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<document id="6C2F4FF7F7A14CFF9290D9A728C87AE7" ID-CLB-Dataset="4031" ID-DOI="10.5852/ejt.2021.750.1365" ID-GBIF-Dataset="818d0bc4-31a6-4319-86a3-6e6b16ab72df" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451747" ID-ZooBank="CE5D8517-AC04-4AF9-BD78-A2D6E21D33A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516667691" checkinUser="felipe" docAuthor="Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z. &amp; Ahrens, Dirk" docDate="2021" docId="038B976CFF86FFE3CA29C82C148DC9EC" docLanguage="en" docName="ejt-750_94-123.pdf" docOrigin="European Journal of Taxonomy 750 (750)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1365" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astaena nigrona L. W. Saylor 1947" docType="treatment" docVersion="10" lastPageNumber="107" masterDocId="FFB2EF14FF8BFFEEC84EC925155FCE04" masterDocTitle="Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor" masterLastPageNumber="123" masterPageNumber="94" pageNumber="107" updateTime="1699290068827" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="89C4D54A2F097C160F093035FF283D1B">Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor</mods:title>
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<mods:namePart id="B35CE68FD12AE34937D9601FFC8E3B21">Pacheco, Thaynara L.</mods:namePart>
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<mods:affiliation id="F5EB793AE257201DE260020A9FF4B89A">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="4ECDE24EE84BDA69EEA7336B9A3447ED" type="email">thay.lpacheco@gmail.com</mods:nameIdentifier>
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<mods:name id="063CE4F4EB31519AAC8F4347615F945C" type="personal">
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<mods:namePart id="C9D488FA50B0E60F67F70BC52F301DEE">Monné, Marcela L.</mods:namePart>
<mods:nameIdentifier id="7C17F82B86F3B0531A1459D70B1C7F97" type="LSID">86B186B4-4297-4380-A98D-FC813B5D53A6&amp;2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="469EC5BBEB93C214B9ECD9FD7E1D51B8">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="CF220B9912E98F7F5B6764B4B06B92B3" type="email">mlmonne2@gmail.com&amp;vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="135F735CA38E69DA2A15323619747D4C">Author</mods:roleTerm>
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<mods:namePart id="DF178E9CFE6C75DA4B25C3B6D54B6DB9">Vaz-De, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="2DB3C160244C41FF8BB633F71476103D" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="004F1AC5273EE6E7A5DB61F70E5724AD">Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A33B8B9FB3E11796DE454AC69565604B" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0602CD651A9AC681B90DACC51A869577">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="4FBB10ED80FB69D88EA29D3763A916CF" type="LSID">DEDCE5CF-AA11-4BBF-A2C6-D7C815019714</mods:nameIdentifier>
<mods:affiliation id="31CBD9067AACFC1B69B7C5DDB6E7ABA2">Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="9C3A00F52CA561AD96651E8A0CDAE484" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="F5F7EDE2D73646AF45DC44E59AF164CF">European Journal of Taxonomy</mods:title>
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<mods:date id="525984661264DECB43D79DE9A36BB8A0">2021</mods:date>
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<mods:number id="B52B6CB2FDAD16AD7BAB51CEE7661F98">2021-05-19</mods:number>
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<mods:identifier id="B5BA50BAE21F86CD8CB68CFACBDD7CA9" type="DOI">10.5852/ejt.2021.750.1365</mods:identifier>
<mods:identifier id="1360E0F156EA6D06EF2A05A18710D0CD" type="GBIF-Dataset">818d0bc4-31a6-4319-86a3-6e6b16ab72df</mods:identifier>
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<treatment id="038B976CFF86FFE3CA29C82C148DC9EC" ID-DOI="http://doi.org/10.5281/zenodo.7401676" ID-GBIF-Taxon="188647624" ID-Zenodo-Dep="7401676" LSID="urn:lsid:plazi:treatment:038B976CFF86FFE3CA29C82C148DC9EC" httpUri="http://treatment.plazi.org/id/038B976CFF86FFE3CA29C82C148DC9EC" lastPageNumber="107" pageId="13" pageNumber="107">
<subSubSection id="C33875F1FF86FFE3CA29C82C1692CF27" box="[615,973,265,291]" pageId="13" pageNumber="107" type="nomenclature">
<paragraph id="8B9D267AFF86FFE3CA29C82C1692CF27" blockId="13.[615,973,265,331]" box="[615,973,265,291]" pageId="13" pageNumber="107">
<heading id="D0D59116FF86FFE3CA29C82C1692CF27" box="[615,973,265,291]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="107" reason="2">
<taxonomicName id="4C225DF9FF86FFE3CA29C82C1692CF27" authority="Saylor, 1947" authorityName="L. W. Saylor" authorityYear="1947" box="[615,973,265,291]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrona">
<emphasis id="B956FA68FF86FFE3CA29C82C166FCF27" bold="true" box="[615,816,265,291]" italics="true" pageId="13" pageNumber="107">Astaena nigrona</emphasis>
Saylor, 1947
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF86FFE3CA8EC815162BCF4F" box="[704,884,304,331]" pageId="13" pageNumber="107" type="description">
<paragraph id="8B9D267AFF86FFE3CA8EC815162BCF4F" blockId="13.[615,973,265,331]" box="[704,884,304,331]" pageId="13" pageNumber="107">
<figureCitation id="13193AFFFF86FFE3CA8EC815161ACF4E" box="[704,837,304,330]" captionStart="Fig" captionStartId="12.[188,231,1858,1884]" captionTargetBox="[244,1343,265,1819]" captionTargetId="figure-97@12.[244,1343,265,1819]" captionTargetPageId="12" captionText="Fig. 3. A–E. Astaena incachaca Saylor, 1946, holotype, ♂ (Type No. 7913; CAS). – F–J. A. maqueta Saylor, 1947, holotype, ♂ (Type No. 7914; CAS). – K–O. A. nigrona Saylor, 1947, holotype, ♂ (Type No. 7915; CAS). A, G, M. Aedeagus, lateral view. B, F, N. Parameres, dorsal view. C, H, O. labels. D, I, L. Habitus, dorsal view. E, J, K. Habitus, lateral view. Scale bars: A–B, F–G, M–N = 0.5 mm; D–E, I–L = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451753" httpUri="https://zenodo.org/record/5451753/files/figure.png" pageId="13" pageNumber="107">Figs 3K–O</figureCitation>
,
<figureCitation id="13193AFFFF86FFE3CB1FC814162BCF4F" box="[849,884,305,331]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[261,1326,295,1750]" captionTargetId="figure-12@26.[241,1347,265,1784]" captionTargetPageId="26" captionText="Fig. 7. Distribution of type localities of the South American Sericini species described by Saylor. A. Astaena abaca Saylor, 1946, A. biciliata Saylor, 1946, A. bogotana Saylor, 1946, A. excisipes Saylor, 1947, A. explaniceps Saylor, 1947, A. incachaca Saylor, 1946, A. maqueta Saylor, 1947, and A. nigrona Saylor, 1947. B. A. fusagona Saylor, 1946, A. pygidia Saylor, 1946, A. rugithorax Saylor, 1946, A. salta Saylor, 1946, A. yungasa Saylor, 1946, and A. zyrota Saylor, 1946. C. Sayloria abcora (Saylor, 1946) comb. nov., S. bicoloripes (Saylor, 1946) comb. nov., and S. pottsi (Saylor, 1946) comb. nov.." figureDoi="http://doi.org/10.5281/zenodo.5451761" httpUri="https://zenodo.org/record/5451761/files/figure.png" pageId="13" pageNumber="107">7A</figureCitation>
</paragraph>
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<subSubSection id="C33875F1FF86FFE3C8F3C853168ECC47" pageId="13" pageNumber="107" type="reference_group">
<paragraph id="8B9D267AFF86FFE3C8F3C8531733CF95" blockId="13.[189,620,374,401]" box="[189,620,374,401]" pageId="13" pageNumber="107">
<treatmentCitationGroup id="AB320154FF86FFE3C8F3C8531733CF95" box="[189,620,374,401]" pageId="13" pageNumber="107">
<treatmentCitation id="0A83006BFF86FFE3C8F3C8531736CF95" author="Saylor L. W." box="[189,617,374,401]" page="435" pageId="13" pageNumber="107" year="1947">
<taxonomicName id="4C225DF9FF86FFE3C8F3C8531736CF95" authority="Saylor, 1947 b: 435" authorityName="L. W. Saylor" authorityPageNumber="435" authorityYear="1947" box="[189,617,374,401]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrona">
<emphasis id="B956FA68FF86FFE3C8F3C8531420CF94" box="[189,383,374,400]" italics="true" pageId="13" pageNumber="107">Astaena nigrona</emphasis>
<bibRefCitation id="EFB35B8BFF86FFE3C9C8C8531736CF95" author="Saylor L. W." box="[390,617,374,401]" pageId="13" pageNumber="107" pagination="433 - 437" refId="ref14000" refString="Saylor L. W. 1947 b. New South American scarab beetles of the genus Astaena. Revista de Entomologia 18: 433 - 437." type="journal article" year="1947">Saylor, 1947b: 435</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3C8991661CFD2" blockId="13.[189,830,443,470]" box="[189,830,443,470]" pageId="13" pageNumber="107">
<treatmentCitationGroup id="AB320154FF86FFE3C8F3C8991661CFD2" box="[189,830,443,470]" pageId="13" pageNumber="107">
<taxonomicName id="4C225DF9FF86FFE3C8F3C8991420CFD1" authority="Frey, 1973" authorityName="L. W. Saylor" authorityYear="1947" box="[189,383,444,470]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrona">
<emphasis id="B956FA68FF86FFE3C8F3C8991420CFD1" box="[189,383,444,470]" italics="true" pageId="13" pageNumber="107">Astaena nigrona</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A83006BFF86FFE3C9D2C8991710CFD2" author="Frey G." box="[412,591,444,470]" page="331" pageId="13" pageNumber="107" year="1973">
<bibRefCitation id="EFB35B8BFF86FFE3C9D2C8991710CFD2" author="Frey G." box="[412,591,444,470]" pageId="13" pageNumber="107" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey 1973: 331</bibRefCitation>
</treatmentCitation>
(identification key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CB27168ECC47" blockId="13.[189,1398,513,579]" pageId="13" pageNumber="107">
<treatmentCitationGroup id="AB320154FF86FFE3C8F3CB27168ECC47" pageId="13" pageNumber="107">
non
<taxonomicName id="4C225DF9FF86FFE3C8BECB27149ECC1F" box="[240,449,514,540]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigroana">
<emphasis id="B956FA68FF86FFE3C8BECB27149ECC1F" box="[240,449,514,540]" italics="true" pageId="13" pageNumber="107">Astaena nigroana</emphasis>
</taxonomicName>
(incorrect subsequent spelling) –
<treatmentCitation id="0A83006BFF86FFE3CB1CCB271155CC18" author="Evans A. V." box="[850,1034,513,540]" page="25" pageId="13" pageNumber="107" year="2003">
<bibRefCitation id="EFB35B8BFF86FFE3CB1CCB271155CC18" author="Evans A. V." box="[850,1034,513,540]" pageId="13" pageNumber="107" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 25</bibRefCitation>
</treatmentCitation>
(checklist). — Evans &amp;
<bibRefCitation id="EFB35B8BFF86FFE3CD7ECB241029CC1F" author="Smith A. B. T. &amp; Evans A. V." box="[1328,1398,513,539]" pageId="13" pageNumber="107" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith</bibRefCitation>
2005: 10 (checklist); 2007: 10 (checklist); 2009: 10 (checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF86FFE3C8F3CB4B11EBCD9A" pageId="13" pageNumber="107" type="materials_examined">
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<emphasis id="B956FA68FF86FFE3C8F3CB4B14B3CC8C" bold="true" box="[189,492,622,648]" pageId="13" pageNumber="107">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF86FFE3C8A4CB841609CD0E" blockId="13.[189,1398,673,778]" pageId="13" pageNumber="107">
<materialsCitation id="3B4A2C27FF86FFE3C8A4CB841609CD0E" ID-GBIF-Occurrence="3383726326" collectionCode="CAS" collectorName="L. W. Saylor Collection" country="Colombia" location="Rio Chil. Colum" municipality="California Academy of Sciences Type" pageId="13" pageNumber="107" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B956FA68FF86FFE3C8A4CB841403CCBF" bold="true" box="[234,348,673,699]" pageId="13" pageNumber="107">
<typeStatus id="549998D8FF86FFE3C8A4CB841403CCBF" box="[234,348,673,699]" pageId="13" pageNumber="107" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F33566EAFF86FFE3C8F3CBED1407CCE6" box="[189,344,712,738]" name="Colombia" pageId="13" pageNumber="107">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF86FFE3C93ACBEC14D2CCE7" box="[372,397,713,739]" pageId="13" pageNumber="107" type="male">♂</specimenCount>
; “
<location id="8EFD70A1FF86FFE3C9EBCBED1731CCE6" LSID="urn:lsid:plazi:treatment:038B976CFF86FFE3CA29C82C148DC9EC:8EFD70A1FF86FFE3C9EBCBED1731CCE6" box="[421,622,712,738]" country="Colombia" municipality="California Academy of Sciences Type" name="Rio Chil. Colum" pageId="13" pageNumber="107">Rio Chil. Colum</location>
/ L.W. Saylor Collection / California Academy of Sciences Type nº 7915 / Holotype
<taxonomicName id="4C225DF9FF86FFE3C9ECCBD517A0CD0E" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1947" box="[418,767,752,778]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrona">
<emphasis id="B956FA68FF86FFE3C9ECCBD5173BCD0D" box="[418,612,752,778]" italics="true" pageId="13" pageNumber="107">Astaena nigrona</emphasis>
L.W. Saylor
</taxonomicName>
”;
<collectionCode id="ED33BEBFFF86FFE3CB5BCBD5160DCD0E" box="[789,850,752,778]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="107" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D267AFF86FFE3C8A4CA1011EBCD9A" blockId="13.[189,1398,821,926]" pageId="13" pageNumber="107">
<materialsCitation id="3B4A2C27FF86FFE3C8A4CA1011EFCD9A" ID-GBIF-Occurrence="3383726335" collectionCode="CAS" collectorName="L. W. Saylor Collection" country="Colombia" location="Bogota" pageId="13" pageNumber="107" specimenCount="1" specimenCount-female="1" stateProvince="Bogota" typeStatus="paratype">
<emphasis id="B956FA68FF86FFE3C8A4CA101403CD4B" bold="true" box="[234,348,821,847]" pageId="13" pageNumber="107">
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<collectingCountry id="F33566EAFF86FFE3C8F3CA781407CD73" box="[189,344,861,887]" name="Colombia" pageId="13" pageNumber="107">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF86FFE3C920CA7814C5CD73" box="[366,410,861,887]" pageId="13" pageNumber="107" type="female">1 ♀</specimenCount>
; “
<collectingRegion id="49E6E898FF86FFE3C9FFCA781756CD73" box="[433,521,861,887]" country="Colombia" name="Bogota" pageId="13" pageNumber="107">Bogota</collectingRegion>
VI- Colum / L.W. Saylor Collection / Collection of the California Academy of Sciences, San Francisco, California / Allotype
<taxonomicName id="4C225DF9FF86FFE3CB4ECAA11102CD9A" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1947" box="[768,1117,900,926]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrona">
<emphasis id="B956FA68FF86FFE3CB4ECAA1169DCD9A" box="[768,962,900,926]" italics="true" pageId="13" pageNumber="107">Astaena nigrona</emphasis>
L.W. Saylor
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”;
<collectionCode id="ED33BEBFFF86FFE3CC3DCAA111EFCD9A" box="[1139,1200,900,926]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="107" type="Museum">CAS</collectionCode>
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.
</paragraph>
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<subSubSection id="C33875F1FF86FFE3C8F3CAEF148DC9EC" pageId="13" pageNumber="107" type="description">
<paragraph id="8B9D267AFF86FFE3C8F3CAEF1483CDE7" blockId="13.[189,476,969,996]" box="[189,476,969,996]" pageId="13" pageNumber="107">
<heading id="D0D59116FF86FFE3C8F3CAEF1483CDE7" bold="true" box="[189,476,969,996]" fontSize="11" level="3" pageId="13" pageNumber="107" reason="3">
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<typeStatus id="549998D8FF86FFE3C8F3CAEF1470CDE0" box="[189,303,970,996]" pageId="13" pageNumber="107" type="holotype">Holotype</typeStatus>
redescription
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</heading>
</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CAD917A0CA13" blockId="13.[189,767,1020,1047]" box="[189,767,1020,1047]" pageId="13" pageNumber="107">
MEASUREMENTS. Length:
<quantity id="4CDA8B9FFF86FFE3C9A8CAD91763CA13" box="[486,572,1020,1047]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="107" unit="mm" value="12.0">12 mm</quantity>
. Width:
<quantity id="4CDA8B9FFF86FFE3CAD0CAD917A4CA13" box="[670,763,1020,1047]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="107" unit="mm" value="5.8">5.8 mm</quantity>
.
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<paragraph id="8B9D267AFF86FFE3C8F3CD671194CAD7" blockId="13.[187,1401,1090,1235]" pageId="13" pageNumber="107">HEAD. Reddish-brown. Labroclypeus, shape of anterior margin weakly sinuated medially. Ocular canthus without terminal setae, ratio to eye: 1/4. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club/remaining antennomeres: 1/1.5; ratio third vs fourth antennomere: 1/3; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1/5.</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CDDB1703C800" blockId="13.[188,1399,1278,1540]" pageId="13" pageNumber="107">PRONOTUM AND ELYTRA. Surface of pronotum and elytra with setae. Pronotum bicolored, reddish-brown with two red spots on laterals, widest at middle, anterior marginal line present, posterior marginal line absent, lateral margin strongly convex and smooth (not serrate), antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally flattened, punctation on disc dense, setae of lateral margin sparse, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, blackish-brown, surface opaque, microsculpture absent, punctation on intervals present, diameter of strial and interval punctation equal.</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CF1517BEC875" blockId="13.[189,1398,1583,1649]" pageId="13" pageNumber="107">ABDOMEN. Lateral carina present. Second visible ventrite with row of setae in medial region. Apical visible ventrite without suture to apical tergite.</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CFB8147CC9A7" blockId="13.[188,1399,1692,1955]" pageId="13" pageNumber="107">LEGS. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1/3. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws asymmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw truncate.</paragraph>
<paragraph id="8B9D267AFF86FFE3C8F3CEEB148DC9EC" blockId="13.[189,466,1998,2024]" box="[189,466,1998,2024]" pageId="13" pageNumber="107">
AEDEAGUS.
<figureCitation id="13193AFFFF86FFE3C906CEEB1491C9EC" box="[328,462,1998,2024]" captionStart="Fig" captionStartId="12.[188,231,1858,1884]" captionTargetBox="[244,1343,265,1819]" captionTargetId="figure-97@12.[244,1343,265,1819]" captionTargetPageId="12" captionText="Fig. 3. A–E. Astaena incachaca Saylor, 1946, holotype, ♂ (Type No. 7913; CAS). – F–J. A. maqueta Saylor, 1947, holotype, ♂ (Type No. 7914; CAS). – K–O. A. nigrona Saylor, 1947, holotype, ♂ (Type No. 7915; CAS). A, G, M. Aedeagus, lateral view. B, F, N. Parameres, dorsal view. C, H, O. labels. D, I, L. Habitus, dorsal view. E, J, K. Habitus, lateral view. Scale bars: A–B, F–G, M–N = 0.5 mm; D–E, I–L = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451753" httpUri="https://zenodo.org/record/5451753/files/figure.png" pageId="13" pageNumber="107">Fig. 3M–N</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
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