitude="50.15167" name="Peninsula" pageId="6" pageNumber="216" stateProvince="Baku">Peninsula</location>
, near Baku;
<geoCoordinate id="EE47AE92B73ED857739288294034FA48" box="[971,1134,1394,1421]" degrees="40.490556" direction="north" orientation="latitude" pageId="6" pageNumber="216" precision="1" value="40.490555">40.490556° N</geoCoordinate>
,
<geoCoordinate id="EE47AE92B73ED857742088294140FA48" box="[1145,1306,1394,1421]" degrees="50.151667" direction="east" orientation="longitude" pageId="6" pageNumber="216" precision="1" value="50.15167">50.151667° E</geoCoordinate>
;
<date id="FFCDEE95B73ED857757D882844A1FA75" pageId="6" pageNumber="216" value="2003-06-18">
<collectingDate id="EF89177DB73ED857757D882844A1FA75" pageId="6" pageNumber="216" value="2003-06-18">18 Jun. 2003</collectingDate>
</date>
;
<collectorName id="2686AD83B73ED857715F88CC45F3FA75" box="[262,425,1430,1457]" pageId="6" pageNumber="216">Y.M. Marusik</collectorName>
leg.;
<collectionCode id="ED625090B73ED85771B288CD4663FA75" box="[491,569,1430,1456]" pageId="6" pageNumber="216">CRBA</collectionCode>
crba000530
</materialsCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="B9071447B73ED85770E4888C4514FA34" bold="true" box="[189,334,1495,1521]" pageId="6" pageNumber="216">Description</emphasis>
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</paragraph>
<paragraph id="8BCCC855B73ED85770B38B5C45F1F9E5" blockId="6.[187,1400,1542,1853]" box="[234,427,1542,1569]" pageId="6" pageNumber="216">
<emphasis id="B9071447B73ED85770B38B5C4570F9E4" bold="true" box="[234,298,1543,1569]" pageId="6" pageNumber="216">Male</emphasis>
(
<figureCitation id="1348D4D0B73ED85771608B5D45FBF9E4" box="[313,417,1542,1569]" captionStart="Figs 35" captionStartId="7.[189,241,1963,1989]" captionTargetBox="[189,1398,1092,1925]" captionTargetId="figure-299@7.[189,1398,1092,1925]" captionTargetPageId="7" captionText="Figs 35. Male Dysdera caspica Dunin, 1990 comb. rev. (crba000530 CRBA), left palp (F broken on 3 and 4). 3. Prolateral view. 4. Anterior view. 5. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663882" httpUri="https://zenodo.org/record/10663882/files/figure.png" pageId="6" pageNumber="216">Figs 35</figureCitation>
<emphasis id="B9071447B73ED85771F88B5D45F1F9E5" bold="true" box="[417,427,1542,1568]" pageId="6" pageNumber="216">)</emphasis>
</paragraph>
<paragraph id="8BCCC855B73ED85770E48B71405CF8F8" blockId="6.[187,1400,1542,1853]" pageId="6" pageNumber="216">PROSOMA. 1.88 long; maximum width 1.47; minimum width 0.95. Uniformly red; slightly wrinkled, anterior region smooth. Frontal border roughly round, ca ½ of carapace length; anterior lateral borders convergent; rounded at maximum dorsal width, posterior lateral borders rounded; posterior margin narrow, rounded. Eye diameters: AME 0.12; PLE 0.11; PME 0.10; AMEs on edge of frontal border, separated from one another by ca 1 diameter, touching PLEs; PMEs ca one-quarter of diameter apart, less than ¼ of PME diameter from PLEs. Labium trapezoid-shaped, its base wider than its distal part, borders slightly curved; longer than wide at base; semi-circular notch at tip. Sternum orange, frontally darker, becoming lighter posteriorly; smooth; uniformly covered with slender black setae.</paragraph>
