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<document id="2431AF383E801DF0914CC6359C5D1D78" ID-DOI="10.5281/zenodo.193258" ID-GBIF-Dataset="85176744-66ec-48ba-860c-671c02718c50" ID-ISSN="1175-5326" ID-Zenodo-Dep="193258" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460129092828" checkinUser="plazi" docAuthor="Johanson, Kjell Arne &amp; Oláh, János" docDate="2010" docId="03CCF00CFF8F1E61FF30F604E8565756" docLanguage="en" docName="zt02338p062.pdf" docOrigin="Zootaxa 2338" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paduniella nandra Johanson &amp; Oláh, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FFF58874FF8D1E62FFA7F31DEC1A551A" masterDocTitle="Descriptions of five new species of Paduniella from Madagascar (Trichoptera: Psychomyiidae)" masterLastPageNumber="62" masterPageNumber="54" pageNumber="56" updateTime="1698235096248" updateUser="plazi">
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<mods:titleInfo id="9760FE703CD523FE215D961B5AFD31DF">
<mods:title id="129EDEC896D9E41B26918C6EE2AD3AC8">Descriptions of five new species of Paduniella from Madagascar (Trichoptera: Psychomyiidae)</mods:title>
</mods:titleInfo>
<mods:name id="F7663F0FA0ABDD415F1BC2427219349D" type="personal">
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<mods:namePart id="27F05FF567C3FE6385BEAB7D1AB7451C">Johanson, Kjell Arne</mods:namePart>
</mods:name>
<mods:name id="4B00836239585CB99B4CE528B434C60F" type="personal">
<mods:role id="5BF6F3E8E8847FEE9A5845E676157EB7">
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<mods:namePart id="FF2C3B9A80C0400D09F25FCD8AD3A539">Oláh, János</mods:namePart>
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<mods:typeOfResource id="EE6F0B485ABAC7D9215822D256BF42C9">text</mods:typeOfResource>
<mods:relatedItem id="2E15D263C2482396B56F14D6EB59E560" type="host">
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<mods:title id="0BB04A954678071E4D0F7ABC0CF84A1E">Zootaxa</mods:title>
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<mods:part id="C13D055A80E4309B5AEE089F563211EA">
<mods:date id="0CA7326A43BA54E9AFA276F5F2262869">2010</mods:date>
<mods:detail id="96C51039E7CD939945BC4D33B4323C28" type="volume">
<mods:number id="A74E7F3DC5FB34BEA7C29AEB33015F31">2338</mods:number>
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<mods:classification id="952A782CE4309960C7FFDA2DC2B6188B">journal article</mods:classification>
<mods:identifier id="734D1549BC575C73E3EBD9881A52749A" type="DOI">10.5281/zenodo.193258</mods:identifier>
<mods:identifier id="94FE23634D070E2B308780F88527F865" type="GBIF-Dataset">85176744-66ec-48ba-860c-671c02718c50</mods:identifier>
<mods:identifier id="D82BF9F508010890FDCAADA820583843" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F992E4198A3C19E10C795C7BFED3C834" type="Zenodo-Dep">193258</mods:identifier>
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<treatment id="03CCF00CFF8F1E61FF30F604E8565756" ID-DOI="http://doi.org/10.5281/zenodo.6205185" ID-GBIF-Taxon="124955619" ID-Zenodo-Dep="6205185" LSID="urn:lsid:plazi:treatment:03CCF00CFF8F1E61FF30F604E8565756" httpUri="http://treatment.plazi.org/id/03CCF00CFF8F1E61FF30F604E8565756" lastPageId="3" lastPageNumber="57" pageId="2" pageNumber="56">
<subSubSection id="C37F1291FF8F1E60FF30F604ECE7504C" pageId="2" pageNumber="56" type="nomenclature">
<paragraph id="8BDA411AFF8F1E60FF30F604EE025029" blockId="2.[151,536,1305,1366]" box="[151,536,1305,1331]" pageId="2" pageNumber="56">
<heading id="D092F676FF8F1E60FF30F604EE025029" bold="true" box="[151,536,1305,1331]" fontSize="11" level="1" pageId="2" pageNumber="56" reason="1">
<emphasis id="B9119D08FF8F1E60FF30F604EE025029" bold="true" box="[151,536,1305,1331]" pageId="2" pageNumber="56">
<taxonomicName id="4C653A99FF8F1E60FF30F604ED605029" box="[151,378,1305,1331]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="nandra" status="sp. nov.">
<emphasis id="B9119D08FF8F1E60FF30F604ED605029" bold="true" box="[151,378,1305,1331]" italics="true" pageId="2" pageNumber="56">Paduniella nandra</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2222073FF8F1E60FE2EF604EE025029" box="[393,536,1305,1331]" pageId="2" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA411AFF8F1E60FF30F621ECE7504C" blockId="2.[151,536,1305,1366]" box="[151,253,1340,1366]" pageId="2" pageNumber="56">
<figureCitation id="135E5D9FFF8F1E60FF30F621ECE7504C" box="[151,253,1340,1366]" captionStart="FIGURES 4 6" captionStartId="3.[151,272,1426,1450]" captionTargetBox="[153,1434,629,1375]" captionTargetId="figure@3.[151,1436,617,1405]" captionTargetPageId="3" captionText="FIGURES 4 6. Paduniella nandra, new species, holotype, male genitalia: 4, lateral; 5, phallic apparatus, lateral; 6, ventral." httpUri="https://zenodo.org/record/193260/files/figure.png" pageId="2" pageNumber="56">Figs 46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8F1E61FF30F69AE90054AD" lastPageId="3" lastPageNumber="57" pageId="2" pageNumber="56" type="description">
<paragraph id="8BDA411AFF8F1E60FF30F69AEF6450F1" blockId="2.[151,1437,1415,2038]" pageId="2" pageNumber="56">
The genitalia of
<taxonomicName id="4C653A99FF8F1E60FEFBF69AEDD550BA" box="[348,463,1415,1440]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="nandra">
<emphasis id="B9119D08FF8F1E60FEFBF69AEDD550BA" box="[348,463,1415,1440]" italics="true" pageId="2" pageNumber="56">P. nandra</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C653A99FF8F1E60FD16F69AEF0F50BA" box="[689,789,1415,1440]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ankya">
<emphasis id="B9119D08FF8F1E60FD16F69AEF0F50BA" box="[689,789,1415,1440]" italics="true" pageId="2" pageNumber="56">P. ankya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C653A99FF8F1E60FCF7F69AEFE250BA" box="[848,1016,1415,1440]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="filamentosa">
<emphasis id="B9119D08FF8F1E60FCF7F69AEFE250BA" box="[848,1016,1415,1440]" italics="true" pageId="2" pageNumber="56">P. filamentosa</emphasis>
</taxonomicName>
in lateral view in the presence of a broad ventral branch of the inferior appendages. It is distinguished from the two species by possessing a tapering main branch of the inferior appendages in lateral view.
</paragraph>
<paragraph id="8BDA411AFF8F1E61FF61F6EAE90054AD" blockId="2.[151,1437,1415,2038]" lastBlockId="3.[151,1437,152,588]" lastPageId="3" lastPageNumber="57" pageId="2" pageNumber="56">
<emphasis id="B9119D08FF8F1E60FF61F6EAEDE0530A" box="[198,506,1527,1552]" italics="true" pageId="2" pageNumber="56">
Male genitalia (
<figureCitation id="135E5D9FFF8F1E60FE26F6EAEDF7530A" box="[385,493,1527,1552]" captionStart="FIGURES 4 6" captionStartId="3.[151,272,1426,1450]" captionTargetBox="[153,1434,629,1375]" captionTargetId="figure@3.[151,1436,617,1405]" captionTargetPageId="3" captionText="FIGURES 4 6. Paduniella nandra, new species, holotype, male genitalia: 4, lateral; 5, phallic apparatus, lateral; 6, ventral." httpUri="https://zenodo.org/record/193260/files/figure.png" pageId="2" pageNumber="56">Figs 46</figureCitation>
).
</emphasis>
Sternite IX narrowly club-shaped in lateral view; both ventral arms with almost straight anterior and posterior margin, 3 times longer than wide, nearly
<quantity id="4C9DECFFFF8F1E60FBA2F501E878532C" box="[1029,1122,1564,1590]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="56" unit="mm" value="0.2">0.2 mm</quantity>
long; without swelling at posterior apex; both dorsal arms narrow, needle-shaped, slightly S-shaped, oriented anterad before curving dorsad, joined with tergite IX at mid-dorsal part of genitalia, about
<quantity id="4C9DECFFFF8F1E60FC61F57AE82A539B" box="[966,1072,1639,1665]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="56" unit="mm" value="0.35">0.35 mm</quantity>
long. Ventral and dorsal arms forming right angle. Tergite IX long, wide, smooth; slightly narrowing distad in lateral view; dorsal and ventral margins nearly straight; nearly
<quantity id="4C9DECFFFF8F1E60FDD2F5AFEEFB53D6" box="[629,737,1714,1740]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="56" unit="mm" value="0.45">0.45 mm</quantity>
long. Superior appendages originating ventroapically on tergite IX; margins parallel, about half as wide as widest part of sternite IX; straight; setae apparently restricted to distal half; apex rounded. Median branch of inferior appendages invisible in lateral view; originating about at mid-length of main branch in ventral view; widening from base into bilobed apex; mesal apical lobe narrow, lateral apical lobe wide; covered by setae, especially along mesal margin; about
<quantity id="4C9DECFFFF8F1E60FAE6F45AE986527B" box="[1345,1436,1863,1889]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="56" unit="mm" value="0.1">0.1 mm</quantity>
long. Ventral lobe of inferior appendages forming rounded lobe in lateral view; forming pair of about
<quantity id="4C9DECFFFF8F1E60FAE5F471E986529C" box="[1346,1436,1900,1926]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="56" unit="mm" value="0.1">0.1 mm</quantity>
long tapering processes covered by setae in ventral view. Main branch of inferior appendages narrowing stepwise apically in lateral view; proximal part about as wide as tergite IX in lateral view; distal one-fourth about as wide as superior appendages in lateral view; generally curving slightly ventrad along entire length; about
<quantity id="4C9DECFFFF8E1E61FF44F385ED4A55A8" box="[227,336,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="57" unit="mm" value="0.35">0.35 mm</quantity>
long. Phallic apparatus connected to sternite IX and tergite IX by long, straight, dorsally narrowing sternal strips; sternal strip with convex anterodorsal margin and nearly straight posteroventral margin, about
<quantity id="4C9DECFFFF8E1E61FE98F3FFEDB255E6" box="[319,424,226,252]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="57" unit="mm" value="0.25">0.25 mm</quantity>
long. Phallobase short, rectangular. Median processes more than
<quantity id="4C9DECFFFF8E1E61FB04F3FFE8E655E6" box="[1187,1276,226,252]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="57" unit="mm" value="0.6">0.6 mm</quantity>
long; needleshaped, with row of dorsal teeth before apex; bow-shaped in lateral view; apex not exceeding phallicata. Phallicata about
<quantity id="4C9DECFFFF8E1E61FEF9F230EDA3545D" box="[350,441,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="57" unit="mm" value="0.5">0.5 mm</quantity>
long; proximal one-third uniformly slender, straight; double as wide at mid-length, widening toward ventrally curved apex, with concave ventral margin, dorsal margin undulating. Paramere separated from phallicata well posterior of phallobase; very thin, straight along its length, except curving at apex; running parallel with proximal part of phallicata, exceeding phallicata; about
<quantity id="4C9DECFFFF8E1E61FBCFF280E8CB54AD" box="[1128,1233,413,439]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="57" unit="mm" value="0.55">0.55 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8E1E61FF63F2DFE8FF5718" pageId="3" pageNumber="57" type="materials_examined">
<paragraph id="8BDA411AFF8E1E61FF63F2DFE87454C6" blockId="3.[151,1437,152,588]" box="[196,1134,450,476]" pageId="3" pageNumber="57">
<emphasis id="B9119D08FF8E1E61FF63F2DFED2C54C6" bold="true" box="[196,310,450,476]" pageId="3" pageNumber="57">
<typeStatus id="54DEFFB8FF8E1E61FF63F2DFED2C54C6" box="[196,310,450,476]" pageId="3" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
male, Namoroka, Vilanandro,
<date id="FFDB67DAFF8E1E61FD07F2DFEEE254C6" box="[672,760,450,476]" pageId="3" pageNumber="57" value="1952-09">ix.1952</date>
[R. Paulian] (
<collectionCode id="ED74D9DFFF8E1E61FC3BF2DFEFE154C6" LSID="urn:lsid:biocol.org:col:34988" box="[924,1019,450,476]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="57">MNHN</collectionCode>
, alcohol).
</paragraph>
<paragraph id="8BDA411AFF8E1E61FF63F2FAE8FF5718" blockId="3.[151,1437,152,588]" box="[196,1253,487,514]" pageId="3" pageNumber="57">
<emphasis id="B9119D08FF8E1E61FF63F2FAED51571B" bold="true" box="[196,331,487,513]" pageId="3" pageNumber="57">
<typeStatus id="54DEFFB8FF8E1E61FF63F2FAED5C571B" box="[196,326,487,513]" pageId="3" pageNumber="57" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="54DEFFB8FF8E1E61FE55F2F5EE415718" box="[498,603,488,514]" pageId="3" pageNumber="57" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D638A93FF8E1E61FDC0F2F5EEC35718" box="[615,729,488,514]" pageId="3" pageNumber="57" type="female">2 females</specimenCount>
(
<collectionCode id="ED74D9DFFF8E1E61FD4FF2F5EF5C5718" LSID="urn:lsid:biocol.org:col:34988" box="[744,838,488,514]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="57">MNHN</collectionCode>
, alcohol),
<specimenCount id="9D638A93FF8E1E61FC67F2F5E80A5718" box="[960,1040,488,514]" pageId="3" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED74D9DFFF8E1E61FBB8F2F5E8685718" box="[1055,1138,488,514]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="3" pageNumber="57">NHRS</collectionCode>
, alcohol).
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8E1E61FF63F110EF14573D" box="[196,782,525,551]" pageId="3" pageNumber="57" type="distribution">
<paragraph id="8BDA411AFF8E1E61FF63F110EF14573D" blockId="3.[151,1437,152,588]" box="[196,782,525,551]" pageId="3" pageNumber="57">
<emphasis id="B9119D08FF8E1E61FF63F110ED7E573D" bold="true" box="[196,356,525,551]" pageId="3" pageNumber="57">Distribution.</emphasis>
<collectingCountry id="F372018AFF8E1E61FECBF110EDE0573D" box="[364,506,525,551]" name="Madagascar" pageId="3" pageNumber="57">Madagascar</collectingCountry>
(Province Mahajanga).
</paragraph>
</subSubSection>
<subSubSection id="C37F1291FF8E1E61FF63F12FE8565756" box="[196,1100,562,588]" pageId="3" pageNumber="57" type="etymology">
<paragraph id="8BDA411AFF8E1E61FF63F12FE8565756" blockId="3.[151,1437,152,588]" box="[196,1100,562,588]" pageId="3" pageNumber="57">
<emphasis id="B9119D08FF8E1E61FF63F12FED4B5756" bold="true" box="[196,337,562,588]" pageId="3" pageNumber="57">Etymology.</emphasis>
The name,
<taxonomicName id="4C653A99FF8E1E61FE7CF12FEE355751" box="[475,559,562,587]" class="Insecta" family="Psychomyiidae" genus="Paduniella" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="nandra">
<emphasis id="B9119D08FF8E1E61FE7CF12FEE355751" box="[475,559,562,587]" italics="true" pageId="3" pageNumber="57">nandra</emphasis>
</taxonomicName>
, is derived from Vilanandro, the
<typeStatus id="54DEFFB8FF8E1E61FC13F12FEFFC5756" box="[948,998,562,588]" pageId="3" pageNumber="57">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>