594 lines
71 KiB
XML
594 lines
71 KiB
XML
<document id="D890D526C76D78598271D9552B518B08" ID-CLB-Dataset="42393" ID-DOI="10.11646/zootaxa.3905.3.10" ID-GBIF-Dataset="f86fcc41-5623-42c3-9b4c-e8affdc8fd25" ID-ISSN="1175-5326" ID-Zenodo-Dep="245312" ID-ZooBank="5A9DBB7C-BCAD-40B2-AFCD-0CFF2D4F643" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460934229620" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2015" docId="03CBD108FFA3E11796C902D38FD7FB33" docLanguage="en" docName="zt03905p446.pdf" docOrigin="Zootaxa 3905 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yola matsikammae Bilton, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="446" masterDocId="FFF2A970FFA2E112965E03158A54FFAE" masterDocTitle="A new species of Yo l a Gozis, 1886 from the Western Cape of South Africa (Coleoptera: Dytiscidae: Bidessini)" masterLastPageNumber="446" masterPageNumber="441" pageNumber="442" updateTime="1698582589653" updateUser="ExternalLinkService">
|
||
<mods:mods id="35A3B512FAD9760EC514732F54CA8DBE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A0EB42F41B17647ED3A8F71D38C02556">
|
||
<mods:title id="1B1A1A2F657C3F5E194ED78379F3ED6D">A new species of Yo l a Gozis, 1886 from the Western Cape of South Africa (Coleoptera: Dytiscidae: Bidessini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="82267D073F493BC35EB56D1E2B5DC78F" type="personal">
|
||
<mods:role id="4AF384B90E6FC5C9B4A12D9447A020D3">
|
||
<mods:roleTerm id="0DC9226869352A2D020EC81C2567784C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A70286F11541D3C163EABE7E2EE45B70">Bilton, David T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B0926A1A23BB50F983C6D6F30BB7DDC9">text</mods:typeOfResource>
|
||
<mods:relatedItem id="445193ACB584EA01A9F2119A3DE32975" type="host">
|
||
<mods:titleInfo id="4A46FB6797BC6C0296DDA28FEBA02656">
|
||
<mods:title id="113FCEEFF611F29A1EBBCFE131375DF9">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9373AF0CD58AE4AE19C478F0DF25328E">
|
||
<mods:date id="941C7FEFE7DB0E047A837AE8FA3B3776">2015</mods:date>
|
||
<mods:detail id="B6203A4C8C61BB64431B1C67AC85FA50" type="volume">
|
||
<mods:number id="C74045798158032BA174288C9718D37E">3905</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BE7B04ECDCF9F07DB1A067ED9E1B6581" type="issue">
|
||
<mods:number id="57836DBB63791B904510B43A59B24AC2">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C875AB08342E55C0944DF9984F8F118B" unit="page">
|
||
<mods:start id="CA3D126D153F745DF04D16DBFA620E53">441</mods:start>
|
||
<mods:end id="0F53EBD65AB9F15F5FB1B4446D5DBCA3">446</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="4FB0DA6682DB0D869363D667138A5F2C">journal article</mods:classification>
|
||
<mods:identifier id="4434064726FD5B78C57A2951A3B04FD9" type="CLB-Dataset">42393</mods:identifier>
|
||
<mods:identifier id="46E1818FCD671FF0E452BE1FE2F7F8FE" type="DOI">10.11646/zootaxa.3905.3.10</mods:identifier>
|
||
<mods:identifier id="D08AC4907DECBF1BE3353B903992C0E1" type="GBIF-Dataset">f86fcc41-5623-42c3-9b4c-e8affdc8fd25</mods:identifier>
|
||
<mods:identifier id="B7F58840B83283972919505F6E1C4942" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="E986B7948FF567D3DD3C8C4A156E44A8" type="Zenodo-Dep">245312</mods:identifier>
|
||
<mods:identifier id="49EFAAF540A02EB540A6A325661CCB8B" type="ZooBank">5A9DBB7C-BCAD-40B2-AFCD-0CFF2D4F643</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CBD108FFA3E11796C902D38FD7FB33" ID-DOI="http://doi.org/10.5281/zenodo.6093402" ID-GBIF-Taxon="119590549" ID-Zenodo-Dep="6093402" LSID="urn:lsid:plazi:treatment:03CBD108FFA3E11796C902D38FD7FB33" httpUri="http://treatment.plazi.org/id/03CBD108FFA3E11796C902D38FD7FB33" lastPageId="5" lastPageNumber="446" pageId="1" pageNumber="442">
|
||
<subSubSection id="C3783395FFA3E11396C902D38B60FDAF" pageId="1" pageNumber="442" type="nomenclature">
|
||
<paragraph id="8BDD601EFFA3E11396C902D38B8AFE4E" blockId="1.[151,478,454,514]" box="[151,478,454,480]" pageId="1" pageNumber="442">
|
||
<heading id="D095D772FFA3E11396C902D38B8AFE4E" bold="true" box="[151,478,454,480]" fontSize="11" level="1" pageId="1" pageNumber="442" reason="1">
|
||
<emphasis id="B916BC0CFFA3E11396C902D38B8AFE4E" bold="true" box="[151,478,454,480]" pageId="1" pageNumber="442">
|
||
<taxonomicName id="4C621B9DFFA3E11396C902D38B2CFE4E" ID-CoL="8LZRZ" box="[151,376,454,480]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA3E11396C902D38B2CFE4E" bold="true" box="[151,376,454,480]" italics="true" pageId="1" pageNumber="442">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2250177FFA3E11397DE02D38B8AFE4E" box="[384,478,454,480]" pageId="1" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E11396C902FC8B60FDAF" blockId="1.[151,478,454,514]" box="[151,308,489,514]" pageId="1" pageNumber="442">
|
||
(
|
||
<figureCitation id="13597C9BFFA3E11396C102FC8AB1FDAF" box="[159,229,489,514]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="1" pageNumber="442">Figs 1</figureCitation>
|
||
and
|
||
<figureCitation id="13597C9BFFA3E113974202FC8B7FFDAF" box="[284,299,489,513]" captionStart="FIGURE 2" captionStartId="3.[151,250,1821,1843]" captionTargetBox="[151,1436,384,1799]" captionTargetId="figure@3.[151,1436,193,1799]" captionTargetPageId="3" captionText="FIGURE 2. Type locality of Yola matsikammae sp. nov., South Africa, Western Cape, Matsikammaberg, permanent stream ca. 1 km SE of Sewefontein Farm. Photo D. T. Bilton." httpUri="https://zenodo.org/record/245314/files/figure.png" pageId="1" pageNumber="442">2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3783395FFA3E11396C901248990FC8C" pageId="1" pageNumber="442" type="materials_examined">
|
||
<paragraph id="8BDD601EFFA3E11396C9012488BCFDC0" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
<emphasis id="B916BC0CFFA3E11396C901248B65FDE4" bold="true" box="[151,305,561,586]" pageId="1" pageNumber="442">
|
||
<typeStatus id="54D9DEBCFFA3E11396C901248A86FDE4" box="[151,210,561,586]" pageId="1" pageNumber="442">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B916BC0CFFA3E113971A01248BB4FDE4" bold="true" box="[324,480,561,586]" pageId="1" pageNumber="442">
|
||
<collectingCountry id="F375208EFFA3E113971A01248BB4FDE4" box="[324,480,561,586]" name="South Africa" pageId="1" pageNumber="442">South Africa</collectingCountry>
|
||
</emphasis>
|
||
, Western Cape, Matsikammaberg, permanent stream ca.
