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<mods:title>Additions To The Colombian Anacroneuria Fauna (Plecoptera: Perlidae) With Descriptions Of Seven New Species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zúñiga, Maria del Carmen</mods:namePart>
<mods:affiliation>Universidad del Valle, Departamento de Biología, Grupo de Investigaciones Entomológicas, Apartado Aéreo 25360, Cali, Colombia E-mail: maczuniga @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">maczuniga@gmail.com</mods:nameIdentifier>
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<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
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<mods:namePart>Cardona, William</mods:namePart>
<mods:affiliation>Fundación EcoAndina / WCS Colombia, Apartado Aéreo 25527, Cali, Colombia E-mail: williamcardona @ gmail. com</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tamaris-Turizo, César</mods:namePart>
<mods:affiliation>Universidad del Magdalena, Facultad de Ciencias Básicas, Programa de Biología, Grupo de Investigaciones en Limnología Neotropical, Santa Marta, Colombia E-mail: cesartamaris @ yahoo. es</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ortega, Oscar E.</mods:namePart>
<mods:affiliation>Universidad Nacional, Facultad de Ciencias, Escuela de Biociencias, Medellín, Colombia E-mail: oeortega @ unalmed. edu. co</mods:affiliation>
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<mods:title>Illiesia</mods:title>
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<mods:date>2007</mods:date>
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<taxonomicName authority="Zuniga &amp; Stark" authorityName="Zuniga &amp; Stark" authorityYear="1999" box="[868,1348,391,416]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="guambiana">
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<emphasis bold="true" box="[868,1159,391,416]" italics="true" pageId="13" pageNumber="140">Anacroneuria guambiana</emphasis>
Zúñiga &amp; Stark
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<taxonomicName authority="Zuniga &amp; Stark" authorityName="Zuniga &amp; Stark" authorityYear="1999" box="[849,1277,460,482]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="guambiana">
<emphasis box="[849,1095,460,482]" italics="true" pageId="13" pageNumber="140">Anacroneuria guambiana</emphasis>
Zúñiga &amp; Stark
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<emphasis box="[1288,1307,461,482]" italics="true" pageId="13" pageNumber="140">in</emphasis>
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.
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(MEUV)
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<emphasis box="[849,1098,554,576]" italics="true" pageId="13" pageNumber="140">Anacroneuria guambiana</emphasis>
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. Female description
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(
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<collectingMunicipality box="[1342,1404,860,884]" pageId="13" pageNumber="140">Same</collectingMunicipality>
data, except 
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<elevation box="[1208,1291,895,919]" metricMagnitude="3" metricUnit="m" metricValue="2.44" pageId="13" pageNumber="140" unit="m" value="2440.0">2440 m</elevation>
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29 
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26 
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, 
<date box="[1032,1200,929,953]" pageId="13" pageNumber="140" value="2007-03-12">
<collectingDate box="[1032,1200,929,953]" pageId="13" pageNumber="140" value="2007-03-12">12 March 2007</collectingDate>
</date>
(
<collectionCode box="[1216,1346,929,953]" pageId="13" pageNumber="140">MHN-ICN</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111187384" collectingDate="2007-07-03" collectionCode="MEUA, BPS, BYU" collectorName="Zuniga &amp; Cardona &amp; J. Chara &amp; L. P. Giraldo &amp; A. M. Chara" country="Colombia" location="Same" pageId="13" pageNumber="140" specimenCount="4" specimenCount-male="4">
<specimenCount box="[1365,1396,929,953]" pageId="13" pageNumber="140" type="male">4♂</specimenCount>
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<location LSID="urn:lsid:plazi:treatment:03D887A7FFFC2B49FCE5FE07F6A41333:8EAE606AFFFC2B49FCADFC4BF11B17B1" box="[812,874,964,988]" country="Colombia" name="Same" pageId="13" pageNumber="140">Same</location>
data, except 
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<collectingDate box="[1025,1152,964,988]" pageId="13" pageNumber="140" value="2007-07-03">3 July 2007</collectingDate>
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, M. del C. 
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, W. 
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. 
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42 
<emphasis bold="true" box="[1178,1184,1067,1091]" italics="true" pageId="13" pageNumber="140">′</emphasis>
35 
<emphasis bold="true" box="[1215,1226,1067,1091]" italics="true" pageId="13" pageNumber="140">″</emphasis>
</geoCoordinate>
, 
<geoCoordinate box="[1236,1380,1067,1091]" degrees="75" direction="west" minutes="30" orientation="longitude" pageId="13" pageNumber="140" precision="15" seconds="03" value="-75.50083">
W 75 
<emphasis bold="true" box="[1292,1302,1067,1091]" italics="true" pageId="13" pageNumber="140">°</emphasis>
30 
<emphasis bold="true" box="[1332,1338,1067,1091]" italics="true" pageId="13" pageNumber="140">′</emphasis>
03 
<emphasis bold="true" box="[1369,1380,1067,1091]" italics="true" pageId="13" pageNumber="140">″</emphasis>
</geoCoordinate>
, 
<date pageId="13" pageNumber="140" value="2007-07-05">
<collectingDate pageId="13" pageNumber="140" value="2007-07-05">5 July 2007</collectingDate>
</date>
(
<collectionCode box="[936,985,1102,1126]" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="13" pageNumber="140" type="Herbarium">BPS</collectionCode>
, 
<collectionCode box="[998,1056,1102,1126]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="13" pageNumber="140" type="University or college">BYU</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111187423" collectingDate="2007-07-15" collectingDateMax="2007-07-16" collectingDateMin="2007-07-15" collectionCode="MEUV" collectorName="C. Zuniga &amp; W. Cardona &amp; G. Zabala &amp; C. Cultid &amp; A. Laina" country="Colombia" county="Municipio de Santuario" elevation="2050" latitude="5.1272225" location="Rio San Rafael" longLatPrecision="21" longitude="-76.00222" municipality="Parque Municipal Natural Planes de San Rafael" pageId="13" pageNumber="140" specimenCount="1" specimenCount-female="1" stateProvince="Risaralda">
<specimenCount box="[1075,1107,1101,1126]" pageId="13" pageNumber="140" type="female">1♀</specimenCount>
, 
<collectingRegion box="[1123,1233,1102,1126]" country="Colombia" name="Risaralda" pageId="13" pageNumber="140">Risaralda</collectingRegion>
, 
<collectingCounty pageId="13" pageNumber="140">Municipio de Santuario</collectingCounty>
, 
<collectingMunicipality pageId="13" pageNumber="140">Parque Municipal Natural Planes de San Rafael</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03D887A7FFFC2B49FCE5FE07F6A41333:8EAE606AFFFC2B49FC03FB1CF65B10C6" box="[898,1066,1171,1195]" country="Colombia" county="Municipio de Santuario" latitude="5.1272225" longLatPrecision="21" longitude="-76.00222" municipality="Parque Municipal Natural Planes de San Rafael" name="Rio San Rafael" pageId="13" pageNumber="140" stateProvince="Risaralda">Río San Rafael</location>
middle basin, 
<quantity box="[1241,1325,1171,1195]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="13" pageNumber="140" unit="m" value="2050.0">
<elevation box="[1241,1325,1171,1195]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="13" pageNumber="140" unit="m" value="2050.0">2050 m</elevation>
</quantity>
, 
<geoCoordinate degrees="05" direction="north" minutes="07" orientation="latitude" pageId="13" pageNumber="140" precision="15" seconds="38" value="5.1272225">
N 05 
<emphasis bold="true" box="[1393,1403,1171,1195]" italics="true" pageId="13" pageNumber="140">°</emphasis>
07 
<emphasis bold="true" box="[838,844,1205,1229]" italics="true" pageId="13" pageNumber="140">′</emphasis>
38 
<emphasis bold="true" box="[878,889,1205,1229]" italics="true" pageId="13" pageNumber="140">″</emphasis>
</geoCoordinate>
, 
<geoCoordinate box="[902,1054,1205,1229]" degrees="76" direction="west" minutes="00" orientation="longitude" pageId="13" pageNumber="140" precision="15" seconds="08" value="-76.00222">
W 76 
<emphasis bold="true" box="[960,970,1205,1229]" italics="true" pageId="13" pageNumber="140">°</emphasis>
00 
<emphasis bold="true" box="[1003,1009,1205,1229]" italics="true" pageId="13" pageNumber="140">′</emphasis>
08 
<emphasis bold="true" box="[1043,1054,1205,1229]" italics="true" pageId="13" pageNumber="140">″</emphasis>
</geoCoordinate>
, 
<collectingMethod box="[1067,1175,1205,1229]" pageId="13" pageNumber="140">light trap</collectingMethod>
, 
<date box="[1187,1361,1205,1229]" pageId="13" pageNumber="140" value="2007-07-15" valueMax="2007-07-16" valueMin="2007-07-15">
<collectingDate box="[1187,1361,1205,1229]" pageId="13" pageNumber="140" value="2007-07-15" valueMax="2007-07-16" valueMin="2007-07-15">15-16 July 2007</collectingDate>
</date>
, M. del 
<collectorName box="[855,971,1240,1264]" pageId="13" pageNumber="140">C. Zúñiga</collectorName>
, 
<collectorName box="[982,1124,1240,1264]" pageId="13" pageNumber="140">W. Cardona</collectorName>
, 
<collectorName box="[1134,1247,1240,1264]" pageId="13" pageNumber="140">G. Zabala</collectorName>
, 
<collectorName box="[1259,1365,1240,1264]" pageId="13" pageNumber="140">C. Cultid</collectorName>
, 
<collectorName pageId="13" pageNumber="140">A. Laina</collectorName>
(
<collectionCode box="[890,974,1275,1299]" pageId="13" pageNumber="140">MEUV</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="140" type="discussion">
<paragraph blockId="13.[812,1404,652,1886]" pageId="13" pageNumber="140">
<emphasis bold="true" box="[812,948,1309,1333]" pageId="13" pageNumber="140">Discussion.</emphasis>
This series of 
<specimenCount box="[1101,1299,1309,1333]" pageId="13" pageNumber="140" type="generic">twelve specimens</specimenCount>
is similar to those reported earlier from this site by 
<bibRefCitation author="Stark, B. P. &amp; C. Zuniga" pageId="13" pageNumber="140" pagination="229 - 237" refId="ref13463" refString="Stark, B. P. &amp; M. del C. Zuniga. 2003. The Anacroneuria guambiana complex of South America (Plecoptera: Perlidae). pp 229 - 237. In: E. Gaino (Ed.). Research update on Ephemeroptera and Plecoptera. University of Perugia, Italy." type="book chapter" year="2003">Stark &amp; Zúñiga (2003)</bibRefCitation>
, however in this same collection from “La Pastora” are 
<specimenCount box="[1021,1157,1413,1437]" pageId="13" pageNumber="140" type="male">eight males</specimenCount>
whose head pattern matches that of 
<taxonomicName authority="Stark &amp; Zuniga" authorityName="Stark &amp; Zuniga" authorityYear="1999" box="[992,1275,1448,1472]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="socapa">
<emphasis box="[992,1092,1448,1471]" italics="true" pageId="13" pageNumber="140">A. socapa</emphasis>
Stark &amp; Zúñiga
</taxonomicName>
, but whose aedeagal features are like those of the males listed above. Four additional “intermediate” males were collected at another site, Natural National Park Los Nevados, El 
<collectingCountry box="[956,1033,1586,1610]" name="Jordan" pageId="13" pageNumber="140">Jordán</collectingCountry>
at 3100 meters, in Risaralda, and 
<specimenCount box="[812,947,1620,1644]" pageId="13" pageNumber="140" type="male">three males</specimenCount>
with the opposite condition (
<taxonomicName authorityName="Stark &amp; Zuniga" authorityYear="1999" box="[1300,1403,1621,1644]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="socapa">
<emphasis box="[1300,1403,1621,1644]" italics="true" pageId="13" pageNumber="140">A. socapa</emphasis>
</taxonomicName>
aedeagus but without head markings) were collected at two additional sites in Risaralda (reported under 
<taxonomicName authorityName="Stark &amp; Zuniga" authorityYear="1999" box="[812,919,1724,1748]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="socapa">
<emphasis box="[812,919,1724,1748]" italics="true" pageId="13" pageNumber="140">A. socapa</emphasis>
</taxonomicName>
). The “La Pastora” site specimens also include several typical males of 
<taxonomicName authorityName="Stark &amp; Zuniga" authorityYear="1999" box="[1223,1332,1759,1782]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="socapa">
<emphasis box="[1223,1332,1759,1782]" italics="true" pageId="13" pageNumber="140">A. socapa</emphasis>
</taxonomicName>
. This suggests the possibility of hybridization at sites in Risaralda, or, perhaps the head pattern character is less reliable than previously thought.
</paragraph>
</subSubSection>
</treatment>
</document>