treatments-xml/data/03/D8/87/03D887A7FFF32B46FF60F9D1F7261344.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4754807" ID-GBIF-Dataset="85ff6002-c514-44a2-997b-aae68c249ba3" ID-GBIF-Taxon="182067289" ID-ISSN="1854-0392" ID-Zenodo-Dep="4754807" checkinTime="1620844093025" checkinUser="felipe" docAuthor="Zúñiga, Maria del Carmen, Stark, Bill P., Cardona, William, Tamaris-Turizo, César &amp; Ortega, Oscar E." docDate="2007" docId="03D887A7FFF32B46FF60F9D1F7261344" docLanguage="en" docName="Illiesia.3.13.127-149.pdf" docOrigin="Illiesia 3 (13)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Anacroneuria otun Zúñiga &amp; Stark &amp; Cardona &amp; Tamaris-Turizo &amp; Ortega 2007, sp. nov." docType="treatment" docVersion="7" lastPageNumber="129" masterDocId="FFE1FFDFFFF12B44FF81FF8FF271146D" masterDocTitle="Additions To The Colombian Anacroneuria Fauna (Plecoptera: Perlidae) With Descriptions Of Seven New Species" masterLastPageNumber="149" masterPageNumber="127" pageId="2" pageNumber="129" updateTime="1632098486936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Additions To The Colombian Anacroneuria Fauna (Plecoptera: Perlidae) With Descriptions Of Seven New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zúñiga, Maria del Carmen</mods:namePart>
<mods:affiliation>Universidad del Valle, Departamento de Biología, Grupo de Investigaciones Entomológicas, Apartado Aéreo 25360, Cali, Colombia E-mail: maczuniga @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">maczuniga@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cardona, William</mods:namePart>
<mods:affiliation>Fundación EcoAndina / WCS Colombia, Apartado Aéreo 25527, Cali, Colombia E-mail: williamcardona @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">williamcardona@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tamaris-Turizo, César</mods:namePart>
<mods:affiliation>Universidad del Magdalena, Facultad de Ciencias Básicas, Programa de Biología, Grupo de Investigaciones en Limnología Neotropical, Santa Marta, Colombia E-mail: cesartamaris @ yahoo. es</mods:affiliation>
<mods:nameIdentifier type="email">cesartamaris@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortega, Oscar E.</mods:namePart>
<mods:affiliation>Universidad Nacional, Facultad de Ciencias, Escuela de Biociencias, Medellín, Colombia E-mail: oeortega @ unalmed. edu. co</mods:affiliation>
<mods:nameIdentifier type="email">oeortega@unalmed.edu.co</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2007</mods:date>
<mods:detail type="volume">
<mods:number>3</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>13</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>127</mods:start>
<mods:end>149</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4754665</mods:identifier>
<mods:identifier type="GBIF-Dataset">85ff6002-c514-44a2-997b-aae68c249ba3</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4754665</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4754807" ID-GBIF-Taxon="182067289" ID-Zenodo-Dep="4754807" LSID="urn:lsid:plazi:treatment:03D887A7FFF32B46FF60F9D1F7261344" httpUri="http://treatment.plazi.org/id/03D887A7FFF32B46FF60F9D1F7261344" lastPageNumber="129" pageId="2" pageNumber="129">
<subSubSection box="[225,733,1629,1654]" pageId="2" pageNumber="129" type="nomenclature">
<paragraph blockId="2.[225,733,1629,1689]" box="[225,733,1629,1654]" pageId="2" pageNumber="129">
<heading bold="true" box="[225,733,1629,1654]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="129" reason="2">
<emphasis bold="true" box="[225,733,1629,1654]" pageId="2" pageNumber="129">
<taxonomicName authority="Zúñiga &amp; Stark &amp; Cardona &amp; Tamaris-Turizo &amp; Ortega, 2007" authorityName="Zúñiga &amp; Stark &amp; Cardona &amp; Tamaris-Turizo &amp; Ortega" authorityYear="2007" box="[225,441,1630,1654]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="otun" status="sp. nov.">
<emphasis bold="true" box="[225,441,1630,1654]" italics="true" pageId="2" pageNumber="129">Anacroneuria otun</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[448,539,1630,1654]" pageId="2" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
, Stark &amp; Zúñiga
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[417,542,1665,1689]" pageId="2" pageNumber="129" type="description">
<paragraph blockId="2.[225,733,1629,1689]" box="[417,542,1665,1689]" pageId="2" pageNumber="129">
(
<figureCitation box="[425,534,1665,1689]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Figs. 6-10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="129" type="materials_examined">
<paragraph blockId="2.[184,776,1727,1959]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[184,415,1727,1751]" pageId="2" pageNumber="129">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111187391" collectingDate="2006-06-27" collectingDateMax="2006-06-28" collectingDateMin="2006-06-27" collectingMethod="light trap" collectionCode="MEUV, BPS" collectorName="C. Zuniga &amp; W. Cardona &amp; G. Zabala &amp; Zuniga" country="Colombia" county="Municipio de Pereira" elevation="3100" latitude="4.7088885" location="El Jordan" longLatPrecision="21" longitude="-75.49361" municipality="Parque Nacional Natural Los Nevados" pageId="2" pageNumber="129" specimenCount="6" specimenCount-male="4" stateProvince="Risaralda" typeStatus="holotype">
<typeStatus box="[423,531,1728,1752]" pageId="2" pageNumber="129">Holotype</typeStatus>
<specimenCount box="[540,559,1727,1751]" pageId="2" pageNumber="129" type="male">♂</specimenCount>
and 
<specimenCount box="[621,653,1727,1752]" pageId="2" pageNumber="129" type="male">3♂</specimenCount>
<typeStatus box="[662,775,1728,1752]" pageId="2" pageNumber="129">paratypes</typeStatus>
from 
<collectingCountry box="[247,393,1762,1786]" name="Colombia" pageId="2" pageNumber="129">COLOMBIA</collectingCountry>
, 
<collectingRegion box="[404,514,1762,1786]" country="Colombia" name="Risaralda" pageId="2" pageNumber="129">Risaralda</collectingRegion>
, 
<collectingCounty box="[526,772,1762,1786]" pageId="2" pageNumber="129">Municipio de Pereira</collectingCounty>
, 
<collectingMunicipality box="[184,615,1797,1821]" pageId="2" pageNumber="129">Parque Nacional Natural Los Nevados</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03D887A7FFF32B46FF60F9D1F7261344:8EAE606AFFF32B46FDF1F88AF0A91370" box="[624,728,1797,1821]" country="Colombia" county="Municipio de Pereira" latitude="4.7088885" longLatPrecision="21" longitude="-75.49361" municipality="Parque Nacional Natural Los Nevados" name="El Jordan" pageId="2" pageNumber="129" stateProvince="Risaralda">El Jordán</location>
, 
<location LSID="urn:lsid:plazi:treatment:03D887A7FFF32B46FF60F9D1F7261344:8EAE606AFFF32B46FD60F88AF3CB1352" country="Colombia" county="Municipio de Pereira" latitude="4.7088885" longLatPrecision="21" longitude="-75.49361" municipality="Parque Nacional Natural Los Nevados" name="Rio Otun headwater basin" pageId="2" pageNumber="129" stateProvince="Risaralda">Río Otún headwater basin</location>
, 
<quantity box="[453,536,1832,1856]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="129" unit="m" value="3100.0">
<elevation box="[453,536,1832,1856]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="129" unit="m" value="3100.0">3100 m</elevation>
</quantity>
, 
<geoCoordinate box="[547,694,1831,1856]" degrees="04" direction="north" minutes="42" orientation="latitude" pageId="2" pageNumber="129" precision="15" seconds="32" value="4.7088885">
N 04 
<emphasis bold="true" box="[601,611,1831,1855]" italics="true" pageId="2" pageNumber="129">°</emphasis>
42 
<emphasis bold="true" box="[644,650,1831,1855]" italics="true" pageId="2" pageNumber="129">′</emphasis>
32 
<emphasis bold="true" box="[683,694,1831,1855]" italics="true" pageId="2" pageNumber="129">″</emphasis>
</geoCoordinate>
, 
<geoCoordinate degrees="75" direction="west" minutes="29" orientation="longitude" pageId="2" pageNumber="129" precision="15" seconds="37" value="-75.49361">
W 75 
<emphasis bold="true" box="[765,775,1831,1855]" italics="true" pageId="2" pageNumber="129">°</emphasis>
29 
<emphasis bold="true" box="[210,216,1866,1890]" italics="true" pageId="2" pageNumber="129">′</emphasis>
37 
<emphasis bold="true" box="[248,259,1866,1890]" italics="true" pageId="2" pageNumber="129">″</emphasis>
</geoCoordinate>
, 
<date box="[271,449,1866,1890]" pageId="2" pageNumber="129" value="2006-06-27" valueMax="2006-06-28" valueMin="2006-06-27">
<collectingDate box="[271,449,1866,1890]" pageId="2" pageNumber="129" value="2006-06-27" valueMax="2006-06-28" valueMin="2006-06-27">27-28 June 2006</collectingDate>
</date>
, 
<collectingMethod box="[459,567,1866,1890]" pageId="2" pageNumber="129">light trap</collectingMethod>
, M. del 
<collectorName box="[655,771,1866,1890]" pageId="2" pageNumber="129">C. Zúñiga</collectorName>
, 
<collectorName box="[184,324,1901,1925]" pageId="2" pageNumber="129">W. Cardona</collectorName>
, 
<collectorName box="[335,447,1901,1925]" pageId="2" pageNumber="129">G. Zabala</collectorName>
, Cardozo- 
<collectorName box="[570,653,1901,1925]" pageId="2" pageNumber="129">Zúñiga</collectorName>
(Holotype and 
<specimenCount box="[234,370,1935,1959]" pageId="2" pageNumber="129" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
: 
<collectionCode box="[380,461,1935,1958]" pageId="2" pageNumber="129">MEUV</collectionCode>
, paratype: 
<collectionCode box="[587,636,1935,1959]" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="2" pageNumber="129" type="Herbarium">BPS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="2.[812,1405,288,1834]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,983,288,312]" pageId="2" pageNumber="129">Adult habitus.</emphasis>
General color dark brown patterned with yellow. Head covered mostly with dark brown but M-line with three prominent pale areas; occiput dusky brown except for pale post- ocellar area. Pronotum brown with narrow median pale band and scattered dark rugosities (
<figureCitation box="[1117,1185,460,485]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Fig. 6</figureCitation>
). Wing membrane transparent, veins dark brown. Legs brown, tibia darker than femora, except narrow, dark, apical femoral band.
</paragraph>
<paragraph blockId="2.[812,1405,288,1834]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,879,599,623]" pageId="2" pageNumber="129">Male.</emphasis>
Forewing length 
<quantity box="[1078,1159,599,623]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="129" unit="mm" value="18.0">18 mm</quantity>
. Hammer cylindrical, height greater than diameter (
<figureCitation box="[1152,1216,633,658]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Fig. 7</figureCitation>
). Aedeagal apex short, slightly longer than wide, and rounded; venter with a pair of small membranous lobes, shoulders broad (
<figureCitation box="[894,952,737,761]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Fig 8</figureCitation>
). Dorsal keel steep, composed of a pair of narrow ridges which end slightly beyond mid length of apical area (
<figureCitation box="[1075,1158,806,830]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Fig. 10</figureCitation>
). Membranous areas dorsad and basal to hooks bulging. Lateral aspect of apex about as wide as long (
<figureCitation box="[1134,1193,875,900]" captionStart="Figs" captionStartId="3.[184,234,1427,1451]" captionTargetBox="[333,1221,295,1369]" captionTargetId="figure-250@3.[353,1235,279,1381]" captionTargetPageId="3" captionText="Figs. 6-10. Anacroneuria otun. 6. Head and pronotum, 7. Male sternum 9, 8. Aedeagus, ventral, 9. Aedeagus, lateral, 10. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4754669" httpUri="https://zenodo.org/record/4754669/files/figure.png" pageId="2" pageNumber="129">Fig 9</figureCitation>
).
</paragraph>
<paragraph blockId="2.[812,1405,288,1834]" box="[812,1032,910,934]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,904,910,934]" pageId="2" pageNumber="129">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="2.[812,1405,288,1834]" box="[812,1014,945,969]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,885,945,969]" pageId="2" pageNumber="129">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="129" type="etymology">
<paragraph blockId="2.[812,1405,288,1834]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,946,980,1004]" pageId="2" pageNumber="129">Etymology.</emphasis>
The species name, used as a noun in apposition, is based on Río Otún, the 
<typeStatus box="[1241,1292,1015,1039]" pageId="2" pageNumber="129">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="129" type="diagnosis">
<paragraph blockId="2.[812,1405,288,1834]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,938,1049,1073]" pageId="2" pageNumber="129">Diagnosis.</emphasis>
The aedeagus of this species is most similar to that of 
<taxonomicName authority="Zuniga &amp; Vimos" authorityName="Zuniga &amp; Vimos" authorityYear="2006" box="[1004,1282,1084,1108]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="cajas">
<emphasis box="[1004,1086,1084,1108]" italics="true" pageId="2" pageNumber="129">A. cajas</emphasis>
Zúñiga &amp; Vimos
</taxonomicName>
, a recently described Ecuadorian species (
<bibRefCitation author="Zuniga, M. &amp; B. P. Stark &amp; J. J. Vasconez &amp; F. Bersosa &amp; D. Vimos" box="[1175,1393,1118,1142]" pageId="2" pageNumber="129" refId="ref13666" refString="Zuniga, M. del C., B. P. Stark., J. J. Vasconez., F. Bersosa. &amp; D. Vimos. 2006. Colombian and Ecuadorian Anacroneuria (Plecoptera: Perlidae): New species, records and life stages. Studies on" type="book" year="2006">Zúñiga et al. 2006</bibRefCitation>
). The aedeagal apex of 
<taxonomicName authorityName="Zuniga &amp; Vimos" authorityYear="2006" box="[1054,1135,1153,1177]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="cajas">
<emphasis box="[1054,1135,1153,1177]" italics="true" pageId="2" pageNumber="129">A. cajas</emphasis>
</taxonomicName>
is more pointed and the dorsal keel is significantly weaker than in 
<emphasis box="[1288,1374,1188,1211]" italics="true" pageId="2" pageNumber="129">
<taxonomicName authorityName="Zúñiga &amp; Stark &amp; Cardona &amp; Tamaris-Turizo &amp; Ortega" authorityYear="2007" box="[1288,1371,1188,1211]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="otun">A. otun</taxonomicName>
.
</emphasis>
In addition, the pronotum of 
<taxonomicName authorityName="Zuniga &amp; Vimos" authorityYear="2006" box="[1115,1197,1222,1246]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="cajas">
<emphasis box="[1115,1197,1222,1246]" italics="true" pageId="2" pageNumber="129">A. cajas</emphasis>
</taxonomicName>
has dark pigment adjacent to the median suture, whereas in 
<taxonomicName authorityName="Zúñiga &amp; Stark &amp; Cardona &amp; Tamaris-Turizo &amp; Ortega" authorityYear="2007" box="[1302,1383,1257,1280]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="otun">
<emphasis box="[1302,1383,1257,1280]" italics="true" pageId="2" pageNumber="129">A. otun</emphasis>
</taxonomicName>
a narrow, median pale band occurs.
</paragraph>
<paragraph blockId="2.[812,1405,288,1834]" pageId="2" pageNumber="129">
<emphasis bold="true" box="[812,1012,1326,1350]" pageId="2" pageNumber="129">Ecological notes.</emphasis>
The 
<typeStatus box="[1076,1127,1326,1350]" pageId="2" pageNumber="129">type</typeStatus>
locality is in the headwaters of Rio Otún in the highest mountains of the Cordillera Central and is part of Región Natural Andina de 
<collectingCountry box="[935,1049,1430,1454]" name="Colombia" pageId="2" pageNumber="129">Colombia</collectingCountry>
. This small second order stream is about 
<quantity box="[910,952,1464,1488]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="129" unit="m" value="8.0">8 m</quantity>
wide and 
<quantity box="[1074,1177,1464,1488]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="129" unit="m" value="0.45" valueMax="0.5" valueMin="0.4">0.4-0.5 m</quantity>
deep, with a mixed substrate composed of boulders, stones, patches of gravel and sands, and riffle zones primarily. The water quality is in optimum environmental condition, the temperature was 10°C and the air, 12°C at the time of collection. Natural riparian vegetation is present, the zone receives 
<quantity box="[1295,1404,1672,1696]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="129" unit="mm" value="1900.0">1900 mm</quantity>
mean annual rainfall and is covered by a very humid montane forest, in accordance with Holdridge life zones system (
<bibRefCitation author="Espinal, L. S. &amp; E. Montenegro" box="[984,1316,1775,1800]" pageId="2" pageNumber="129" refId="ref13193" refString="Espinal, L. S. &amp; E. Montenegro. 1963. Formaciones vegetales de Colombia. Memoria explicativa del mapa ecologico. Instituto Geografico Agustin Codazzi. 201 pp." type="book" year="1963">
Espinal &amp; 
<collectingCountry box="[1110,1251,1776,1800]" name="Serbia" pageId="2" pageNumber="129">Montenegro</collectingCountry>
1963
</bibRefCitation>
). These specimens were collected during the dry season.
</paragraph>
</subSubSection>
</treatment>
</document>