<paragraph id="8BCCC855B73ED85770E48A3F4710F82C" blockId="6.[188,1399,1890,2025]" pageId="6" pageNumber="216">CHELICERAE. 0.84 long, ca ½ of carapace length in dorsal view; fang 0.80 long; paturon proximal dorsal and ventral side scantly covered with piligerous granulations. Cheliceral inner groove short, ca ⅓ of cheliceral length; armed with three teeth and lamina at base; B= M&gt; D; D triangular, located roughly at centre of groove; B close to basal lamina; M close to B.</paragraph>
<paragraph id="8BCCC855B73FD85670E48C5245A9FDD3" blockId="7.[187,1400,265,534]" pageId="7" pageNumber="217">LEGS. Orange. Lengths of leg segments (leg 2 missing): fe1 1.45; pa1 0.91; ti1 1.11; me1 1.14; ta1 0.42; total 5.05; fe3 0.98; pa3 0.58; ti3 0.60; me3 0.85; ta3 0.29; total 3.30; fe4 1.33; pa4 0.78; ti4 1.01; me4 1.21; ta4 0.38; total 4.73; fe Pdp 0.88; pa Pdp 0.54; ti Pdp 0.42; ta Pdp 0.53; total 2.37; leg formula: 1&gt;4&gt;3. Fe3d spineless; pa3 spineless; tb3d spines arranged in one band; proximal 0; medial-proximal 0; medial-distal 0; distal 1.0.0; tb3v spineless; with one terminal spine on forward margin. Fe4d spineless; pa4 spineless; tb4d spineless; tb4v spineless; with one terminal spine at anterior border. Dorsal side of frontal legs covered with small thin grains; ventral side of pedipalp covered with setae, lacking grains. Claws with 8 teeth or less.</paragraph>
<paragraph id="8BCCC855B73FD85670E48F6646EAFDBC" blockId="7.[187,1398,572,633]" pageId="7" pageNumber="217">OPISTHOSOMA. Cream-coloured. Abdominal dorsal hairs 0.03 long; thin, slightly curved, not compressed, pointed; uniformly and thickly distributed.</paragraph>
<paragraph id="8BCCC855B73FD85670E48FFB40CCFC68" blockId="7.[188,1401,672,941]" pageId="7" pageNumber="217">
PALP (
<figureCitation id="1348D4D0B73FD856715A8FFB4528FD7E" box="[259,370,672,699]" captionStart="Figs 35" captionStartId="7.[189,241,1963,1989]" captionTargetBox="[189,1398,1092,1925]" captionTargetId="figure-299@7.[189,1398,1092,1925]" captionTargetPageId="7" captionText="Figs 35. Male Dysdera caspica Dunin, 1990 comb. rev. (crba000530 CRBA), left palp (F broken on 3 and 4). 3. Prolateral view. 4. Anterior view. 5. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663882" httpUri="https://zenodo.org/record/10663882/files/figure.png" pageId="7" pageNumber="217">FIGS 35</figureCitation>
). T markedly shorter than ED; external distal border straight; internal sloped backward. ED slightly bent in lateral view; internal distal border not expanded. IS and ES equally developed, fused basally; IS and ES continuous to tip, bent ventrally. ED tip straight in lateral view. C present, long; distal border rounded, smooth, slightly expanded, projected over ED external part. AC absent. LF absent. L poorly developed; external border sclerotized, not folded. LA absent. F present, distally curved. AL present, very poorly developed; proximal border in posterior view fused with DH. Base of P fused to T; slightly sloped upwards; lateral length from ⅓ to ⅖ of T width; ridge present, perpendicular to T; not expanded, upper margin smooth; not distally projected; posterior margin not folded.
</paragraph>
<paragraph id="8BCCC855B73FD85670B38E8F451FFC2B" blockId="7.[189,573,980,1041]" box="[234,325,980,1006]" pageId="7" pageNumber="217">
<emphasis id="B9071447B73FD85670B38E8F451FFC2B" bold="true" box="[234,325,980,1006]" pageId="7" pageNumber="217">Female</emphasis>
</paragraph>
<paragraph id="8BCCC855B73FD85670E48EAC4667FBD4" blockId="7.[189,573,980,1041]" box="[189,573,1015,1041]" pageId="7" pageNumber="217">
Not examined. See
<bibRefCitation id="EFE2B5A4B73FD85671FA8EAC4663FBD4" author="Dunin" box="[419,569,1015,1041]" firstAuthor="Dunin" pageId="7" pageNumber="217" pagination="141 - 148" refId="ref11789" refString="Dunin P. M. 1990. Spiders of the genus Dysdera (Aranei, Haplogynae, Dysderidae) from Azerbaijan. Z oologicheskii Zhurnal 69 (6): 141 - 148." type="journal article" year="1990">Dunin, 1990</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0C98DDB73FD85670E48AF040FAF82C" ID-DOI="http://doi.org/10.5281/zenodo.10663882" ID-Zenodo-Dep="10663882" httpUri="https://zenodo.org/record/10663882/files/figure.png" pageId="7" pageNumber="217" startId="7.[189,241,1963,1989]" targetBox="[189,1398,1092,1925]" targetPageId="7">
<paragraph id="8BCCC855B73FD85670E48AF040FAF82C" blockId="7.[189,1399,1963,2025]" pageId="7" pageNumber="217">
<emphasis id="B9071447B73FD85670E48AF04576F800" bold="true" box="[189,300,1963,1989]" pageId="7" pageNumber="217">Figs 35.</emphasis>
Male
<taxonomicName id="4C73B3D6B73FD856712F8AF0468FF800" authority="Dunin, 1990" authorityName="Dunin" authorityYear="1990" box="[374,725,1963,1989]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="217" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B9071447B73FD856712F8AF04661F800" box="[374,571,1963,1989]" italics="true" pageId="7" pageNumber="217">Dysdera caspica</emphasis>
<bibRefCitation id="EFE2B5A4B73FD856721B8AF0468FF800" author="Dunin" box="[578,725,1963,1989]" firstAuthor="Dunin" pageId="7" pageNumber="217" pagination="141 - 148" refId="ref11789" refString="Dunin P. M. 1990. Spiders of the genus Dysdera (Aranei, Haplogynae, Dysderidae) from Azerbaijan. Z oologicheskii Zhurnal 69 (6): 141 - 148." type="journal article" year="1990">Dunin, 1990</bibRefCitation>
</taxonomicName>
comb. rev. (crba000530 CRBA), left palp (F broken on 3 and 4).
<emphasis id="B9071447B73FD856714F8A95457FF82D" bold="true" box="[278,293,1998,2024]" pageId="7" pageNumber="217">3</emphasis>
. Prolateral view.
<emphasis id="B9071447B73FD85671AB8A95465BF82D" bold="true" box="[498,513,1998,2024]" pageId="7" pageNumber="217">4</emphasis>
. Anterior view.
<emphasis id="B9071447B73FD85672E48A954696F82D" bold="true" box="[701,716,1998,2024]" pageId="7" pageNumber="217">5</emphasis>
. Retrolateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3699BDEB730D85970E48C53451CFE97" box="[189,342,264,338]" pageId="8" pageNumber="218" type="distribution">
<paragraph id="8BCCC855B730D85970E48C53450CFEE7" blockId="8.[189,342,264,290]" box="[189,342,264,290]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85970E48C53450CFEE7" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="8" pageNumber="218" reason="3">
<emphasis id="B9071447B730D85970E48C53450CFEE7" bold="true" box="[189,342,264,290]" pageId="8" pageNumber="218">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B730D85970E48C63451CFE97" blockId="8.[189,326,312,338]" box="[189,326,312,338]" pageId="8" pageNumber="218">
<collectingCountry id="F36488C5B730D85970E48C634518FE97" box="[189,322,312,338]" name="Azerbaijan" pageId="8" pageNumber="218">Azerbaijan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>