161 lines
20 KiB
XML
161 lines
20 KiB
XML
<document id="8FB4EEEDB93561D04B2187CDCC2C0D8C" ID-CLB-Dataset="29087" ID-DOI="10.11646/zootaxa.4455.3.4" ID-GBIF-Dataset="c9bf5059-abe4-414e-a7c2-f3f9e75f2979" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457467" ID-ZooBank="69209FF7-80B0-409D-A257-0B88C65E903F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262799757" checkinUser="plazi" docAuthor="Kamiński, Marcin J. & Schoeman, Colin S." docDate="2018" docId="038BC66EFFDE965FF7ABFBE2FEEAFCE7" docLanguage="en" docName="zootaxa.4455.3.4.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaxius pseudoloensus Kamiński & Schoeman 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="482" masterDocId="FFB2BE16FFD49654F73CFFF7FF95FFF2" masterDocTitle="Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)" masterLastPageNumber="485" masterPageNumber="471" pageNumber="481" updateTime="1698576062251" updateUser="ExternalLinkService">
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<mods:mods id="C1CB85AC46E4BAE7A1F705902B603BE2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0AF4CFB73ACE3F99AC5416EDF3CBCC4B">
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<mods:title id="340FC9E9774B29F7176A9D29DBA27D71">Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)</mods:title>
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<mods:name id="298E892A2316BCD560E5BC519B594989" type="personal">
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<mods:namePart id="F46A021C194B4CA1BDB4F162B6C37F55">Kamiński, Marcin J.</mods:namePart>
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<mods:roleTerm id="8EF834FDC37D3252EEBF0E2CA545ED06">Author</mods:roleTerm>
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<mods:namePart id="D88B6F45E4B5B6D4F46AA5EB545E453D">Schoeman, Colin S.</mods:namePart>
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<mods:typeOfResource id="A2B70289C0A6A856E03DB11469C6B264">text</mods:typeOfResource>
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<mods:title id="DB9BFCA7E261CAEC42D03EF62F898039">Zootaxa</mods:title>
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<mods:part id="E1AA3CBAB218DF80FE8CCCE3DBDF40DC">
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<mods:date id="DD5F2422A66C227BE186BB046AE36006">2018</mods:date>
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<mods:number id="B4758B1E8D431817B84B352F679A45CD">2018-08-03</mods:number>
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<mods:number id="456D323060C107B7395C68D38A3EF262">4455</mods:number>
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<mods:classification id="C16724FD5B7CCB931672E52AB5287512">journal article</mods:classification>
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<treatment id="038BC66EFFDE965FF7ABFBE2FEEAFCE7" ID-DOI="http://doi.org/10.5281/zenodo.6490575" ID-GBIF-Taxon="148691415" ID-Zenodo-Dep="6490575" LSID="urn:lsid:plazi:treatment:038BC66EFFDE965FF7ABFBE2FEEAFCE7" httpUri="http://treatment.plazi.org/id/038BC66EFFDE965FF7ABFBE2FEEAFCE7" lastPageId="11" lastPageNumber="482" pageId="10" pageNumber="481">
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<subSubSection id="C33824F3FFDE965EF7ABFBE2FB31FBA2" pageId="10" pageNumber="481" type="nomenclature">
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<paragraph id="8B9D7778FFDE965EF7ABFBE2FD8DFBDD" blockId="10.[151,1188,1045,1141]" box="[151,536,1045,1071]" pageId="10" pageNumber="481">
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<heading id="D0D5C014FFDE965EF7ABFBE2FD8DFBDD" bold="true" box="[151,536,1045,1071]" fontSize="11" level="1" pageId="10" pageNumber="481" reason="1">
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<emphasis id="B956AB6AFFDE965EF7ABFBE2FD8DFBDD" bold="true" box="[151,536,1045,1071]" pageId="10" pageNumber="481">
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<taxonomicName id="4C220CFBFFDE965EF7ABFBE2FE26FBDD" authority="Kamiński & Schoeman, 2018" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[151,435,1045,1071]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="pseudoloensus" status="sp. nov.">
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<emphasis id="B956AB6AFFDE965EF7ABFBE2FE26FBDD" bold="true" box="[151,435,1045,1071]" italics="true" pageId="10" pageNumber="481">Anaxius pseudoloensus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651611FFDE965EF686FBE2FD8DFBDD" box="[442,536,1045,1071]" pageId="10" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9D7778FFDE965EF7ABFBC0FB31FBA2" blockId="10.[151,1188,1045,1141]" box="[151,1188,1079,1104]" pageId="10" pageNumber="481">http://zoobank.org/urn:lsid:zoobank.org:act:13054B1E-F5D9-43A0-BAE0-BAA29297C843</paragraph>
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</subSubSection>
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||
<subSubSection id="C33824F3FFDE965EF7ABFBABFEDAFB87" box="[151,335,1116,1141]" pageId="10" pageNumber="481" type="description">
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<paragraph id="8B9D7778FFDE965EF7ABFBABFEDAFB87" blockId="10.[151,1188,1045,1141]" box="[151,335,1116,1141]" pageId="10" pageNumber="481">
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(
