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<document id="9C74C3760B0532A28381A89FED4123DD" ID-DOI="10.11646/zootaxa.3608.2.1" ID-GBIF-Dataset="dc20b97c-cdf4-467d-8ce9-98e912fe060c" ID-ISSN="1175-5326" ID-Zenodo-Dep="223235" ID-ZooBank="0E56C847-5694-4315-9A51-B4F207CA4F91" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460569345297" checkinUser="plazi" docAuthor="Criscione, Francesco &amp; Köhler, Frank" docDate="2013" docId="03D887BF0012250C929B10DC7757FC7D" docLanguage="en" docName="zt03608p115.pdf" docOrigin="Zootaxa 3608 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camaenidae Pilsbry 1895" docType="treatment" docVersion="9" lastPageNumber="105" masterDocId="FFE1FFC700112508920C14147605FFF2" masterDocTitle="Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae)" masterLastPageNumber="115" masterPageNumber="101" pageNumber="104" updateTime="1698321793467" updateUser="plazi">
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<mods:title id="E3830FE58FDFE700B2207420CF48F830">Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="EB92EC4E520BA8EE1204F0FD3380C28D">Criscione, Francesco</mods:namePart>
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<mods:namePart id="E11D0FC7E81EF66BA43FC5FCCB934ECE">Köhler, Frank</mods:namePart>
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<mods:title id="D35F094B8ADF35417038F8C1DC9DDB9B">Zootaxa</mods:title>
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<mods:date id="9A36007FB0B4CD7195DE35A54FCC5DB8">2013</mods:date>
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<treatment id="03D887BF0012250C929B10DC7757FC7D" ID-DOI="http://doi.org/10.5281/zenodo.6153543" ID-GBIF-Taxon="119558296" ID-Zenodo-Dep="6153543" LSID="urn:lsid:plazi:treatment:03D887BF0012250C929B10DC7757FC7D" httpUri="http://treatment.plazi.org/id/03D887BF0012250C929B10DC7757FC7D" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
<subSubSection id="C36B65220012250B929B10DC72F6FAD4" pageId="3" pageNumber="104" type="nomenclature">
<paragraph id="8BCE36A90012250B929B10DC77DEFB10" blockId="3.[151,475,1120,1250]" box="[151,475,1224,1250]" pageId="3" pageNumber="104">
<heading id="D08681C50012250B929B10DC77DEFB10" bold="true" box="[151,475,1224,1250]" fontSize="11" level="1" pageId="3" pageNumber="104" reason="1">
<taxonomicName id="4C714D2A0012250B929B10DC77DEFB10" ID-CoL="7NG9L" authority="Pilsbry, 1895" authorityName="Pilsbry" authorityYear="1895" box="[151,475,1224,1250]" class="Gastropoda" family="Camaenidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="104" phylum="Mollusca" rank="family">
<emphasis id="B905EABB0012250B929B10DC77DEFB10" bold="true" box="[151,475,1224,1250]" pageId="3" pageNumber="104">Camaenidae Pilsbry, 1895</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCE36A90012250B929B110472F6FAD4" blockId="3.[151,1267,1296,1318]" box="[151,1267,1296,1318]" pageId="3" pageNumber="104">
<taxonomicName id="4C714D2A0012250B929B110477CEFAD7" authority="Kohler, 2011" authorityName="Kohler" authorityYear="2011" box="[151,459,1296,1318]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="104" phylum="Mollusca" rank="genus">
<emphasis id="B905EABB0012250B929B11047744FAD4" box="[151,321,1296,1318]" italics="true" pageId="3" pageNumber="104">Australocosmica</emphasis>
Köhler, 2011
</taxonomicName>
<typeStatus id="54CA880B0012250B93DD11057406FAD4" box="[465,515,1297,1318]" pageId="3" pageNumber="104">Type</typeStatus>
species.
<taxonomicName id="4C714D2A0012250B9053110475FDFAD7" authority="Kohler, 2011" authorityName="Kohler" authorityYear="2011" box="[607,1016,1296,1318]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="104" phylum="Mollusca" rank="species" species="augustae">
<emphasis id="B905EABB0012250B90531104756EFAD4" box="[607,875,1296,1318]" italics="true" pageId="3" pageNumber="104">Australocosmica augustae</emphasis>
Köhler, 2011
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C36B65220012250B929B11477759F874" pageId="3" pageNumber="104" type="description">
<paragraph id="8BCE36A90012250B929B11477759F874" blockId="3.[151,1437,1362,1998]" pageId="3" pageNumber="104">
<emphasis id="B905EABB0012250B929B11477762FA99" box="[151,359,1363,1387]" italics="true" pageId="3" pageNumber="104">Generic diagnosis</emphasis>
. Shell between
<quantity id="4C899B4C0012250B9029114674C2FA98" box="[549,711,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.7" metricValueMin="0.75" pageId="3" pageNumber="104" unit="mm" value="12.25" valueMax="17.0" valueMin="7.5">7.5 to 17 mm</quantity>
in height and about
<quantity id="4C899B4C0012250B91B211467252FA98" box="[958,1111,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.8" metricValueMin="1.1" pageId="3" pageNumber="104" unit="mm" value="14.5" valueMax="18.0" valueMin="11.0">11 to 18 mm</quantity>
in diameter (
<tableCitation id="C6F303120012250B96FB1146734BFA98" box="[1271,1358,1362,1386]" captionStart="TABLE 1" captionStartId="4.[151,239,155,179]" captionText="TABLE 1. Shell dimensions (mm) and whorl counts of Australocosmica species recognised in the present work." pageId="3" pageNumber="104">Table 1</tableCitation>
), with broadly conical to semi-globose shape. Whorls wide in cross-section, convex, with well-rounded to slightly angulate periphery, slightly to moderately shouldered underneath deeply incised suture. Sculpture of moderately to well-pronounced, dense, regularly spaced axial ribs, becoming sharply less prominent on shell base. Microsculpture of small tubercles arranged in irregular spiral rows. Aperture simple, rounded, with no nodes, with thinly callused parietal wall. Outer lip thin, slightly expanded. Colour of shell horn to brown. Mantle roof with black mottled to spotted pigmentation. Penial retractor muscle attached to penial apex near entrance of vas deferens. Vas deferens entering penial complex apically, connect to lumen of penial chamber through a conspicuously thickened, furrowed, collar-like vergic papilla. Penial sheath well-developed, extending entire length, thin proximally, thick distally. No epiphallus present. Penis rather straight, with well-developed sheath, comparatively thick, reaching from base to well above apex of penis. Inner penial wall may support marked longitudinal pilasters, dense pustulation, or a combination of both. Vagina more or less tubular, usually only slightly inflated. Free oviduct straight, as long as bursa copulatrix. Bursa connected with spermoviduct by delicate connective tissue, reaching distal base of spermoviduct; duct and head well differentiated. Albumen gland elongate. Talon situated within anterior quarter of albumen gland. Species aestivate as free sealers (updated, based on Köhler, 2011).
</paragraph>
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<subSubSection id="C36B65220012250C92CB13877341FF1A" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104" type="discussion">
<paragraph id="8BCE36A90012250B92CB138775AEF83C" blockId="3.[151,1437,1362,1998]" pageId="3" pageNumber="104">
<emphasis id="B905EABB0012250B92CB1387772CF859" box="[199,297,1939,1963]" italics="true" pageId="3" pageNumber="104">Remarks</emphasis>
. All
<typeStatus id="54CA880B0012250B936F138677D5F858" box="[355,464,1938,1962]" pageId="3" pageNumber="104" type="holotype">holotypes</typeStatus>
are dissected wet specimens, whose shell was cracked to permit access to soft tissues. References to size are intended as relative to other congeneric species.
</paragraph>
<caption id="DF0E66210015250C929B148F734BFF40" box="[151,1358,155,179]" pageId="4" pageNumber="105">
<paragraph id="8BCE36A90015250C929B148F734BFF40" blockId="4.[151,1358,155,179]" box="[151,1358,155,179]" pageId="4" pageNumber="105">
<emphasis id="B905EABB0015250C929B148F770FFF41" bold="true" box="[151,266,155,179]" pageId="4" pageNumber="105">TABLE 1.</emphasis>
Shell dimensions (mm) and whorl counts of
<taxonomicName id="4C714D2A0015250C90FE148F75A0FF40" box="[754,933,155,178]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="genus">
<emphasis id="B905EABB0015250C90FE148F75A0FF40" box="[754,933,155,178]" italics="true" pageId="4" pageNumber="105">Australocosmica</emphasis>
</taxonomicName>
species recognised in the present work.
</paragraph>
</caption>
<paragraph id="8BCE36A90015250C929314DA7341FF1A" blockId="4.[159,1390,206,829]" box="[159,1348,206,232]" pageId="4" pageNumber="105">
<emphasis id="B905EABB0015250C929314DA7341FF1A" bold="true" box="[159,1348,206,232]" pageId="4" pageNumber="105">Species MS H D N</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36B65220015250C92A914EB7757FC7D" pageId="4" pageNumber="105" type="description">
<paragraph id="8BCE36A90015250C92A914EB736BFCCF" blockId="4.[159,1390,206,829]" pageId="4" pageNumber="105">
<taxonomicName id="4C714D2A0015250C92A914EB773AFEE6" box="[165,319,255,276]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="bernoullienis" status="sp. nov.">
<emphasis id="B905EABB0015250C92A914EB773AFEE6" box="[165,319,255,276]" italics="true" pageId="4" pageNumber="105">A. bernoullienis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C934814EB777DFEE6" box="[324,376,255,276]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 78) 46 10.0 12.2 12.9 16.1 4.5 5.2 (11.2 ± 0.6) (14.1 ± 0.8) (4.9 ± 0.2)
<taxonomicName id="4C714D2A0015250C92A915407728FE9B" box="[165,301,340,361]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="buffonensis" status="sp. nov.">
<emphasis id="B905EABB0015250C92A915407728FE9B" box="[165,301,340,361]" italics="true" pageId="4" pageNumber="105">A. buffonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C933F15407762FE9B" box="[307,359,340,361]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 77) 101 7.7 13.6 10.6 16.5 4.1 5.5 (10.2 ± 1.5) (13.2 ± 1.4) (4.7 ± 0.3)
<taxonomicName id="4C714D2A0015250C92A915BD773AFE4C" box="[165,319,425,446]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="crassicostata" status="sp. nov.">
<emphasis id="B905EABB0015250C92A915BD773AFE4C" box="[165,319,425,446]" italics="true" pageId="4" pageNumber="105">A. crassicostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C934815BD777DFE4C" box="[324,376,425,446]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 79) 62 10.1 13.8 13.0 15.9 4.6 5.4 (11.8 ± 0.7) (14.8 ± 0.5) (5.0 ± 0.2)
<taxonomicName id="4C714D2A0015250C92A915EA76F5FDE1" box="[165,240,510,531]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="nana" status="sp. nov.">
<emphasis id="B905EABB0015250C92A915EA76F5FDE1" box="[165,240,510,531]" italics="true" pageId="4" pageNumber="105">A. nana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C92FA15EA772FFDE1" box="[246,298,510,531]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 80) 40 7.5 9.9 12.0 13.7 4.0 4.7 (8.8 ± 0.5) (12.6 ± 0.4) (4.4 ± 0.2)
<taxonomicName id="4C714D2A0015250C92A916407701FD9B" box="[165,260,596,617]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="pallida" status="sp. nov.">
<emphasis id="B905EABB0015250C92A916407701FD9B" box="[165,260,596,617]" italics="true" pageId="4" pageNumber="105">A. pallida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C93061647773BFD9A" box="[266,318,595,616]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 76) 58 11.5 14.4 13.7 16.1 5.2 5.6 (12.8 ± 0.6) (14.8 ± 0.6) (5.4 ± 0.1)
<taxonomicName id="4C714D2A0015250C92A916BD7709FD4C" box="[165,268,681,702]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="105" phylum="Mollusca" rank="species" species="rotunda" status="sp. nov.">
<emphasis id="B905EABB0015250C92A916BD7709FD4C" box="[165,268,681,702]" italics="true" pageId="4" pageNumber="105">A. rotunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C00015250C931E16BC7743FD4F" box="[274,326,680,701]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
(NSP 40) 19 10.7 14.0 14.1 17.8 4.5 5.3 (11.8 ± 0.9) (15.2 ± 0.9) (4.8 ± 0.2) Total 326 7.514.4 10.617.8 4.05.6 (11.0 ± 1.6) (13.9 ± 1.3) (4.9 ± 0.4)
</paragraph>
<paragraph id="8BCE36A90015250C929B174E7757FC7D" blockId="4.[151,1436,858,911]" pageId="4" pageNumber="105">Given are: minmax (average ± standard deviation). Abbreviations: MS, number of measured shells; H, height, D, diameter, N, number of whorls.</paragraph>
</subSubSection>
</treatment>
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