|
||
<quantity id="4C9ACDFBFFA3E113922501248EECFDE4" box="[1147,1208,561,586]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="442" unit="km" value="1.0">1 km</quantity>
|
||
SE of Sewefontein Farm,
|
||
<geoCoordinate id="EE5606D9FFA3E11396BE01438BD4FDC0" box="[224,384,598,622]" direction="south" orientation="latitude" pageId="1" pageNumber="442" precision="1" value="-31.696688">31°41'48.08''S</geoCoordinate>
|
||
<geoCoordinate id="EE5606D9FFA3E11397D601438879FDC0" box="[392,557,598,622]" direction="east" orientation="longitude" pageId="1" pageNumber="442" precision="1" value="18.83012">18°49'48.43''E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D64AB97FFA3E1139466014088D3FDC3" box="[568,647,597,622]" pageId="1" pageNumber="442" type="generic">
|
||
<quantity id="4C9ACDFBFFA3E1139466014088D6FDC3" box="[568,642,597,622]" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="1" pageNumber="442" unit="m" value="617.0">617 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
(
|
||
<figureCitation id="13597C9BFFA3E11394CB0140888FFDC0" box="[661,731,597,622]" captionStart="FIGURE 2" captionStartId="3.[151,250,1821,1843]" captionTargetBox="[151,1436,384,1799]" captionTargetId="figure@3.[151,1436,193,1799]" captionTargetPageId="3" captionText="FIGURE 2. Type locality of Yola matsikammae sp. nov., South Africa, Western Cape, Matsikammaberg, permanent stream ca. 1 km SE of Sewefontein Farm. Photo D. T. Bilton." httpUri="https://zenodo.org/record/245314/files/figure.png" pageId="1" pageNumber="442">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E1139699016C8E75FD74" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
<emphasis id="B916BC0CFFA3E1139699016C8B26FD3C" bold="true" box="[199,370,633,658]" pageId="1" pageNumber="442">
|
||
<typeStatus id="54D9DEBCFFA3E1139699016C8B56FD3C" box="[199,258,633,658]" pageId="1" pageNumber="442">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B916BC0CFFA3E11397D8016C8BA7FD3C" bold="true" box="[390,499,633,658]" pageId="1" pageNumber="442">
|
||
<typeStatus id="54D9DEBCFFA3E11397D8016C8BA7FD3C" box="[390,499,633,658]" pageId="1" pageNumber="442" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(male): “
|
||
<date id="FFDC46DEFFA3E1139438016C88B6FD3C" box="[614,738,633,658]" pageId="1" pageNumber="442" value="2014-09-22">22/ix/2014</date>
|
||
<collectingCountry id="F375208EFFA3E11394B0016C89D6FD3C" box="[750,898,633,658]" name="South Africa" pageId="1" pageNumber="442">South Africa</collectingCountry>
|
||
WC// Matsikammaberg stream
|
||
<quantity id="4C9ACDFBFFA3E11392A0016F8F6FFD3C" box="[1278,1339,633,658]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="442" unit="km" value="1.0">1 km</quantity>
|
||
// SE of Sewefontein Farm// permanent D T Bilton leg.” (genitalia extracted and mounted in
|
||
<collectionCode id="ED73F8DBFFA3E113921B018B8EC2FD1B" box="[1093,1174,670,693]" pageId="1" pageNumber="442">DMHF</collectionCode>
|
||
on same card) with red printed
|
||
<typeStatus id="54D9DEBCFFA3E11396B301D48B1BFD74" box="[237,335,705,730]" pageId="1" pageNumber="442" type="holotype">holotype</typeStatus>
|
||
label “
|
||
<typeStatus id="54D9DEBCFFA3E11397C301D48852FD74" box="[413,518,705,730]" pageId="1" pageNumber="442" type="holotype">Holotype</typeStatus>
|
||
//
|
||
<taxonomicName id="4C621B9DFFA3E113944101D488A4FD77" box="[543,752,705,729]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA3E113944101D488A4FD77" box="[543,752,705,729]" italics="true" pageId="1" pageNumber="442">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA3E11394A901D48905FD74" bold="true" box="[759,849,705,730]" pageId="1" pageNumber="442">
|
||
<taxonomicNameLabel id="A2250177FFA3E11394A901D48905FD74" box="[759,849,705,730]" pageId="1" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
// Bilton” (
|
||
<collectionCode id="ED73F8DBFFA3E113959701D78E41FD74" box="[969,1045,706,730]" pageId="1" pageNumber="442">ISAM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E113969901F08990FC8C" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
<emphasis id="B916BC0CFFA3E113969901F08B1CFD50" bold="true" box="[199,328,741,766]" pageId="1" pageNumber="442">
|
||
<typeStatus id="54D9DEBCFFA3E113969901F08B17FD50" box="[199,323,741,766]" pageId="1" pageNumber="442" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D64AB97FFA3E113970C01F38B23FD50" box="[338,375,742,766]" pageId="1" pageNumber="442" type="generic">3♂</specimenCount>
|
||
, 9♀ same data as
|
||
<typeStatus id="54D9DEBCFFA3E113940E01F088E6FD50" box="[592,690,741,766]" pageId="1" pageNumber="442" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED73F8DBFFA3E113949A01F38947FD50" box="[708,787,742,766]" pageId="1" pageNumber="442">CDTB</collectionCode>
|
||
,
|
||
<collectionCode id="ED73F8DBFFA3E113957F01F38921FD50" box="[801,885,742,766]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="1" pageNumber="442">DMSA</collectionCode>
|
||
,
|
||
<collectionCode id="ED73F8DBFFA3E11395DD01F38998FD50" box="[899,972,742,766]" pageId="1" pageNumber="442">ISAM</collectionCode>
|
||
,
|
||
<collectionCode id="ED73F8DBFFA3E113958701F38E75FD53" box="[985,1057,742,765]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="442">NMW</collectionCode>
|
||
,
|
||
<collectionCode id="ED73F8DBFFA3E113927101F38E29FD50" LSID="urn:lsid:biocol.org:col:34248" box="[1071,1149,742,766]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="442">SANC</collectionCode>
|
||
,
|
||
<collectionCode id="ED73F8DBFFA3E11392D501F38EAEFD50" box="[1163,1274,742,766]" pageId="1" pageNumber="442">OUMNH</collectionCode>
|
||
). All with red printed
|
||
<typeStatus id="54D9DEBCFFA3E11396B3001F8B19FC8C" box="[237,333,778,802]" pageId="1" pageNumber="442" type="paratype">paratype</typeStatus>
|
||
labels “
|
||
<typeStatus id="54D9DEBCFFA3E11397F9001F885EFC8C" box="[423,522,778,802]" pageId="1" pageNumber="442" type="paratype">Paratype</typeStatus>
|
||
//
|
||
<taxonomicName id="4C621B9DFFA3E113947C001C88A0FC8F" box="[546,756,777,801]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA3E113947C001C88A0FC8F" box="[546,756,777,801]" italics="true" pageId="1" pageNumber="442">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA3E11394A5001C8900FC8C" bold="true" box="[763,852,777,802]" pageId="1" pageNumber="442">
|
||
<taxonomicNameLabel id="A2250177FFA3E11394A5001C8900FC8C" box="[763,852,777,802]" pageId="1" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
// Bilton”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3783395FFA3E1169699003889D9F9B6" lastPageId="4" lastPageNumber="445" pageId="1" pageNumber="442" type="description">
|
||
<paragraph id="8BDD601EFFA3E113969900388F47FCC4" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
<emphasis id="B916BC0CFFA3E113969900388B05FCE8" bold="true" box="[199,337,813,838]" pageId="1" pageNumber="442">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="54D9DEBCFFA3E11397C30038885EFCE8" box="[413,522,813,838]" pageId="1" pageNumber="442" type="holotype">Holotype</typeStatus>
|
||
: body length (to elytral apices)
|
||
<quantity id="4C9ACDFBFFA3E1139536003B8995FCEB" box="[872,961,814,838]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="442" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; maximum width (elytra)
|
||
<quantity id="4C9ACDFBFFA3E11392BD00388F1EFCEB" box="[1251,1354,813,838]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="442" unit="mm" value="1.25">1.25 mm</quantity>
|
||
; elytral length
|
||
<quantity id="4C9ACDFBFFA3E11396BA00448B1FFCC7" box="[228,331,849,874]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="442" unit="mm" value="1.65">1.65 mm</quantity>
|
||
. Same values for
|
||
<typeStatus id="54D9DEBCFFA3E113944F004788D5FCC4" box="[529,641,850,874]" pageId="1" pageNumber="442" type="paratype">paratypes</typeStatus>
|
||
:
|
||
<quantity id="4C9ACDFBFFA3E11394D500448976FCC7" box="[651,802,849,874]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.45" metricValueMin="2.3" pageId="1" pageNumber="442" unit="mm" value="2.375" valueMax="2.45" valueMin="2.3">2.3–2.45 mm</quantity>
|
||
,
|
||
<quantity id="4C9ACDFBFFA3E1139570004789E1FCC7" box="[814,949,850,874]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="442" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
and
|
||
<quantity id="4C9ACDFBFFA3E11395B500448ED4FCC7" box="[1003,1152,849,874]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.85" metricValueMin="1.6" pageId="1" pageNumber="442" unit="mm" value="1.725" valueMax="1.85" valueMin="1.6">1.6–1.85 mm</quantity>
|
||
respectively.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E1139699006088CCFAB7" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
Colour (
|
||
<figureCitation id="13597C9BFFA3E113977800608B3EFC20" box="[294,362,885,910]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="1" pageNumber="442">Fig. 1</figureCitation>
|
||
): Dorsal surface predominantly yellowish brown, straw coloured. Head and pronotum darker than ground colour of elytra. Frons darker than clypeus; infuscated along inner border of eyes. Front and hind margins of pronotum with a black band; anterior band present over central 4/5 of width, between projections of anterior angles. Posterior band more well-developed between lateral plicae, here extending for approximately 1/5 of pronotal length. Elytra with diffuse dark markings, boundaries rather ill-defined. Anterior margins with narrow dark band, laterally almost reaching shoulders. Suture dark to apex, with diffuse dark markings expanded laterally from suture; interrupted in centre and reaching lateral keel anteriorly and elytral margin posteriorly (
|
||
<figureCitation id="13597C9BFFA3E113936D07588F22FBC8" box="[1331,1398,1101,1126]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="1" pageNumber="442">Fig. 1</figureCitation>
|
||
B). Maxillary palpi pale straw yellow, last segment infuscated. Fore and mid legs straw yellow with femuro-tibial junctions and tarsi infuscated.
|
||
<collectingCountry id="F375208EFFA3E11397A80780887AFB00" box="[502,558,1173,1198]" name="India" pageId="1" pageNumber="442">Hind</collectingCountry>
|
||
legs same but femuro-tibial and tibio-tarsal junctions infuscated, along with apical 1/3–1/2 of tarsal segments. Antennae with segments 1–5 pale yellow, 6–10 with apices infuscated (more so in later segments) and segment 11 infuscated throughout. Venter reddish brown, abdomen dark pitchy brown, particularly towards lateral margins and apex.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E113969906308FD4F994" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">Head: Broad, with large eyes. Clypeus somewhat thickened and evenly rounded, lacking anterior border, and not separated from rest of head. Head raised in front of eye, above antennal insertion, into a distinct, low tubercle. Frons slightly elevated in centre. Cervical line well marked. Head in front of cervical line with coarse, often confluent punctures, punctures particularly large mediodistally on frons; many punctures bearing small, peg-like setae. Anterior half of frons and clypeus with smaller sparse punctures. Centre of frons and clypeus shining, lacking microreticulation. Areas immediately inside eyes with shallow, isodiametric microreticulation, particularly anteriorly and around frontal tubercles, giving a duller appearance. Anterior margin of clypeus shining, but with traces of transverse microreticulation. Head behind cervical line duller, with strong transverse microreticulation.</paragraph>
|
||
<paragraph id="8BDD601EFFA3E113969905538E8AF898" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
Pronotum: Transverse rectangular. Sides rounded; broadest just before middle.
|
||
<collectingCountry id="F375208EFFA3E11392DA05508EE8F9F0" box="[1156,1212,1605,1630]" name="India" pageId="1" pageNumber="442">Hind</collectingCountry>
|
||
margins distinctly bisinuate around middle. Lateral plicae well developed, relatively deep, and open interiorly. Plicae deepest at posterior ends; sinuate and becoming shallow anteriorly, here extending to approximately 1/3 away from front margin of pronotum. Posterior 1/3 of disc between plicae depressed, especially in centre, surface here raised abruptly to level of remainder of disc. Pronotal surface shining, lacking microreticulation, but roughened due to dense, coarse punctation. Punctures largest and closest towards centre, smaller towards lateral margins. Punctures bearing flattened, white recumbent setae in centre; peg-like towards posterior and lateral margins.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA3E113969904548F57F844" blockId="1.[151,1437,561,2026]" pageId="1" pageNumber="442">
|
||
Elytra: Together forming an elongate oval, with point of maximum width just behind middle. Lateral margin rounded to front angles, parallel over central 1/3, and tapered towards apex in posterior 1/2. Each elytron with three keels. Discal keel very strong, gradually elevated both anteriorly and posteriorly and of even height over central 4/ 5, occupying ca. 2/3 of elytron (
|
||
<figureCitation id="13597C9BFFA3E11397A704B8886CF868" box="[505,568,1965,1990]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="1" pageNumber="442">Fig. 1</figureCitation>
|
||
B). Median keel lower, running from shoulder to just in front of posterior end of discal keel. Lateral keel slightly lower still, same length as median keel. Elytra shining, lacking
|
||
</paragraph>
|
||
<caption id="DF1D3096FFA0E11096C904808EEEF864" httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="2" pageNumber="443" targetBox="[160,1410,202,1892]" targetPageId="2">
|
||
<paragraph id="8BDD601EFFA0E11096C904808EEEF864" blockId="2.[151,1436,1941,1994]" pageId="2" pageNumber="443">
|
||
<emphasis id="B916BC0CFFA0E11096C904808B42F804" bold="true" box="[151,278,1941,1963]" pageId="2" pageNumber="443">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C621B9DFFA0E110977E04808BB5F804" box="[288,481,1941,1962]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA0E110977E04808BB5F804" box="[288,481,1941,1962]" italics="true" pageId="2" pageNumber="443">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA0E11097B204808814F804" bold="true" box="[492,576,1941,1962]" pageId="2" pageNumber="443">
|
||
<taxonomicNameLabel id="A2250177FFA0E11097B204808814F804" box="[492,576,1941,1962]" pageId="2" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A–B = 1 mm; C = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1D3096FFA1E11196C9040888CCF8FC" httpUri="https://zenodo.org/record/245314/files/figure.png" pageId="3" pageNumber="444" targetBox="[151,1436,384,1799]" targetPageId="3">
|
||
<paragraph id="8BDD601EFFA1E11196C9040888CCF8FC" blockId="3.[151,1436,1821,1874]" pageId="3" pageNumber="444">
|
||
<emphasis id="B916BC0CFFA1E11196C904088B46F89C" bold="true" box="[151,274,1821,1843]" pageId="3" pageNumber="444">FIGURE 2.</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="4C621B9DFFA1E11197E004088828F89C" box="[446,636,1821,1842]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA1E11197E004088828F89C" box="[446,636,1821,1842]" italics="true" pageId="3" pageNumber="444">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA1E11194DC04088886F89C" bold="true" box="[642,722,1821,1842]" pageId="3" pageNumber="444">
|
||
<taxonomicNameLabel id="A2250177FFA1E11194DC04088886F89C" box="[642,722,1821,1842]" pageId="3" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, South Africa, Western Cape, Matsikammaberg, permanent stream ca. 1 km SE of Sewefontein Farm. Photo D. T. Bilton.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDD601EFFA1E11196C9046B8B18F870" blockId="3.[151,1437,1918,2015]" pageId="3" pageNumber="444">microreticulation. Punctate, punctures close, coarse and shallow anteriorly, becoming finer and sparser posteriorly. Punctures almost confluent in anterior 1/2, between discal and medial keels. Each puncture bearing a flattened decumbent seta.</paragraph>
|
||
<paragraph id="8BDD601EFFA6E1169699038D8ED7FC4E" blockId="4.[151,1437,151,1560]" pageId="4" pageNumber="445">Venter: Mentum and prementum smooth and shining, lacking microreticulation. Anterior margin of prementum with short, stout bristles. Anterior and lateral margins of mentum thickened and somewhat darkened. Gula and genae shining, lacking microreticulation. Gular sutures weak; gula with scattered shallow medium punctures. Subgenal ridges well-marked, area between ridge and eye with open, isodiametric microreticulation. Pronotal hypomera smooth and shining, with scattered shallow punctures bearing long golden setae, and traces of transverse wrinkles posteriorly. Proepisternum shining, minutely wrinkled, especially posteriorly. Prosternum shining, strongly punctate, especially in centre where punctures are deeper and almost confluent in places. Punctures in centre of prosternum with long, fine golden setae. Prosternal process raised, neck of process making an angle of approx. 70° with prosternum. Process lanceolate, with broad, shallow central groove furnished with dense, coarse, punctures bearing long, decumbent setae. Neck of process with sparse small punctures bearing hairlike, golden setae. Prosternal process meeting anterior projection of metaventrite. Metaventrite and metathoracic anepisternum shining, with sparse, shallow punctures; traces of microreticulation visible towards centre of metaventrite. Punctures somewhat larger on metaventrite and deeper anteriorly, each bearing a short, flattened, decumbent seta. Metacoxa shining, with large, shallow, almost confluent punctures bearing short, flattened, decumbent setae. Elytral epipleura shining, with close, coarse, setose punctures which are almost confluent in posterior 1/2. Metacoxal lines well marked, subparallel, extending forwards as lines onto metaventrites, these converging towards its anterior margin. Metacoxal processes with shallow open central channel extending forwards onto metaventrite; foveate at junction of metacoxae and metaventrite. Processes punctate, with coarse, shallow punctures, each bearing a short, flat, recumbent seta. Apex of metacoxal processes fused to abdomen; sinuate, lobes weakly developed and not covering leg insertions. Abdominal ventrites 2–4 fused, junctions weakly visible. Abdominal ventrites shining, 2–4 densely punctate; punctures smaller and more distinct in centre, larger and more confluent laterally; bearing flattened, decumbent setae. Abdominal ventrites 5–7 with smaller, sparser punctures, each bearing a golden recumbent seta, longer than those on preceding ventrites. Abdominal ventrites 5–6 with central tuft of long, golden, hair-like setae, extending almost to abdominal apex.</paragraph>
|
||
<paragraph id="8BDD601EFFA6E116969900FE89F8FB89" blockId="4.[151,1437,151,1560]" pageId="4" pageNumber="445">
|
||
Aedeagus: Median lobe characteristically shaped (
|
||
<figureCitation id="13597C9BFFA6E116954F00FE8901FBAD" box="[785,853,1003,1028]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="4" pageNumber="445">Fig. 1</figureCitation>
|
||
C); apex flattened and spathulate in ventral view. Parameres (
|
||
<figureCitation id="13597C9BFFA6E1169745071B8B0DFB89" box="[283,345,1038,1063]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="4" pageNumber="445">Fig. 1</figureCitation>
|
||
C) two-segmented, with bluntly curved, hooked tips.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E116969907278F67FB1B" blockId="4.[151,1437,151,1560]" pageId="4" pageNumber="445">
|
||
Females:
|
||
<collectingCountry id="F375208EFFA6E116976907278B02FBE4" box="[311,342,1074,1098]" name="American Samoa" pageId="4" pageNumber="445">As</collectingCountry>
|
||
males except head, pronotum and elytra entirely microreticulate between punctures, giving a duller appearance. Venter duller, with fine microreticulation throughout. Venter paler than in male, yellowish brown throughout except for neck of prosternal process and segmental junctions, which are slightly infuscated. Fore and mid tarsi slightly narrower and noticeably shorter than in males, particularly last tarsal segment.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E116969907AA8824FB52" blockId="4.[151,1437,151,1560]" pageId="4" pageNumber="445">
|
||
<emphasis id="B916BC0CFFA6E116969907AA8B11FB76" bold="true" box="[199,325,1215,1240]" pageId="4" pageNumber="445">Variability</emphasis>
|
||
.
|
||
<typeStatus id="54D9DEBCFFA6E116970607D48B91FB77" box="[344,453,1217,1241]" pageId="4" pageNumber="445" type="paratype">Paratypes</typeStatus>
|
||
vary slightly in size (see above) and the development of the elytral pattern, some specimens being darker than the
|
||
<typeStatus id="54D9DEBCFFA6E116945807F68838FB52" box="[518,620,1251,1276]" pageId="4" pageNumber="445" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E1169699061389D9F9B6" blockId="4.[151,1437,151,1560]" pageId="4" pageNumber="445">
|
||
<emphasis id="B916BC0CFFA6E116969906138F76FAB1" bold="true" box="[199,1314,1286,1311]" pageId="4" pageNumber="445">
|
||
Differential diagnosis and key to southern African members of the
|
||
<taxonomicName id="4C621B9DFFA6E116924406138E9BFAB1" box="[1050,1231,1286,1311]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="bicarinata">
|
||
<emphasis id="B916BC0CFFA6E116924406138E9BFAB1" bold="true" box="[1050,1231,1286,1311]" italics="true" pageId="4" pageNumber="445">Yola bicarinata</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</emphasis>
|
||
. The new species is apparently a member of the
|
||
<taxonomicName id="4C621B9DFFA6E116941B063F88EEFAEC" box="[581,698,1322,1346]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="bicarinata">
|
||
<emphasis id="B916BC0CFFA6E116941B063F88EEFAEC" box="[581,698,1322,1346]" italics="true" pageId="4" pageNumber="445">bicarinata</emphasis>
|
||
</taxonomicName>
|
||
species group, sensu
|
||
<bibRefCitation id="EFF31DEFFFA6E11695F3063F8E08FAED" author="Bistrom" box="[941,1116,1322,1347]" pageId="4" pageNumber="445" refString="Bistrom, O. (1983) Revision of the genera Yola Des Gozis and Yolina Guignot. Acta Zoologica Fennica, 176, 1 - 67." type="journal article" year="1983">Biström (1983)</bibRefCitation>
|
||
, having three distinct elytral keels with discal keels that descend gradually to the elytral surface (
|
||
<figureCitation id="13597C9BFFA6E11695D50658899DFAC8" box="[907,969,1357,1382]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="4" pageNumber="445">Fig. 1</figureCitation>
|
||
B). Within this group
|
||
<taxonomicName id="4C621B9DFFA6E11692E1065A8F20FAC8" box="[1215,1396,1358,1382]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA6E11692E1065A8F20FAC8" box="[1215,1396,1358,1382]" italics="true" pageId="4" pageNumber="445">
|
||
<collectingCountry id="F375208EFFA6E11692E1065A8E84FAC8" box="[1215,1232,1359,1382]" name="Italy" pageId="4" pageNumber="445">Y</collectingCountry>
|
||
. matsikammae
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA6E116932406588A9CFA24" bold="true" pageId="4" pageNumber="445">
|
||
<taxonomicNameLabel id="A2250177FFA6E116932406588A9CFA24" pageId="4" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
would key to the Palaearctic
|
||
<taxonomicName id="4C621B9DFFA6E116947A0667892EFA24" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" box="[548,890,1393,1418]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="bicarinata">
|
||
<emphasis id="B916BC0CFFA6E116947A066788E3FA27" box="[548,695,1393,1417]" italics="true" pageId="4" pageNumber="445">
|
||
<collectingCountry id="F375208EFFA6E116947A06678861FA27" box="[548,565,1394,1417]" name="Italy" pageId="4" pageNumber="445">Y</collectingCountry>
|
||
. bicarinata
|
||
</emphasis>
|
||
(Latreille, 1804)
|
||
</taxonomicName>
|
||
using
|
||
<bibRefCitation id="EFF31DEFFFA6E116959506648E2FFA24" author="Bistrom" box="[971,1147,1393,1418]" pageId="4" pageNumber="445" refString="Bistrom, O. (1983) Revision of the genera Yola Des Gozis and Yolina Guignot. Acta Zoologica Fennica, 176, 1 - 67." type="journal article" year="1983">Biström (1983)</bibRefCitation>
|
||
, which at
|
||
<quantity id="4C9ACDFBFFA6E11692A806678FD4FA27" box="[1270,1408,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="4" pageNumber="445" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
|
||
is smaller than the new species, as well as being more strongly rounded, and differing in genital anatomy (median lobe of aedeagus pointed and needle-like in ventral view in
|
||
<emphasis id="B916BC0CFFA6E116957006AC8981FA7E" box="[814,981,1464,1488]" italics="true" pageId="4" pageNumber="445">
|
||
<collectingCountry id="F375208EFFA6E116957006AC896BFA7E" box="[814,831,1465,1488]" name="Italy" pageId="4" pageNumber="445">Y</collectingCountry>
|
||
. bicarinata—
|
||
</emphasis>
|
||
see
|
||