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<figureCitation id="13196BFDFFDE965EF7A3FBABFF6AFB86" box="[159,255,1116,1141]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="10" pageNumber="481">Figs. 1G</figureCitation>
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,
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<figureCitation id="13196BFDFFDE965EF637FBABFEB9FB86" box="[267,300,1116,1140]" captionStart="FIGURE 3" captionStartId="7.[151,250,1440,1462]" captionTargetBox="[200,1386,725,1434]" captionTargetId="figure@7.[200,1386,725,1434]" captionTargetPageId="7" captionText="FIGURE 3. Aedeagal tegmen morphology. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457473/files/figure.png" pageId="10" pageNumber="481">3G</figureCitation>
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,
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<figureCitation id="13196BFDFFDE965EF604FBABFED3FB87" box="[312,326,1116,1141]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="10" pageNumber="481">5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33824F3FFDE965EF7ABFB53FC57FB4F" box="[151,962,1188,1213]" pageId="10" pageNumber="481" type="materials_examined">
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<paragraph id="8B9D7778FFDE965EF7ABFB53FC57FB4F" blockId="10.[151,1437,1188,2005]" box="[151,962,1188,1213]" pageId="10" pageNumber="481">
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<emphasis id="B956AB6AFFDE965EF7ABFB53FE1DFB4F" bold="true" box="[151,392,1188,1213]" pageId="10" pageNumber="481">Type data. Holotype</emphasis>
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, male (TMSA): “Espungabera / 80.K.Jutha 1954”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33824F3FFDE965EF7FBFB30FDE7FAF7" pageId="10" pageNumber="481" type="etymology">
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<paragraph id="8B9D7778FFDE965EF7FBFB30FDE7FAF7" blockId="10.[151,1437,1188,2005]" pageId="10" pageNumber="481">
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<emphasis id="B956AB6AFFDE965EF7FBFB30FEDBFB12" bold="true" box="[199,334,1223,1248]" pageId="10" pageNumber="481">Etymology.</emphasis>
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The name highlights the morphological resemblance of this species to representatives of the genus
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<taxonomicName id="4C220CFBFFDE965EF7ABFB1AFE16FAF6" authority="Lucas, 1920" authorityName="Lucas" authorityYear="1920" box="[151,387,1260,1284]" class="Insecta" family="Tenebrionidae" genus="Loensus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="genus">
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<emphasis id="B956AB6AFFDE965EF7ABFB1AFF66FAF6" box="[151,243,1261,1284]" italics="true" pageId="10" pageNumber="481">Loensus</emphasis>
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Lucas, 1920
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</taxonomicName>
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(
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<taxonomicName id="4C220CFBFFDE965EF6ADFB1BFE60FAF7" authorityName="Eschscholtz" authorityYear="1829" box="[401,501,1260,1285]" class="Insecta" family="Tenebrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="tribe" tribe="Pedinini">Pedinini</taxonomicName>
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:
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<taxonomicName id="4C220CFBFFDE965EF6C3FB1BFDF2FAF7" baseAuthorityName="Mortensen" baseAuthorityYear="1939" box="[511,615,1260,1285]" class="Echinoidea" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="481" phylum="Echinodermata" rank="subOrder" subOrder="Pedinina">Pedinina</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33824F3FFDE965EF7FBFAF8FDC0FA4A" pageId="10" pageNumber="481" type="diagnosis">
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<paragraph id="8B9D7778FFDE965EF7FBFAF8FDC0FA4A" blockId="10.[151,1437,1188,2005]" pageId="10" pageNumber="481">
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<emphasis id="B956AB6AFFDE965EF7FBFAF8FED4FADA" bold="true" box="[199,321,1295,1320]" pageId="10" pageNumber="481">Diagnosis.</emphasis>
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This species is similar to
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<taxonomicName id="4C220CFBFFDE965EF55AFAE7FC92FADA" authorityName="Koch" authorityYear="1958" box="[614,775,1296,1320]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="campbellae">
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<emphasis id="B956AB6AFFDE965EF55AFAE7FC92FADA" box="[614,775,1296,1320]" italics="true" pageId="10" pageNumber="481">A. campbellae</emphasis>
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</taxonomicName>
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by having,
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<emphasis id="B956AB6AFFDE965EF4B0FAE7FC66FADA" box="[908,1011,1296,1320]" italics="true" pageId="10" pageNumber="481">inter alia</emphasis>
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, protibia with shallow median cavity, widened protarsus, with subsequent tarsomeres slightly narrower, and not apically depressed intercoxal process of prostenum (lateral view).
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<taxonomicName id="4C220CFBFFDE965EF681FAAFFD56FA82" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[445,707,1368,1392]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="pseudoloensus">
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<emphasis id="B956AB6AFFDE965EF681FAAFFD56FA82" box="[445,707,1368,1392]" italics="true" pageId="10" pageNumber="481">Anaxius pseudoloensus</emphasis>
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</taxonomicName>
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can be easily distinguished by its smaller body size (
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<quantity id="4CDADA9DFFDE965EF219FAA0FAEBFA9D" box="[1317,1406,1367,1392]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="481" unit="mm" value="4.5">4.5 mm</quantity>
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in
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<taxonomicName id="4C220CFBFFDE965EF7ABFA8BFEAFFA66" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[151,314,1404,1428]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="pseudoloensus">
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<emphasis id="B956AB6AFFDE965EF7ABFA8BFEAFFA66" box="[151,314,1404,1428]" italics="true" pageId="10" pageNumber="481">pseudoloensus</emphasis>
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</taxonomicName>
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; 6.5–7.0 in
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<taxonomicName id="4C220CFBFFDE965EF6F7FA8BFDD9FA66" authorityName="Koch" authorityYear="1958" box="[459,588,1404,1428]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="campbellae">
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<emphasis id="B956AB6AFFDE965EF6F7FA8BFDD9FA66" box="[459,588,1404,1428]" italics="true" pageId="10" pageNumber="481">campbellae</emphasis>
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</taxonomicName>
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), more slender elytra (
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<figureCitation id="13196BFDFFDE965EF46BFA8BFC21FA66" box="[855,948,1404,1429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="10" pageNumber="481">Figs 1B</figureCitation>
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vs 1G), and simple mesofemora in males (with preapical denticle in
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<taxonomicName id="4C220CFBFFDE965EF6FFFA57FDD1FA4A" authorityName="Koch" authorityYear="1958" box="[451,580,1440,1464]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="campbellae">
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<emphasis id="B956AB6AFFDE965EF6FFFA57FDD1FA4A" box="[451,580,1440,1464]" italics="true" pageId="10" pageNumber="481">campbellae</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33824F3FFDE965FF7FBFA33FA93FD3F" lastPageId="11" lastPageNumber="482" pageId="10" pageNumber="481" type="description">
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<paragraph id="8B9D7778FFDE965EF7FBFA33FC17FA2E" blockId="10.[151,1437,1188,2005]" box="[199,898,1476,1501]" pageId="10" pageNumber="481">
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<emphasis id="B956AB6AFFDE965EF7FBFA33FECCFA2F" bold="true" box="[199,345,1476,1501]" pageId="10" pageNumber="481">Description.</emphasis>
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Body—length ca.
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<quantity id="4CDADA9DFFDE965EF510FA33FD10FA2E" box="[556,645,1476,1501]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="481" unit="mm" value="4.5">4.5 mm</quantity>
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. Habitus as in
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<figureCitation id="13196BFDFFDE965EF414FA33FCEBFA2E" box="[808,894,1476,1501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="10" pageNumber="481">Fig. 1G</figureCitation>
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.
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</paragraph>
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<paragraph id="8B9D7778FFDE965EF7FBFA10FB12F8D2" blockId="10.[151,1437,1188,2005]" pageId="10" pageNumber="481">
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Dorsal side of head covered with fine punctures (0.5–1.0 diameters apart). Frontoclypeal suture not visible. Canthus rounded at base; narrower than head at level of eyes. Clypeal emargination shallow (clypeal emargination width: depth ratio ca. 8.0). Labrum wide (width: length ratio ca. 1.9); emarginate medially; with basal membrane exposed. Ligula exposed, not covered by mentum. Mentum widest in apical part; with well-developed medial keel; with exposed lateral lobes; not covering cardo or basistipes; apically with median indentation. Submentum pentagonal; short. Apical segment of maxillary palpus trapezoidal; with sensory field occupying whole proximal edge. Second segment of maxillary palpus elongate (length: width ratio ca. 2.4). Palpifer located laterally on basistipes. Anterior tentorial pit circular. Eye not dorsally shielded by any keel. Margin between maxillary fossa and antenna insertion well marked. Antenna not available for study (broken off
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<typeStatus id="5499C9DAFFDE965EF328F8F0FBEEF8D2" box="[1044,1147,1799,1824]" pageId="10" pageNumber="481" type="holotype">holotype</typeStatus>
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).
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</paragraph>
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<paragraph id="8B9D7778FFDE965FF7FBF8DAFEE6FF0A" blockId="10.[151,1437,1188,2005]" lastBlockId="11.[151,1437,151,789]" lastPageId="11" lastPageNumber="482" pageId="10" pageNumber="481">Pronotum slightly narrower than elytra (elytra: pronotum width ratio ca. 1.1); relatively narrow (length: width ratio of pronotal disc ca. 0.8); disc shiny, covered with fine punctures (0.5–1.0 diameters apart). Lateral sides of pronotal disc straight at basal half. Disc without apophyseal depressions. Basal, anterior, and lateral emargination of pronotal disc present. Base of pronotum straight. Posterior angles not protruding beyond base. Pronotal hypomeron dull; covered with very fine longitudinal rugosities; without submarginal indentation. Intercoxal process of prosternum not depressed apically (lateral view); apically with oval depression on ventral surface; slightly protruding towards first abdominal segment. Procoxae narrowly separated (procoxa: intercoxal process width ratio ca. 2.0).</paragraph>
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<paragraph id="8B9D7778FFDF965FF7FBFEF3FB12FE7A" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="482">Five elytral intervals finely marked; remaining ones indistinguishable. First interval elevated in disc. Elytron covered by fine punctures (0.1 diameters apart; sometimes confluent). Elytral base straight; not emarginate. Elytral humerus rounded. Epipleuron slightly narrowing apically, then of constant width; basally covering all elytral intervals (ventral view). Scutellum relatively large; triangular. Metathoracic wings absent.</paragraph>
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<paragraph id="8B9D7778FFDF965FF7FBFE63FAF5FE06" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="482">Metaventrite extremely short (metacoxal cavity: metaventrite (between insertions of meso- and metacoxae) length ratio ca. 8.0). Process of first abdominal ventrite wide (distances between mesocoxae: metacoxae ratio ca. 0.9). Fifth abdominal ventrite without submarginal sulcus; covered with fine punctures (4–5 diameters apart).</paragraph>
|
||
<paragraph id="8B9D7778FFDF965FF7FBFE08FD26FD92" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="482">
|
||
Male protarsus widened, with subsequent tarsomeres slightly narrower. Protibia with shallow longitudinal cavity. Metafemora with fringe of setae distributed on whole length. Other leg parts simple. Meso- and metatibiae not avaliable for study (broken out of
|
||
<typeStatus id="5499C9DAFFDF965FF57CFDB0FD32FD92" box="[576,679,583,608]" pageId="11" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9D7778FFDF965FF7FBFD9BFA93FD3F" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="482">Aedeagal tegmen widest in middle; unipartite; without basal gap. Apical part divided (up to 0.10 of whole tegmen length). Clava straight. Penis of even width along most of its length; with apical part not covered by tegmen, exposed dorsally. Basal apophyses of penis relatively short (ca. 0.4 of whole tegmen length).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33824F3FFDF965FF7FBFD20FEEAFCE7" pageId="11" pageNumber="482" type="distribution">
|
||
<paragraph id="8B9D7778FFDF965FF7FBFD20FEEAFCE7" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="482">
|
||
<emphasis id="B956AB6AFFDF965FF7FBFD20FEF5FD02" bold="true" box="[199,352,727,752]" pageId="11" pageNumber="482">Distribution.</emphasis>
|
||
This species has been collected in the following ecoregion of
|
||
<collectingCountry id="F33537E8FFDF965FF30FFD20FB52FD02" box="[1075,1223,727,752]" name="Mozambique" pageId="11" pageNumber="482">Mozambique</collectingCountry>
|
||
(
|
||
<figureCitation id="13196BFDFFDF965FF3E4FD20FA8AFD02" box="[1240,1311,727,752]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="11" pageNumber="482">Fig. 5</figureCitation>
|
||
): Southern Miombo woodlands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |