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<document id="57558A02E0F0D0A4CBE014DBCC5CD28C" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFCD1F5BFF50FB37FF1C8F9E" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax gracilis Burret 1929" docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="51" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="98EECB0B7578140FD3F268D43EED77AF">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:title id="DE6355B1665125DE608870494774E5F9">Phytotaxa</mods:title>
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<mods:classification id="EC200EEF36EA34521F7D34BDF943F5E2">journal article</mods:classification>
<mods:identifier id="AA06A93938C92C249F24C95B96DA8FF4" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
<mods:identifier id="9D97038B3298B401A8AFC5528E022F43" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03B387DAFFCD1F5BFF50FB37FF1C8F9E" ID-DOI="http://doi.org/10.5281/zenodo.8389305" ID-Zenodo-Dep="8389305" LSID="urn:lsid:plazi:treatment:03B387DAFFCD1F5BFF50FB37FF1C8F9E" httpUri="http://treatment.plazi.org/id/03B387DAFFCD1F5BFF50FB37FF1C8F9E" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
<subSubSection id="C3006547FFCD1F5DFF50FB37FE378CEB" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8BA536CCFFCD1F5DFF50FB37FE378CEB" blockId="50.[136,1452,1166,1265]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF50FB37FD658CB3" bold="true" box="[136,668,1166,1193]" pageId="50" pageNumber="51">
1.17.
<taxonomicName id="4C1A4D4FFFCD1F5DFF1FFB37FD6E8CB3" ID-CoL="WLYH" authority="Burret (1929: 14)" authorityName="Burret" authorityPageNumber="14" authorityYear="1929" box="[199,663,1166,1193]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B96EEADEFFCD1F5DFF1FFB37FE388CB2" bold="true" box="[199,449,1166,1193]" italics="true" pageId="50" pageNumber="51">Coccothrinax gracilis</emphasis>
<bibRefCitation id="EF8B4B3DFFCD1F5DFE1FFB37FD6E8CB3" author="Burret, M." box="[455,663,1166,1192]" pageId="50" pageNumber="51" pagination="1 - 42" refId="ref61294" refString="Burret, M. (1929) Palmae Cubenses et Domingenses a Cl. E. L. Ekman 1914 - 1928 lectae. Kongliga Svenska Vetenskapsakademiens Handlingar 6 (7): 1 - 42." type="journal article" year="1929">Burret (1929: 14)</bibRefCitation>
</taxonomicName>
.
</emphasis>
<materialsCitation id="3B723C91FFCD1F5DFD7BFB36FE378CEB" collectingDate="1925-03-26" collectionCode="E, S, EHH, B" collectorName="de Paix" country="Haiti" location="Vallee des Trois Rivieres" municipality="Port de Paix" pageId="50" pageNumber="51" specimenCount="3" typeStatus="lectotype">
<typeStatus id="54A1886EFFCD1F5DFD7BFB36FCEA8CB2" box="[675,787,1167,1193]" pageId="50" pageNumber="51">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F30D765CFFCD1F5DFC26FB37FBB78CB3" box="[1022,1102,1166,1192]" name="Haiti" pageId="50" pageNumber="51">HAITI</collectingCountry>
.
<location id="8EC56017FFCD1F5DFB80FB37FA8B8CB3" LSID="urn:lsid:plazi:treatment:03B387DAFFCD1F5BFF50FB37FF1C8F9E:8EC56017FFCD1F5DFB80FB37FA8B8CB3" box="[1112,1394,1166,1192]" country="Haiti" municipality="Port de Paix" name="Vallee des Trois Rivieres" pageId="50" pageNumber="51">Vallée des Trois Rivières</location>
,
<collectingMunicipality id="6BC1ACB6FFCD1F5DFAA5FB36FF198CD7" pageId="50" pageNumber="51">
Port
<collectorName id="26EF531AFFCD1F5DFF50FB0BFF198CD7" box="[136,224,1202,1228]" pageId="50" pageNumber="51">de Paix</collectorName>
</collectingMunicipality>
,
<date id="FFA4100CFFCD1F5DFF34FB0BFE658CD6" box="[236,412,1202,1229]" pageId="50" pageNumber="51" value="1925-03-26">
<collectingDate id="EFE0E9E4FFCD1F5DFF34FB0BFE658CD6" box="[236,412,1202,1229]" pageId="50" pageNumber="51" value="1925-03-26">26 March 1925</collectingDate>
</date>
,
<emphasis id="B96EEADEFFCD1F5DFE71FB0AFDAE8CD7" box="[425,599,1202,1228]" italics="true" pageId="50" pageNumber="51">
<collectionCode id="ED0BAE09FFCD1F5DFE71FB0AFE478CD7" box="[425,446,1203,1228]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="50" pageNumber="51" type="Herbarium">E</collectionCode>
. Ekman 3586
</emphasis>
(
<typeStatus id="54A1886EFFCD1F5DFDBFFB0BFD298CD7" box="[615,720,1202,1228]" pageId="50" pageNumber="51">lectotype</typeStatus>
<collectionCode id="ED0BAE09FFCD1F5DFD00FB0BFD118CD7" box="[728,744,1202,1228]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="50" pageNumber="51" type="Herbarium">S</collectionCode>
<emphasis id="B96EEADEFFCD1F5DFD37FB0AFCEF8CD7" box="[751,790,1203,1228]" italics="true" pageId="50" pageNumber="51">n.v.</emphasis>
,
<collectionCode id="ED0BAE09FFCD1F5DFCFDFB0BFCCC8CD7" box="[805,821,1202,1228]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="50" pageNumber="51" type="Herbarium">S</collectionCode>
image!, isolectotypes
<collectionCode id="ED0BAE09FFCD1F5DFBECFB0AFB978CD6" box="[1076,1134,1203,1229]" country="Haiti" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15290" lsid="urn:lsid:biocol.org:col:15290" name="Université d'Etat d'Haiti" pageId="50" pageNumber="51" type="Herbarium">EHH</collectionCode>
<emphasis id="B96EEADEFFCD1F5DFBAEFB0AFB618CD7" box="[1142,1176,1203,1228]" italics="true" pageId="50" pageNumber="51">n.v</emphasis>
.,
<collectionCode id="ED0BAE09FFCD1F5DFB76FB0AFB118CD6" box="[1198,1256,1203,1229]" country="Haiti" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15290" lsid="urn:lsid:biocol.org:col:15290" name="Université d'Etat d'Haiti" pageId="50" pageNumber="51" type="Herbarium">EHH</collectionCode>
image!, US!, the
<typeStatus id="54A1886EFFCD1F5DFF50FB6FFF138CEB" box="[136,234,1238,1264]" pageId="50" pageNumber="51">holotype</typeStatus>
at
<collectionCode id="ED0BAE09FFCD1F5DFED4FB6EFEE68CEA" box="[268,287,1239,1265]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="50" pageNumber="51" type="Herbarium">B</collectionCode>
was destroyed)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFCD1F5DFF50FAA7FDE58F63" pageId="50" pageNumber="51" type="description">
<paragraph id="8BA536CCFFCD1F5DFF50FAA7FDE58F63" blockId="50.[136,1452,1310,1984]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF50FAA7FF298D23" bold="true" box="[136,208,1310,1336]" pageId="50" pageNumber="51">Stems</emphasis>
length not recorded,
<quantity id="4CE29B29FFCD1F5DFE67FAA7FDF28D22" box="[447,523,1310,1337]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="50" pageNumber="51" unit="cm" value="7.6">7.6 cm</quantity>
diameter, branching not recorded.
<emphasis id="B96EEADEFFCD1F5DFC4BFAA7FC1C8D23" bold="true" box="[915,997,1310,1336]" pageId="50" pageNumber="51">Leaves</emphasis>
not recorded; leaf sheath fibers 1.1(1.0 1.3) mm diameter, stout, loosely woven and forming a loose, hexagonal mesh, initially forming ligules at the apices; petioles 5.4(4.36.5) mm diameter just below the apex; palmans 4.6(2.75.5) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 25(2426) per leaf, the middle ones 33.8(27.042.5) cm long and 2.1(2.02.3) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively long and narrow, tapering from base to apex, scarcely folded, flexible and not leathery, a shoulder or constriction absent or poorly developed, the apices thin, deeply splitting and breaking off; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, semi-persistent, interlocking, fimbriate hairs without an obvious center, without or with poorly developed transverse veinlets.
<emphasis id="B96EEADEFFCD1F5DFF28F93FFE608EBB" bold="true" box="[240,409,1670,1696]" pageId="50" pageNumber="51">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences number not recorded; proximalmost rachillae straight, 4.7(3.85.5) cm long and 1.0(0.61.3) mm diameter in fruit; rachillae uneven at or near anthesis with lines of warty outgrowths, these often becoming more pronounced as fruits develop; stamens not recorded; fruit pedicels
<quantity id="4CE29B29FFCD1F5DFDF1F8AFFD4B8F2A" box="[553,690,1814,1841]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="50" pageNumber="51" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
long;
<emphasis id="B96EEADEFFCD1F5DFD26F8AFFCC68F2B" bold="true" box="[766,831,1814,1840]" pageId="50" pageNumber="51">fruits</emphasis>
6.2(5.86.6) mm long and 7.2(7.17.2) mm diameter, wine color; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFCD1F5BFF65F83BFF1C8F9E" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BA536CCFFCD1F5DFF65F83BFF0F8FDB" blockId="50.[136,1452,1310,1984]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF65F83BFDFC8F87" bold="true" box="[189,517,1922,1948]" pageId="50" pageNumber="51">Distribution and habitat:—</emphasis>
Northwestern
<collectingCountry id="F30D765CFFCD1F5DFD7CF83BFD258F87" box="[676,732,1922,1948]" name="Haiti" pageId="50" pageNumber="51">Haiti</collectingCountry>
(
<collectingRegion id="49DEF82EFFCD1F5DFD32F83BFC8B8F87" box="[746,882,1922,1948]" country="Haiti" name="Nord-Ouest" pageId="50" pageNumber="51">Nord-Ouest</collectingRegion>
) (
<figureCitation id="13212A49FFCD1F5DFC51F83BFC208F87" box="[905,985,1922,1948]" captionStart="FIGURE 13" captionStartId="52.[136,229,1506,1528]" captionTargetBox="[155,1429,198,1477]" captionTargetId="figure-27@52.[150,1434,193,1482]" captionTargetPageId="52" captionText="FIGURE 13. Distribution of Coccothrinax gracilis, C. gundlachii, C. hioramii, and C. jimenezii" figureDoi="http://doi.org/10.5281/zenodo.8389385" httpUri="https://zenodo.org/record/8389385/files/figure.png" pageId="50" pageNumber="51">Fig. 13</figureCitation>
) on limestone terraces near the sea at low elevation.
</paragraph>
<caption id="DF656644FFCC1F5CFF50F8C4FDBF8FAC" ID-DOI="http://doi.org/10.5281/zenodo.8389383" ID-Zenodo-Dep="8389383" httpUri="https://zenodo.org/record/8389383/files/figure.png" pageId="51" pageNumber="52" startId="51.[136,214,1917,1938]" targetBox="[151,1433,211,1891]" targetPageId="51" targetType="plate">
<paragraph id="8BA536CCFFCC1F5CFF50F8C4FDBF8FAC" blockId="51.[136,1452,1917,1975]" pageId="51" pageNumber="52">
<emphasis id="B96EEADEFFCC1F5CFF50F8C4FF0C8F89" bold="true" box="[136,245,1917,1938]" pageId="51" pageNumber="52">PLATE 13</emphasis>
.
<taxonomicName id="4C1A4D4FFFCC1F5CFED9F8C4FE028F88" box="[257,507,1917,1939]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="species" species="gonaivensis">
<emphasis id="B96EEADEFFCC1F5CFED9F8C4FE028F88" box="[257,507,1917,1939]" italics="true" pageId="51" pageNumber="52">Coccothrinax gonaivensis</emphasis>
</taxonomicName>
, Haiti, with wedge-shaped leaf blades and segments not pendulous at the apices, giving the leaf a flat appearance. (Image by Eladio Fernández).
</paragraph>
</caption>
<caption id="DF656644FFCB1F5BFF50FA5BFBDF8DE3" ID-DOI="http://doi.org/10.5281/zenodo.8389385" ID-Zenodo-Dep="8389385" box="[136,1062,1506,1528]" httpUri="https://zenodo.org/record/8389385/files/figure.png" pageId="52" pageNumber="53" startId="52.[136,229,1506,1528]" targetBox="[155,1429,198,1477]" targetPageId="52" targetType="figure">
<paragraph id="8BA536CCFFCB1F5BFF50FA5BFBDF8DE3" blockId="52.[136,1062,1506,1528]" box="[136,1062,1506,1528]" pageId="52" pageNumber="53">
<emphasis id="B96EEADEFFCB1F5BFF50FA5BFEFA8DEC" bold="true" box="[136,259,1506,1528]" pageId="52" pageNumber="53">FIGURE 13</emphasis>
. Distribution of
<emphasis id="B96EEADEFFCB1F5BFE7BFA5BFC7C8DEC" box="[419,901,1506,1527]" italics="true" pageId="52" pageNumber="53">
<taxonomicName id="4C1A4D4FFFCB1F5BFE7BFA5BFD818DEC" authorityName="Burret" authorityYear="1929" box="[419,632,1506,1527]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="gracilis">Coccothrinax gracilis</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFCB1F5BFD59FA5BFCFF8DEC" box="[641,774,1506,1527]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="gundlachii">C. gundlachii</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFCB1F5BFCC8FA5BFC788DEC" box="[784,897,1506,1527]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="hioramii">C. hioramii</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C1A4D4FFFCB1F5BFC6BFA5BFBDF8DE3" authorityName="Mejia &amp; Garcia" authorityYear="2013" box="[947,1062,1506,1528]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="jimenezii">C. jimenezii</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BA536CCFFCB1F5BFF65F99FFF1F8ECA" blockId="52.[136,1452,1574,1925]" pageId="52" pageNumber="53">
<emphasis id="B96EEADEFFCB1F5BFF65F99FFE538E5A" bold="true" box="[189,426,1574,1601]" pageId="52" pageNumber="53">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFCB1F5BFD10F99FFC458E5B" authorityName="Burret" authorityYear="1929" box="[712,956,1574,1600]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B96EEADEFFCB1F5BFD10F99FFC458E5B" box="[712,956,1574,1600]" italics="true" pageId="52" pageNumber="53">Coccothrinax gracilis</emphasis>
</taxonomicName>
has a unique combination of character states and is recognized as a phylogenetic species. It is a member of a group of nine Hispaniolan species, differing in its leaf shape, leaf segment shape and indumentum, and seed surfaces (see notes under
<taxonomicName id="4C1A4D4FFFCB1F5BFBCDF9D6FB528E93" authorityName="Mejia &amp; Garcia" authorityYear="1997" box="[1045,1195,1646,1673]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="boschiana">
<emphasis id="B96EEADEFFCB1F5BFBCDF9D6FB528E93" box="[1045,1195,1646,1673]" italics="true" pageId="52" pageNumber="53">C. boschiana</emphasis>
</taxonomicName>
). For the group, it has unusually large fruits. It is known only from the
<typeStatus id="54A1886EFFCB1F5BFD77F92AFD268EB6" box="[687,735,1683,1709]" pageId="52" pageNumber="53">type</typeStatus>
locality and does not now occur there (Eladio Fernández, pers. comm.).
</paragraph>
<paragraph id="8BA536CCFFCB1F5BFF65F962FF1C8F9E" blockId="52.[136,1452,1574,1925]" pageId="52" pageNumber="53">
A few unidentified specimens may also belong here. Two specimens (
<emphasis id="B96EEADEFFCB1F5BFC19F962FB568EEE" box="[961,1199,1755,1781]" italics="true" pageId="52" pageNumber="53">Zanoni 28372, 33498</emphasis>
), both from Presquile du St. Nicolas from northwestern
<collectingCountry id="F30D765CFFCB1F5BFDDEF947FDB88F03" box="[518,577,1790,1816]" name="Haiti" pageId="52" pageNumber="53">Haiti</collectingCountry>
, from arid areas on limestone terraces near the sea, bear some resemblance to
<taxonomicName id="4C1A4D4FFFCB1F5BFF50F89AFF078F27" authorityName="Burret" authorityYear="1929" box="[136,254,1826,1853]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B96EEADEFFCB1F5BFF50F89AFF078F27" box="[136,254,1826,1853]" italics="true" pageId="52" pageNumber="53">C. gracilis</emphasis>
</taxonomicName>
and occur about
<quantity id="4CE29B29FFCB1F5BFE66F89BFDFC8F27" box="[446,517,1826,1852]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="km" value="60.0">60 km</quantity>
west of its locality. They lack fruits and remain unidentified. Two sterile specimens (
<emphasis id="B96EEADEFFCB1F5BFF49F8FFFE808F7A" box="[145,377,1862,1889]" italics="true" pageId="52" pageNumber="53">Leonard 7172, 8525</emphasis>
), both from St. Michel de lAtalaye in north-central
<collectingCountry id="F30D765CFFCB1F5BFC07F8FFFBE38F7B" box="[991,1050,1862,1888]" name="Haiti" pageId="52" pageNumber="53">Haiti</collectingCountry>
, also bear some resemblance to
<taxonomicName id="4C1A4D4FFFCB1F5BFA4AF8FEFF278F9F" authorityName="Burret" authorityYear="1929" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B96EEADEFFCB1F5BFA4AF8FEFF278F9F" italics="true" pageId="52" pageNumber="53">C. gracilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>