<bibRefCitation id="EFF31DEFFFA6E116925E06AD8EF0FA7E" author="Bistrom" box="[1024,1188,1464,1489]" pageId="4" pageNumber="445" refString="Bistrom, O. (1983) Revision of the genera Yola Des Gozis and Yolina Guignot. Acta Zoologica Fennica, 176, 1 - 67." type="journal article" year="1983">Biström, 1983</bibRefCitation>
|
||
). The new species can be distinguished from other southern African members of the
|
||
<taxonomicName id="4C621B9DFFA6E116951706C989EAFA5A" box="[841,958,1500,1524]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="bicarinata">
|
||
<emphasis id="B916BC0CFFA6E116951706C989EAFA5A" box="[841,958,1500,1524]" italics="true" pageId="4" pageNumber="445">bicarinata</emphasis>
|
||
</taxonomicName>
|
||
group as follows (for detailed descriptions and illustrations of previously described species see
|
||
<bibRefCitation id="EFF31DEFFFA6E11694BC06EA89D4F9B6" author="Bistrom" box="[738,896,1535,1560]" pageId="4" pageNumber="445" refString="Bistrom, O. (1983) Revision of the genera Yola Des Gozis and Yolina Guignot. Acta Zoologica Fennica, 176, 1 - 67." type="journal article" year="1983">Biström 1983</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3783395FFA6E11796C905538F0EFE68" lastPageId="5" lastPageNumber="446" pageId="4" pageNumber="445" type="key">
|
||
<key id="FFF3372FFFA6E11696C905538FC8F858" pageId="4" pageNumber="445">
|
||
<keyStep id="309678BBFFA6E11696C905538FCFF938" pageId="4" pageNumber="445">
|
||
<paragraph id="8BDD601EFFA6E11696C905538FC3F9D7" blockId="4.[151,1436,1606,2038]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C905538FC3F9D7" pageId="4" pageNumber="445">
|
||
1 Median keel of elytra relatively low, equal in height to lateral keel; both keels indistinct. Median lobe of aedeagus with elongate, parallel-sided, needle-like apical projection in ventral view. Total length 1.8–2.0 mm. ...
|
||
<taxonomicName id="4C621B9DFFA6E116923A05768FC3F9D7" ID-CoL="5CRYC" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" box="[1124,1431,1635,1657]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="subopaca">
|
||
<emphasis id="B916BC0CFFA6E116923A05768EA4F9D7" box="[1124,1264,1635,1657]" italics="true" pageId="4" pageNumber="445">Yola subopaca</emphasis>
|
||
Régimbart, 1895
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E11696C905948FCFF938" blockId="4.[151,1436,1606,2038]" box="[151,1435,1665,1686]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C905948FCFF938" box="[151,1435,1665,1686]" pageId="4" pageNumber="445">- Median elytral keel more strongly elevated than lateral keel. Both keels distinct.................................... 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309678BBFFA6E11696C9058B8FCFF940" pageId="4" pageNumber="445">
|
||
<paragraph id="8BDD601EFFA6E11696C9058B8FC8F97F" blockId="4.[151,1436,1606,2038]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C9058B8FC8F97F" pageId="4" pageNumber="445">
|
||
2 Median lobe of aedeagus in lateral view with apex distinctly curved in a ventral direction. All three elytral keels distinct; discal keel higher than median keel, lateral keel lower than median. Total length 1.8–2.0 mm.......
|
||
<taxonomicName id="4C621B9DFFA6E116922405AE8FC8F97F" ID-CoL="5CRXJ" authority="Regimbart, 1906" authorityName="Regimbart" authorityYear="1906" box="[1146,1436,1723,1745]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="dilatata">
|
||
<emphasis id="B916BC0CFFA6E116922405AE8EA1F97F" box="[1146,1269,1723,1745]" italics="true" pageId="4" pageNumber="445">Yola dilatata</emphasis>
|
||
Régimbart, 1906
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E11696C905CC8FCFF940" blockId="4.[151,1436,1606,2038]" box="[151,1435,1753,1774]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C905CC8FCFF940" box="[151,1435,1753,1774]" pageId="4" pageNumber="445">- Median lobe of aedeagus straight, or curved dorsally in lateral view.............................................. 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309678BBFFA6E11696C905E38FC8F858" pageId="4" pageNumber="445">
|
||
<paragraph id="8BDD601EFFA6E11696C905E38FC8F8CD" blockId="4.[151,1436,1606,2038]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C905E38FC8F8CD" pageId="4" pageNumber="445">
|
||
3 Median elytral keel only slightly more strongly elevated than lateral keel. Pronotum broadest at posterior corners, or just anterior to them. Pronotal disc flat between plicae. Median lobe of aedeagus in lateral view relatively broad almost until apex; apex at most slightly expanded and triangular in ventral view. Size smaller, total length
|
||
<quantity id="4C9ACDFBFFA6E11695AB04248E3FF8E8" box="[1013,1131,1841,1862]" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="4" pageNumber="445" unit="mm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9–2.3 mm</quantity>
|
||
; maximum width
|
||
<quantity id="4C9ACDFBFFA6E116934704248FDBF8E8" box="[1305,1423,1841,1862]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="4" pageNumber="445" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1–1.4 mm</quantity>
|
||
.............................................................................
|
||
<taxonomicName id="4C621B9DFFA6E116922A04588FC8F8CD" ID-CoL="5CRXQ" authority="Regimbart, 1906" authorityName="Regimbart" authorityYear="1906" box="[1140,1436,1869,1891]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis id="B916BC0CFFA6E116922A04588EA0F8CD" box="[1140,1268,1869,1891]" italics="true" pageId="4" pageNumber="445">Yola frontalis</emphasis>
|
||
Régimbart, 1906
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA6E11696C904798FC8F858" blockId="4.[151,1436,1606,2038]" pageId="4" pageNumber="445">
|
||
<keyLead id="3093C32BFFA6E11696C904798FC8F858" pageId="4" pageNumber="445">
|
||
- Median elytral keel distinctly more strongly elevated than lateral keel. Pronotum broadest just before middle. Posterior 1/3 of pronotal disc depressed between plicae, especially in centre; surface here raised abruptly to level of remainder of disc (
|
||
<figureCitation id="13597C9BFFA6E116932A049C8ABEF815" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="4" pageNumber="445">Fig. 1</figureCitation>
|
||
A). Median lobe of aedeagus (
|
||
<figureCitation id="13597C9BFFA6E116944B04B38818F815" box="[533,588,1958,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1963]" captionTargetBox="[160,1410,202,1892]" captionTargetId="figure@2.[151,1436,193,1919]" captionTargetPageId="2" captionText="FIGURE 1. Yola matsikammae sp. nov. A) holotype habitus, dorsal view; B) holotype habitus, lateral view; C) holotype median lobe of aedeagus, lateral and ventral views, right paramere. Scale bar A – B = 1 mm; C = 0.5 mm." httpUri="https://zenodo.org/record/245313/files/figure.png" pageId="4" pageNumber="445">Fig. 1</figureCitation>
|
||
C) in lateral view narrowing approx. 1/6 before apex, then parallel-sided to apex; apex strongly expanded, spathulate, in ventral view. Size larger, total length
|
||
<quantity id="4C9ACDFBFFA6E11695D704D18E5EF877" box="[905,1034,1988,2009]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.45" metricValueMin="2.3" pageId="4" pageNumber="445" unit="mm" value="2.375" valueMax="2.45" valueMin="2.3">2.3–2.45 mm</quantity>
|
||
; maximum width
|
||
<quantity id="4C9ACDFBFFA6E11692E604D18F63F877" box="[1208,1335,1988,2009]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" pageId="4" pageNumber="445" unit="mm" value="1.325" valueMax="1.4" valueMin="1.25">1.25–1.4 mm</quantity>
|
||
.......................................................................................
|
||
<taxonomicName id="4C621B9DFFA6E11692CB04F58F1DF858" ID-CoL="8LZRZ" box="[1173,1353,2016,2038]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA6E11692CB04F58F1DF858" box="[1173,1353,2016,2038]" italics="true" pageId="4" pageNumber="445">Yola matsikammae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA6E116931104F58FC8F858" bold="true" box="[1359,1436,2016,2038]" pageId="4" pageNumber="445">
|
||
<taxonomicNameLabel id="A2250177FFA6E116931104F58FC8F858" box="[1359,1436,2016,2038]" pageId="4" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<paragraph id="8BDD601EFFA7E117969903828F0EFE68" blockId="5.[151,1437,151,1182]" pageId="5" pageNumber="446">
|
||
Superficially, in both shape and dorsal colouration, the new taxon resembles species of
|
||
<taxonomicName id="4C621B9DFFA7E11792E5038D8F1EFF1E" ID-CoL="7GBL" box="[1211,1354,152,176]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E11792E5038D8F1EFF1E" box="[1211,1354,152,176]" italics="true" pageId="5" pageNumber="446">Sharphydrus</emphasis>
|
||
</taxonomicName>
|
||
Omer- Cooper, 1958, especially
|
||
<taxonomicName id="4C621B9DFFA7E11797EA03AE8874FF7D" ID-CoL="8M23Q" box="[436,544,187,211]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="B916BC0CFFA7E11797EA03AE8874FF7D" box="[436,544,187,211]" italics="true" pageId="5" pageNumber="446">S. brincki</emphasis>
|
||
</taxonomicName>
|
||
, with which it was microsympatric (i.e. co-occurred in the same microhabitat). It can readily be distinguished from
|
||
<taxonomicName id="4C621B9DFFA7E117946E03CB88C8FF5B" ID-CoL="8M23Q" box="[560,668,221,245]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="B916BC0CFFA7E117946E03CB88C8FF5B" box="[560,668,221,245]" italics="true" pageId="5" pageNumber="446">S. brincki</emphasis>
|
||
</taxonomicName>
|
||
by the presence of three elytral keels rather than just the discal one. The discal keel of
|
||
<taxonomicName id="4C621B9DFFA7E117973102148872FEB6" ID-CoL="8LZRZ" box="[367,550,256,280]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA7E117973102148872FEB6" box="[367,550,256,280]" italics="true" pageId="5" pageNumber="446">
|
||
<collectingCountry id="F375208EFFA7E117973102148BD4FEB6" box="[367,384,257,280]" name="Italy" pageId="5" pageNumber="446">Y</collectingCountry>
|
||
. matsikammae
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA7E117947103EA88DFFEB6" bold="true" box="[559,651,255,280]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2250177FFA7E117947103EA88DFFEB6" box="[559,651,255,280]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also much stronger, the dorsal surface shinier, and the dorsal and ventral punctures larger than in
|
||
<taxonomicName id="4C621B9DFFA7E117945D02368824FE95" ID-CoL="8M23Q" box="[515,624,291,315]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="B916BC0CFFA7E117945D02368824FE95" box="[515,624,291,315]" italics="true" pageId="5" pageNumber="446">S. brincki</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the median lobe of the aedeagus lacks the tripartite apical structure characteristic of known
|
||
<taxonomicName id="4C621B9DFFA7E1179440025088F9FEF3" ID-CoL="7GBL" box="[542,685,325,349]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E1179440025088F9FEF3" box="[542,685,325,349]" italics="true" pageId="5" pageNumber="446">Sharphydrus</emphasis>
|
||
</taxonomicName>
|
||
species (see
|
||
<bibRefCitation id="EFF31DEFFFA7E11795120250898EFEF0" author="Bilton" box="[844,986,325,350]" pageId="5" pageNumber="446" refString="Bilton, D. T. (2013) A taxonomic revision of South African Sharphydrus with the description of two new species (Coleoptera: Dytiscidae: Bidessini). Zootaxa, 3750 (1), 26 - 36. http: // dx. doi. org / 10.11646 / 10.11646 / zootaxa. 3750.1.2" type="journal article" year="2013">Bilton 2013</bibRefCitation>
|
||
).
|
||
<collectingCountry id="F375208EFFA7E11795AF02538E44FEF0" box="[1009,1040,326,350]" name="American Samoa" pageId="5" pageNumber="446">As</collectingCountry>
|
||
discussed by
|
||
<bibRefCitation id="EFF31DEFFFA7E11792E602508F3CFEF0" author="Bistrom" box="[1208,1384,325,350]" pageId="5" pageNumber="446" refString="Bistrom, O. (1988) Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta Zoologica Fennica, 184, 1 - 41." type="journal article" year="1988">Biström (1988)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFF31DEFFFA7E11796C902728B66FE2E" author="Bilton" box="[151,306,359,384]" pageId="5" pageNumber="446" refString="Bilton, D. T. (2013) A taxonomic revision of South African Sharphydrus with the description of two new species (Coleoptera: Dytiscidae: Bidessini). Zootaxa, 3750 (1), 26 - 36. http: // dx. doi. org / 10.11646 / 10.11646 / zootaxa. 3750.1.2" type="journal article" year="2013">Bilton (2013)</bibRefCitation>
|
||
, the distinction between
|
||
<taxonomicName id="4C621B9DFFA7E1179412027D882EFE2E" ID-CoL="89X8" box="[588,634,360,384]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E1179412027D882EFE2E" box="[588,634,360,384]" italics="true" pageId="5" pageNumber="446">Yola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621B9DFFA7E11794E8027D8911FE2E" ID-CoL="7GBL" box="[694,837,360,384]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E11794E8027D8911FE2E" box="[694,837,360,384]" italics="true" pageId="5" pageNumber="446">Sharphydrus</emphasis>
|
||
</taxonomicName>
|
||
is not well-defined.
|
||
<bibRefCitation id="EFF31DEFFFA7E117926E02728E9DFE2E" author="Bilton" box="[1072,1225,359,384]" pageId="5" pageNumber="446" refString="Bilton, D. T. (2013) A taxonomic revision of South African Sharphydrus with the description of two new species (Coleoptera: Dytiscidae: Bidessini). Zootaxa, 3750 (1), 26 - 36. http: // dx. doi. org / 10.11646 / 10.11646 / zootaxa. 3750.1.2" type="journal article" year="2013">Bilton (2013)</bibRefCitation>
|
||
suggested that the tripartite structure of the median lobe of the aedeagus represented a unique synapomorphy of this group, although the inter-relationships between
|
||
<taxonomicName id="4C621B9DFFA7E11797A802B888D1FE6B" ID-CoL="7GBL" box="[502,645,429,453]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E11797A802B888D1FE6B" box="[502,645,429,453]" italics="true" pageId="5" pageNumber="446">Sharphydrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621B9DFFA7E11794E202B888BEFE6B" ID-CoL="89X8" box="[700,746,429,453]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E11794E202B888BEFE6B" box="[700,746,429,453]" italics="true" pageId="5" pageNumber="446">Yola</emphasis>
|
||
</taxonomicName>
|
||
, including the new species, require further exploration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3783395FFA7E117969902DA8966FD16" pageId="5" pageNumber="446" type="distribution">
|
||
<paragraph id="8BDD601EFFA7E117969902DA8966FD16" blockId="5.[151,1437,151,1182]" pageId="5" pageNumber="446">
|
||
<emphasis id="B916BC0CFFA7E117969902DA8B0DFE46" bold="true" box="[199,345,463,488]" pageId="5" pageNumber="446">Distribution</emphasis>
|
||
. To date known from the
|
||
<typeStatus id="54D9DEBCFFA7E11794D502C588EFFE46" box="[651,699,464,488]" pageId="5" pageNumber="446">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13597C9BFFA7E117957502DA8927FE46" box="[811,883,463,488]" captionStart="FIGURE 2" captionStartId="3.[151,250,1821,1843]" captionTargetBox="[151,1436,384,1799]" captionTargetId="figure@3.[151,1436,193,1799]" captionTargetPageId="3" captionText="FIGURE 2. Type locality of Yola matsikammae sp. nov., South Africa, Western Cape, Matsikammaberg, permanent stream ca. 1 km SE of Sewefontein Farm. Photo D. T. Bilton." httpUri="https://zenodo.org/record/245314/files/figure.png" pageId="5" pageNumber="446">Fig. 2</figureCitation>
|
||
), a permanent stream surrounded by Bokkeveld Sandstone Fynbos (sensu
|
||
<bibRefCitation id="EFF31DEFFFA7E11797E402E788A5FDA5" author="Mucina" box="[442,753,498,523]" pageId="5" pageNumber="446" refString="Murcina, L. & Rutherford, M. C. (2006) (Eds.), The vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) on the Matsikammaberg in the northern part of the Western Cape province of
|
||
<collectingCountry id="F375208EFFA7E117970001008BA5FD80" box="[350,497,533,558]" name="South Africa" pageId="5" pageNumber="446">South Africa</collectingCountry>
|
||
. The Matsikammaberg is a striking inselberg, reaching just over
|
||
<quantity id="4C9ACDFBFFA7E117929C01038F4AFD83" box="[1218,1310,534,558]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="446" unit="m" value="1000.0">1,000 m</quantity>
|
||
in altitude, with
|
||
<quantity id="4C9ACDFBFFA7E117968E012D8B43FDE1" box="[208,279,568,592]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="446" unit="m" value="700.0">700 m</quantity>
|
||
high sandstone cliffs towering over the dry Knersvlakte plains of Namaqualand. The mountain forms a mesic island in an otherwise semi-arid landscape, annual rainfall reaching
|
||
<quantity id="4C9ACDFBFFA7E11795BB014F8E17FDDC" box="[997,1091,602,627]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="446" unit="mm" value="550.0">550 mm</quantity>
|
||
in the east contrasting with as little as
|
||
<quantity id="4C9ACDFBFFA7E11796B101688B6AFD3B" box="[239,318,637,662]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="446" unit="mm" value="50.0">50 mm</quantity>
|
||
per year on the plains below. The Matsikammaberg is consequently home to a diverse flora, 10% of which is regionally, and 4% locally endemic (
|
||
<bibRefCitation id="EFF31DEFFFA7E11794C8018A8972FD16" author="Helme" box="[662,806,671,696]" pageId="5" pageNumber="446" refString="Helme, N. (2004) The vegetation of Matsikamma Mountain, with some comments on endemicity. PlantLife, 30, 5 - 16." type="journal article" year="2004">Helme 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3783395FFA7E117969901D78FD7FB33" pageId="5" pageNumber="446" type="etymology">
|
||
<paragraph id="8BDD601EFFA7E117969901D789C2FCB1" blockId="5.[151,1437,151,1182]" pageId="5" pageNumber="446">
|
||
<emphasis id="B916BC0CFFA7E117969901D78B12FD75" bold="true" box="[199,326,706,731]" pageId="5" pageNumber="446">Etymology</emphasis>
|
||
. Named after the Matsikammaberg, on which the
|
||
<typeStatus id="54D9DEBCFFA7E117952101D689FBFD75" box="[895,943,707,731]" pageId="5" pageNumber="446">type</typeStatus>
|
||
locality is situated. Matsikamma is a Khoi- San word which translates as “full of pools”, in apparent reference to the abundance of seasonal rock pools on the mountain plateau. The specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BDD601EFFA7E1179699003F8FD7FB33" blockId="5.[151,1437,151,1182]" pageId="5" pageNumber="446">
|
||
<emphasis id="B916BC0CFFA7E1179699003F8B70FCED" bold="true" box="[199,292,810,835]" pageId="5" pageNumber="446">Ecology</emphasis>
|
||
. Specimens of
|
||
<taxonomicName id="4C621B9DFFA7E1179788003E88D9FCED" box="[470,653,811,835]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="matsikammae" status="sp. nov.">
|
||
<emphasis id="B916BC0CFFA7E1179788003E88D9FCED" box="[470,653,811,835]" italics="true" pageId="5" pageNumber="446">
|
||
<collectingCountry id="F375208EFFA7E1179788003E8BB3FCEC" box="[470,487,811,834]" name="Italy" pageId="5" pageNumber="446">Y</collectingCountry>
|
||
. matsikammae
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BC0CFFA7E11794CB003F88A5FCED" bold="true" box="[661,753,810,835]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2250177FFA7E11794CB003F88A5FCED" box="[661,753,810,835]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were taken from the margins of deep (ca.
|
||
<quantity id="4C9ACDFBFFA7E1179285003F8F04FCED" box="[1243,1360,810,835]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="446" unit="cm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 cm</quantity>
|
||
), clear pools in a permanent stream section, over sand and gravel. Here they co-occurred with a range of water beetles, including
|
||
<taxonomicName id="4C621B9DFFA7E117975200658833FC26" authority="Aube, 1838" authorityName="Aube" authorityYear="1838" box="[268,615,879,904]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="dregei">
|
||
<emphasis id="B916BC0CFFA7E117975200658B84FC26" box="[268,464,880,904]" italics="true" pageId="5" pageNumber="446">Hydaticus dregei</emphasis>
|
||
Aubé, 1838
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117942900658935FC26" box="[631,865,880,904]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="B916BC0CFFA7E117942900658935FC26" box="[631,865,880,904]" italics="true" pageId="5" pageNumber="446">Sharphydrus brincki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117952A00658F26FC26" authority="Omer-Cooper, 1965" authorityName="Omer-Cooper" authorityYear="1965" box="[884,1394,879,904]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="consuetus">
|
||
<emphasis id="B916BC0CFFA7E117952A00658E2AFC26" box="[884,1150,880,904]" italics="true" pageId="5" pageNumber="446">Canthyporus consuetus</emphasis>
|
||
Omer-Cooper, 1965
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11793DC00658B9FFC05" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="lateralis">
|
||
<emphasis id="B916BC0CFFA7E11793DC00658AA1FC05" italics="true" pageId="5" pageNumber="446">C. lateralis</emphasis>
|
||
(Boheman, 1848)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117978200868931FC05" authority="Omer-Cooper, 1965" authorityName="Omer-Cooper" authorityYear="1965" box="[476,869,914,939]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="nebulosus">
|
||
<emphasis id="B916BC0CFFA7E117978200868825FC05" box="[476,625,915,939]" italics="true" pageId="5" pageNumber="446">C. nebulosus</emphasis>
|
||
Omer-Cooper, 1965
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117952B00868F3AFC05" authority="Omer-Cooper, 1965" authorityName="Omer-Cooper" authorityYear="1965" box="[885,1390,914,939]" class="Insecta" family="Dytiscidae" genus="Hydropeplus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B916BC0CFFA7E117952B00868E29FC04" box="[885,1149,915,939]" italics="true" pageId="5" pageNumber="446">Hydropeplus montanus</emphasis>
|
||
Omer-Cooper, 1965
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11793DF00868BB1FC60" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Dytiscidae" genus="Hydropeplus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="trimaculatus">
|
||
<emphasis id="B916BC0CFFA7E11793DF00868B71FC63" italics="true" pageId="5" pageNumber="446">H. trimaculatus</emphasis>
|
||
(Laporte, 1835)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621B9DFFA7E117947B00A088FAFC63" box="[549,686,949,973]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E117947B00A088FAFC63" box="[549,686,949,973]" italics="true" pageId="5" pageNumber="446">Laccophilus</emphasis>
|
||
</taxonomicName>
|
||
sp. Other beetles taken in the same stream reach, but in other microhabitats, were
|
||
<taxonomicName id="4C621B9DFFA7E11797D900CD8941FC5E" authority="Reichart, 1976" authorityName="Reichart" authorityYear="1976" box="[391,789,983,1008]" class="Insecta" family="Torridincolidae" genus="Delevea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="bertrandi">
|
||
<emphasis id="B916BC0CFFA7E11797D900CD880EFC5E" box="[391,602,984,1008]" italics="true" pageId="5" pageNumber="446">Delevea bertrandi</emphasis>
|
||
Reichart, 1976
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117957800CD8EAAFC5E" authority="Thunberg, 1781" authorityName="Thunberg" authorityYear="1781" box="[806,1278,983,1008]" class="Insecta" family="Gyrinidae" genus="Aulonogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="capensis">
|
||
<emphasis id="B916BC0CFFA7E117957800CD8E77FC41" box="[806,1059,984,1008]" italics="true" pageId="5" pageNumber="446">Aulonogyrus capensis</emphasis>
|
||
(Thunberg, 1781)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117934E00CD8B1EFBBD" authority="Modeer, 1776" authorityName="Modeer" authorityYear="1776" class="Insecta" family="Gyrinidae" genus="Aulonogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="formosus">
|
||
<emphasis id="B916BC0CFFA7E117934E00CD8FC8FC5E" box="[1296,1436,984,1008]" italics="true" pageId="5" pageNumber="446">A. formosus</emphasis>
|
||
(Modeer, 1776)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117970700EE88C8FBBD" authority="Aube, 1838" authorityName="Aube" authorityYear="1838" box="[345,668,1018,1043]" class="Insecta" family="Gyrinidae" genus="Aulonogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="marginatus">
|
||
<emphasis id="B916BC0CFFA7E117970700EE8BACFBBC" box="[345,504,1019,1042]" italics="true" pageId="5" pageNumber="446">A. marginatus</emphasis>
|
||
(Aubé, 1838)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11794F500EE8E4CFBBD" authority="Aube, 1838" authorityName="Aube" authorityYear="1838" box="[683,1048,1018,1043]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="B916BC0CFFA7E11794F500EE89D3FBBC" box="[683,903,1019,1042]" italics="true" pageId="5" pageNumber="446">Dineutus punctatus</emphasis>
|
||
Aubé, 1838
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117927800EE8FCCFBBD" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[1062,1432,1018,1043]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="capensis">
|
||
<emphasis id="B916BC0CFFA7E117927800EE8F55FBBC" box="[1062,1281,1019,1043]" italics="true" pageId="5" pageNumber="446">Copelatus capensis</emphasis>
|
||
Sharp, 1882
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11796C9070B8827FB98" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[151,627,1053,1078]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="glabriventris">
|
||
<emphasis id="B916BC0CFFA7E11796C9070B8BFCFB9B" box="[151,424,1053,1077]" italics="true" pageId="5" pageNumber="446">Anacaena glabriventris</emphasis>
|
||
Komarek, 2004
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11794D9070B899BFB98" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[647,975,1053,1078]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="reducta">
|
||
<emphasis id="B916BC0CFFA7E11794D9070B8950FB9B" box="[647,772,1053,1077]" italics="true" pageId="5" pageNumber="446">A. reducta</emphasis>
|
||
Komarek, 2004
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11795BA070B8FCCFB98" authority="Knisch, 1924" authorityName="Knisch" authorityYear="1924" box="[996,1432,1053,1078]" class="Insecta" family="Hydrophilidae" genus="Crenitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="zimmermanni">
|
||
<emphasis id="B916BC0CFFA7E11795BA070B8EB1FB9B" box="[996,1253,1054,1077]" italics="true" pageId="5" pageNumber="446">Crenitis zimmermanni</emphasis>
|
||
Knisch, 1924
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11796C907558B43FBF6" box="[151,279,1088,1112]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E11796C907558B43FBF6" box="[151,279,1088,1112]" italics="true" pageId="5" pageNumber="446">Helochares</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<emphasis id="B916BC0CFFA7E117970707558821FBF6" box="[345,629,1088,1112]" italics="true" pageId="5" pageNumber="446">Mesoceration umbrosum</emphasis>
|
||
Perkins, 2008,
|
||
<emphasis id="B916BC0CFFA7E117956707558987FBF9" box="[825,979,1088,1111]" italics="true" pageId="5" pageNumber="446">Mesoceration</emphasis>
|
||
<emphasis id="B916BC0CFFA7E11795BF072A8E15FBF6" bold="true" box="[993,1089,1087,1112]" pageId="5" pageNumber="446">
|
||
sp.
|
||
<date id="FFDC46DEFFA7E117924E072A8E15FBF6" box="[1040,1089,1087,1112]" pageId="5" pageNumber="446">nov.</date>
|
||
</emphasis>
|
||
1,
|
||
<emphasis id="B916BC0CFFA7E117922F07558F5FFBF9" box="[1137,1291,1088,1111]" italics="true" pageId="5" pageNumber="446">Mesoceration</emphasis>
|
||
<emphasis id="B916BC0CFFA7E1179347072A8F2DFBF6" bold="true" box="[1305,1401,1087,1112]" pageId="5" pageNumber="446">
|
||
sp.
|
||
<date id="FFDC46DEFFA7E1179316072A8F2DFBF6" box="[1352,1401,1087,1112]" pageId="5" pageNumber="446">nov.</date>
|
||
</emphasis>
|
||
2,
|
||
<taxonomicName id="4C621B9DFFA7E11796C907768806FBD5" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[151,594,1122,1147]" class="Insecta" family="Hydraenidae" genus="Prosthetops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="wolfbergensis">
|
||
<emphasis id="B916BC0CFFA7E11796C907768BE9FBD5" box="[151,445,1123,1147]" italics="true" pageId="5" pageNumber="446">Prosthetops wolfbergensis</emphasis>
|
||
<bibRefCitation id="EFF31DEFFFA7E117979A07778806FBD5" author="Bilton" box="[452,594,1122,1147]" pageId="5" pageNumber="446" refString="Bilton, D. T. (2013) A taxonomic revision of South African Sharphydrus with the description of two new species (Coleoptera: Dytiscidae: Bidessini). Zootaxa, 3750 (1), 26 - 36. http: // dx. doi. org / 10.11646 / 10.11646 / zootaxa. 3750.1.2" type="journal article" year="2013">Bilton, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E1179403077689AAFBD5" authority="Perkins, 2004" authorityName="Perkins" authorityYear="2004" box="[605,1022,1122,1147]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="endroedyi">
|
||
<emphasis id="B916BC0CFFA7E11794030776890FFBD5" box="[605,859,1123,1147]" italics="true" pageId="5" pageNumber="446">Pneuminion endroedyi</emphasis>
|
||
Perkins, 2004
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117925707768FCCFBD5" authority="Perkins, 1997" authorityName="Perkins" authorityYear="1997" box="[1033,1432,1122,1147]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="velamen">
|
||
<emphasis id="B916BC0CFFA7E117925707768EA0FBD5" box="[1033,1268,1123,1147]" italics="true" pageId="5" pageNumber="446">Pneuminion velamen</emphasis>
|
||
Perkins, 1997
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E11796C90793897CFB30" authority="Perkins & Balfour-Browne, 1994" authorityName="Perkins & Balfour-Browne" authorityYear="1994" box="[151,808,1157,1182]" class="Insecta" family="Hydraenidae" genus="Discozantaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="genuvela">
|
||
<emphasis id="B916BC0CFFA7E11796C907938BFDFB33" box="[151,425,1157,1181]" italics="true" pageId="5" pageNumber="446">Discozantaena genuvela</emphasis>
|
||
Perkins & Balfour-Browne, 1994
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C621B9DFFA7E117956D07908EB5FB30" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[819,1249,1157,1182]" class="Insecta" family="Elmidae" genus="Elpidelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="capensis">
|
||
<emphasis id="B916BC0CFFA7E117956D07908E47FB33" box="[819,1043,1157,1181]" italics="true" pageId="5" pageNumber="446">Elpidelmis capensis</emphasis>
|
||
(Grouvelle, 1890)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621B9DFFA7E117934607938F08FB33" box="[1304,1372,1158,1181]" class="Insecta" family="Dryopidae" genus="Strina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BC0CFFA7E117934607938F08FB33" box="[1304,1372,1158,1181]" italics="true" pageId="5" pageNumber="446">Strina</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |