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<document id="B04465EDCD4085F95B763F9B448AC2D8" ID-CLB-Dataset="299055" ID-DOI="10.5281/zenodo.11067355" ID-GBIF-Dataset="67ec2de4-1816-4969-8132-f6796d3afa06" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067355" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052343267" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2021" docId="03AF0B0CFFF0FF86FF9FFEA1A586FD97" docLanguage="en" docName="ProcCalifAcadSci.67.4.85-182.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (4)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Broscosoma parvum Kavanaugh and Liang 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="136" masterDocId="FF967374FFDFFFB5FFE4FFCFA106FFEE" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae)." masterLastPageNumber="182" masterPageNumber="85" pageNumber="132" updateTime="1719540814554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="667694B0EC1D5E27CA76127E86344FC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BAF321EF9BF430F16B88C68B53A2C4D9">
<mods:title id="85E0058934F632F1C7D95E0ADA3ED29C">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Broscini (Coleoptera: Carabidae).</mods:title>
</mods:titleInfo>
<mods:name id="9A09C105633524530DDE63BB4FB9023C" type="personal">
<mods:role id="A286982CE9F06D7CDF69FD5250B99AE0">
<mods:roleTerm id="541B19FD80AEE7A0D6AA2A4FEC42497F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="045A3E548E1F2B5F9474014C2CF8BBEF">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="1AFCB21150EFB6AC70352718417E0B22">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="3C14A8B21AFF5A87B0700C407D96A3B3" type="personal">
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<mods:roleTerm id="67B48C8AFA5F85E1F83FFF99DF8F63BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="309425E2FDBD265D0B2032D2B0A58B2E">Liang, Hongbin</mods:namePart>
<mods:affiliation id="D05BD0E6925365C3ED0BF6826F92C560">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="921C26F3650BEB99B47330C298B6B5A6">text</mods:typeOfResource>
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<mods:titleInfo id="EEE34898D4CEFD2D55BAD3111D3DF3ED">
<mods:title id="C2061C803F5FAB892E879E0AFF00968B">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="4446895BCDAB54155482E66B7C8B4F8B">
<mods:date id="796AEFA56999904129EEAD076E34A182">2021</mods:date>
<mods:detail id="1D41C29FD7A72BDC3A57C9A4F828E001" type="pubDate">
<mods:number id="0814BA8D9CAB71D39301DB5C219509C2">2021-09-30</mods:number>
</mods:detail>
<mods:detail id="96EB14D27A7F2B8664655CF947B93609" type="volume">
<mods:number id="0DBA3A70CB9560C5E30C036601E239EA">67</mods:number>
</mods:detail>
<mods:detail id="CBE6FAA85194DDAE24437596DBB1554B" type="issue">
<mods:number id="15B5C4A49F8C230701693E6AE3390ADF">4</mods:number>
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<mods:start id="429834E5FE096A3EB1D72504764D7085">85</mods:start>
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<mods:classification id="5B332B2168D375C69684446A2FDA197A">journal article</mods:classification>
<mods:identifier id="283BA60BF0B91B1F1D2F551CE04F4F2A" type="CLB-Dataset">299055</mods:identifier>
<mods:identifier id="59FAA5E8C1187563F6ADA57E3F65A6BE" type="DOI">10.5281/zenodo.11067355</mods:identifier>
<mods:identifier id="A82CA5B56460EC924BAFBC979F2EA331" type="GBIF-Dataset">67ec2de4-1816-4969-8132-f6796d3afa06</mods:identifier>
<mods:identifier id="DFCD465F368041AFBAED82B6EE3B604F" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="92DCE05D2815A5B8C3552F0424341BCC" type="Zenodo-Dep">11067355</mods:identifier>
</mods:mods>
<treatment id="03AF0B0CFFF0FF86FF9FFEA1A586FD97" ID-DOI="http://doi.org/10.5281/zenodo.12575578" ID-GBIF-Taxon="233814473" ID-Zenodo-Dep="12575578" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97" httpUri="http://treatment.plazi.org/id/03AF0B0CFFF0FF86FF9FFEA1A586FD97" lastPageId="51" lastPageNumber="136" pageId="47" pageNumber="132">
<subSubSection id="C31CE991FFF0FF9AFF9FFEA1A3E2FE68" box="[123,740,366,390]" pageId="47" pageNumber="132" type="nomenclature">
<paragraph id="8BB9BA1AFFF0FF9AFF9FFEA1A3E2FE68" blockId="47.[123,740,366,420]" box="[123,740,366,390]" pageId="47" pageNumber="132">
<heading id="D0F10D76FFF0FF9AFF9FFEA1A3E2FE68" box="[123,740,366,390]" fontSize="10" level="2" pageId="47" pageNumber="132" reason="7">
<emphasis id="B9726608FFF0FF9AFF9FFEA1A3E2FE68" bold="true" box="[123,740,366,390]" pageId="47" pageNumber="132">
9.
<taxonomicName id="4C06C199FFF0FF9AFF72FEA1A387FE68" authority="Kavanaugh and Liang" authorityName="Kavanaugh and Liang" authorityYear="2021" box="[150,641,366,390]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="parvum" status="sp. nov.">
<emphasis id="B9726608FFF0FF9AFF72FEA1A07EFE6B" bold="true" box="[150,376,366,389]" italics="true" pageId="47" pageNumber="132">Broscosoma parvum</emphasis>
Kavanaugh and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF0FF9AFD6AFEA1A3E2FE68" box="[654,740,366,390]" pageId="47" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF0FF9AFF9FFE41A09CFE4A" box="[123,410,398,420]" pageId="47" pageNumber="132" type="description">
<paragraph id="8BB9BA1AFFF0FF9AFF9FFE41A09CFE4A" blockId="47.[123,740,366,420]" box="[123,410,398,420]" pageId="47" pageNumber="132">
<figureCitation id="133DA69FFFF0FF9AFF9FFE41A1DBFE4A" box="[123,221,398,420]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="47" pageNumber="132">Figures 6f</figureCitation>
,
<figureCitation id="133DA69FFFF0FF9AFF0EFE41A004FE4A" box="[234,258,398,420]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="47" pageNumber="132">26</figureCitation>
,
<figureCitation id="133DA69FFFF0FF9AFEEAFE41A020FE4D" box="[270,294,398,419]" captionStart="FIGURE" captionStartId="49.[171,183,963,982]" captionTargetBox="[144,1168,235,949]" captionTargetId="figure-1710@49.[120,1176,216,957]" captionTargetPageId="49" captionText="FIGURE 27. Male genitalia, Broscosoma parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067409" httpUri="https://zenodo.org/record/11067409/files/figure.png" pageId="47" pageNumber="132">27</figureCitation>
,
<figureCitation id="133DA69FFFF0FF9AFED7FE41A050FE4D" box="[307,342,398,420]" captionStart="FIGURE" captionStartId="85.[171,183,1590,1609]" captionTargetBox="[148,1148,216,1572]" captionTargetId="figure-67@85.[148,1148,216,1572]" captionTargetPageId="85" captionText="FIGURE 45. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. Snowmelt stream on north-facing slope above North Fork of Yamu He, 8.5 km W of Shibali, 3100-3200 m, 27.18315º/098.71921º, Lumadeng Township, Fugong County, Yunnan (habitat for Broscosoma parvum sp. nov.); b. 0.6 km N of Dizhengdang village on Dulongjiang, 1880 m, 28.084º/098.327º, Dulongjiang Township, Gongshan County, Yunnan (habitat for Broscosoma purpureum sp. nov.). Photos by David H. Kavanaugh" figureDoi="http://doi.org/10.5281/zenodo.11067449" httpUri="https://zenodo.org/record/11067449/files/figure.png" pageId="47" pageNumber="132">45a</figureCitation>
,
<figureCitation id="133DA69FFFF0FF9AFE86FE41A09CFE4A" box="[354,410,398,420]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="88.[171,183,995,1014]" captionStartId-1="88.[184,196,1682,1701]" captionStartId-2="89.[171,183,1461,1480]" captionTargetBox-0="[130,1164,233,969]" captionTargetBox-1="[130,1162,1063,1660]" captionTargetBox-2="[140,1154,249,1428]" captionTargetId-0="figure-57@88.[120,1176,215,989]" captionTargetId-1="figure-332@88.[120,1176,1054,1668]" captionTargetId-2="figure-68@89.[120,1176,216,1448]" captionTargetPageId-0="88" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." captionText-2="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi-0="http://doi.org/10.5281/zenodo.11067455" figureDoi-1="http://doi.org/10.5281/zenodo.11067457" figureDoi-2="http://doi.org/10.5281/zenodo.11067459" httpUri-0="https://zenodo.org/record/11067455/files/figure.png" httpUri-1="https://zenodo.org/record/11067457/files/figure.png" httpUri-2="https://zenodo.org/record/11067459/files/figure.png" pageId="47" pageNumber="132">48-50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF0FF9AFF4FFE78A01CFB21" pageId="47" pageNumber="132" type="materials_examined">
<paragraph id="8BB9BA1AFFF0FF9AFF4FFE78A39DFBC1" blockId="47.[123,1174,439,1712]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FFE78A054FE21" bold="true" box="[171,338,439,463]" pageId="47" pageNumber="132">Type material.</emphasis>
<materialsCitation id="3B6EB047FFF0FF9AFEBEFE78A037FDA1" ID-GBIF-Occurrence="4899905513" collectingDate="2004-05-09" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCode="male, CASENT 1022788" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54BD04B8FFF0FF9AFEBEFE78A0BBFE21" box="[346,445,439,463]" pageId="47" pageNumber="132" type="holotype">Holotype</typeStatus>
, a
<specimenCode id="DBA01261FFF0FF9AFE05FE78A313FE21" box="[481,533,439,463]" pageId="47" pageNumber="132">male</specimenCode>
, in
<collectionCode id="ED1722DFFFF0FF9AFDA5FE78A36BFE21" box="[577,621,439,463]" pageId="47" pageNumber="132">IOZ</collectionCode>
, labeled: “
<specimenCode id="DBA01261FFF0FF9AFD02FE78A2B5FE21" box="[742,947,439,463]" pageId="47" pageNumber="132">CASENT 1022788</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF0FF9AFC3FFE78A529FE21" box="[987,1071,439,463]" name="China" pageId="47" pageNumber="132">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF0FF9AFBD9FE77A588FE21" box="[1085,1166,440,463]" country="China" name="Yunnan" pageId="47" pageNumber="132">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF0FF9AFF9FFE17A026FE01" box="[123,288,471,496]" pageId="47" pageNumber="132">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF0FF9AFECAFE18A316FE01" box="[302,528,471,495]" pageId="47" pageNumber="132">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A0BA338FFF0FF9AFDFAFE18A271FE01" box="[542,887,471,495]" location="Shibali on Shibali" pageId="47" pageNumber="132">
<quantity id="4CFE17FFFFF0FF9AFDFAFE18A36EFE01" box="[542,616,471,495]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="47" pageNumber="132" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali
</locationDeviation>
road,
<location id="8ED9ECC1FFF0FF9AFC5FFE18A19CFDE1" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFF0FF9AFC5FFE18A19CFDE1" country="China" county="Fugong County" latitude="27.18326" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" name="North Fork of Yamu He" pageId="47" pageNumber="132" stateProvince="Yunnan">North Fork of Yamu He</location>
” / “
<geoCoordinate id="EE32DCDDFFF0FF9AFF2CFE37A041FDE1" box="[200,327,503,527]" degrees="27. 18326" direction="north" orientation="latitude" pageId="47" pageNumber="132" precision="1" value="27.18326">N27.18326º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF0FF9AFEABFE37A0DFFDE1" box="[335,473,503,527]" degrees="098. 72002" direction="east" orientation="longitude" pageId="47" pageNumber="132" precision="1" value="98.72002">E098.72002º</geoCoordinate>
,
<quantity id="4CFE17FFFFF0FF9AFE03FE38A337FDE1" box="[487,561,503,527]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">
<elevation id="002B5D29FFF0FF9AFE03FE38A337FDE1" box="[487,561,503,527]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">3100m</elevation>
</quantity>
,
<date id="FFB89CDAFFF0FF9AFDDBFE38A3C4FDE1" box="[575,706,503,528]" pageId="47" pageNumber="132" value="2004-05-09">
<collectingDate id="EFFC6532FFF0FF9AFDDBFE38A3C4FDE1" box="[575,706,503,528]" pageId="47" pageNumber="132" value="2004-05-09">9 May 2004</collectingDate>
</date>
, Stop #DHK2004-042,
<collectorName id="26F3DFCCFFF0FF9AFC20FE37A57FFDE1" box="[964,1145,503,527]" pageId="47" pageNumber="132">D.H. Kavanaugh</collectorName>
&amp;
<collectorName id="26F3DFCCFFF0FF9AFF9FFDD7A1F4FDC1" box="[123,242,535,559]" pageId="47" pageNumber="132">H.B. Liang</collectorName>
collectors”/ HOLOTYPE
<taxonomicName id="4C06C199FFF0FF9AFDEAFDD7A2C2FDC1" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[526,964,535,560]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="furvum" status="sp. nov.">
<emphasis id="B9726608FFF0FF9AFDEAFDD7A3E2FDC1" box="[526,740,535,559]" italics="true" pageId="47" pageNumber="132">Broscosoma furvum</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFF0FF9AFC2EFDD7A51BFDC1" box="[970,1053,536,559]" pageId="47" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [red label]
</materialsCitation>
.
<materialsCitation id="3B6EB047FFF0FF9AFEDBFDF7A0A7FD81" ID-GBIF-Occurrence="4899905448" collectingDate="2004-05-09" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCode="CASENT 1022789" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54BD04B8FFF0FF9AFEDBFDF7A0A1FDBE" box="[319,423,568,592]" pageId="47" pageNumber="132" type="paratype">Paratypes</typeStatus>
(a total of 6):
<specimenCount id="9D007193FFF0FF9AFDA5FDF7A3BEFDA1" box="[577,696,567,591]" count="1" pageId="47" pageNumber="132" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFF0FF9AFD23FDF8A3F5FDA1" box="[711,755,567,591]" pageId="47" pageNumber="132">IOZ</collectionCode>
) labeled: same as holotype except first label “
<specimenCode id="DBA01261FFF0FF9AFF21FD98A092FD81" box="[197,404,599,623]" pageId="47" pageNumber="132">CASENT 1022789</specimenCode>
”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF0FF9AFE57FD97A1E8FD21" ID-GBIF-Occurrence="4899905494" collectingDate="2004-05-07" collectionCode="CAS" collectorName=". H. Kavanaugh" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCode="CASENT 1023771" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF0FF9AFE57FD97A31EFD81" box="[435,536,599,623]" count="1" pageId="47" pageNumber="132" type="male">one male</specimenCount>
(
<collectionCode id="ED1722DFFFF0FF9AFDC8FD98A366FD81" box="[556,608,599,623]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="132" type="Museum">CAS</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF0FF9AFD3BFD98A2A8FD81" box="[735,942,599,623]" pageId="47" pageNumber="132">CASENT 1023771</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF0FF9AFC3CFD98A52AFD81" box="[984,1068,599,623]" name="China" pageId="47" pageNumber="132">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF0FF9AFBD9FD97A588FD81" box="[1085,1166,600,623]" country="China" name="Yunnan" pageId="47" pageNumber="132">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF0FF9AFF9FFDB7A026FD61" box="[123,288,631,656]" pageId="47" pageNumber="132">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF0FF9AFECAFDB8A316FD61" box="[302,528,631,655]" pageId="47" pageNumber="132">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A0BA338FFF0FF9AFDFAFDB8A271FD61" box="[542,887,631,655]" location="Shibali on Shibali" pageId="47" pageNumber="132">
<quantity id="4CFE17FFFFF0FF9AFDFAFDB8A36EFD61" box="[542,616,631,655]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="47" pageNumber="132" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali
</locationDeviation>
road,
<location id="8ED9ECC1FFF0FF9AFC5FFDB8A19CFD41" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFF0FF9AFC5FFDB8A19CFD41" country="China" county="Fugong County" latitude="27.18326" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" name="North Fork of Yamu He" pageId="47" pageNumber="132" stateProvince="Yunnan">North Fork of Yamu He</location>
” / “
<geoCoordinate id="EE32DCDDFFF0FF9AFF2EFD57A04FFD41" box="[202,329,663,687]" degrees="27. 18326" direction="north" orientation="latitude" pageId="47" pageNumber="132" precision="1" value="27.18326">N27.18326º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF0FF9AFEB5FD57A0DDFD41" box="[337,475,663,687]" degrees="098. 72002" direction="east" orientation="longitude" pageId="47" pageNumber="132" precision="1" value="98.72002">E098.72002º</geoCoordinate>
,
<quantity id="4CFE17FFFFF0FF9AFE0EFD58A332FD41" box="[490,564,663,687]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">
<elevation id="002B5D29FFF0FF9AFE0EFD58A332FD41" box="[490,564,663,687]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">3100m</elevation>
</quantity>
,
<date id="FFB89CDAFFF0FF9AFDA0FD57A3CEFD41" box="[580,712,663,688]" pageId="47" pageNumber="132" value="2004-05-07">
<collectingDate id="EFFC6532FFF0FF9AFDA0FD57A3CEFD41" box="[580,712,663,688]" pageId="47" pageNumber="132" value="2004-05-07">7 May 2004</collectingDate>
</date>
, Stop #DHK2004-038B, D
<collectorName id="26F3DFCCFFF0FF9AFC17FD57A593FD41" box="[1011,1173,663,687]" pageId="47" pageNumber="132">.H. Kavanaugh</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF0FF9AFF1BFD77A51CFD21" ID-GBIF-Occurrence="4899905478" box="[255,1050,695,719]" collectingDate="2005-08-16" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang" country="China" county="Fugong County" elevation="3150" latitude="27.18315" location="North Fork Yamu He" longLatPrecision="1" longitude="98.71921" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCode="CASENT 1019759" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF0FF9AFF1BFD77A064FD21" box="[255,354,695,719]" count="1" pageId="47" pageNumber="132" type="male">one male</specimenCount>
and
<specimenCount id="9D007193FFF0FF9AFE7FFD77A315FD21" box="[411,531,695,719]" count="1" pageId="47" pageNumber="132" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFF0FF9AFDC1FD78A35FFD21" box="[549,601,695,719]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="132" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1722DFFFF0FF9AFD8CFD78A392FD21" box="[616,660,695,719]" pageId="47" pageNumber="132">IOZ</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF0FF9AFCF4FD78A2DBFD21" box="[784,989,695,719]" pageId="47" pageNumber="132">CASENT 1019759</specimenCode>
” and
</materialsCitation>
<materialsCitation id="3B6EB047FFF0FF9AFBC6FD78A558FCC1" ID-GBIF-Occurrence="4899905309" collectingDate="2005-08-16" collectionCode="CAS, IOZ" collectorName="D. H. Kavanaugh &amp; H. B. Liang" country="China" county="Fugong County" elevation="3150" latitude="27.18315" location="North Fork Yamu He" longLatPrecision="1" longitude="98.71921" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCode="CASENT 1019754&quot;" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
“
<specimenCode id="DBA01261FFF0FF9AFBCAFD78A1E2FD01" pageId="47" pageNumber="132">CASENT 1019754”</specimenCode>
, respectively “
<collectingCountry id="F311FA8AFFF0FF9AFE68FD18A0E6FD01" box="[396,480,727,751]" name="China" pageId="47" pageNumber="132">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF0FF9AFE15FD17A344FD01" box="[497,578,728,751]" country="China" name="Yunnan" pageId="47" pageNumber="132">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF0FF9AFDB0FD17A3FBFD01" box="[596,765,727,752]" pageId="47" pageNumber="132">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF0FF9AFCEAFD18A2F5FD01" box="[782,1011,727,751]" pageId="47" pageNumber="132">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A0BA338FFF0FF9AFBE0FD18A038FCE1" location="Shibali on Shibali" pageId="47" pageNumber="132">
<quantity id="4CFE17FFFFF0FF9AFBE0FD18A554FD01" box="[1028,1106,727,751]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="47" pageNumber="132" unit="km" value="8.5">8.5 km</quantity>
W of Shibali on Shibali
</locationDeviation>
road, south bank of
<location id="8ED9ECC1FFF0FF9AFDC5FD38A201FCE1" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFF0FF9AFDC5FD38A201FCE1" box="[545,775,759,783]" country="China" county="Fugong County" latitude="27.18315" longLatPrecision="1" longitude="98.71921" municipality="Lumadeng Township" name="North Fork Yamu He" pageId="47" pageNumber="132" stateProvince="Yunnan">North Fork Yamu He</location>
” / “
<geoCoordinate id="EE32DCDDFFF0FF9AFCD3FD37A2B1FCE1" box="[823,951,759,783]" degrees="27. 18315" direction="north" orientation="latitude" pageId="47" pageNumber="132" precision="1" value="27.18315">N27.18315º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF0FF9AFC5AFD37A54EFCE1" box="[958,1096,759,783]" degrees="098. 71921" direction="east" orientation="longitude" pageId="47" pageNumber="132" precision="1" value="98.71921">E098.71921º</geoCoordinate>
,
<quantity id="4CFE17FFFFF0FF9AFBB3FD38A1C3FCC1" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="47" pageNumber="132" unit="m" value="3150.0" valueMax="3200.0" valueMin="3100.0">
<elevation id="002B5D29FFF0FF9AFBB3FD38A1C3FCC1" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="47" pageNumber="132" unit="m" value="3150.0" valueMax="3200.0" valueMin="3100.0">3100- 3200m</elevation>
</quantity>
,
<date id="FFB89CDAFFF0FF9AFF31FCD8A084FCC1" box="[213,386,791,816]" pageId="47" pageNumber="132" value="2005-08-16">
<collectingDate id="EFFC6532FFF0FF9AFF31FCD8A084FCC1" box="[213,386,791,816]" pageId="47" pageNumber="132" value="2005-08-16">16 August 2005</collectingDate>
</date>
, Stop #DHK2005-090,
<collectorName id="26F3DFCCFFF0FF9AFD6EFCD7A246FCC1" box="[650,832,791,815]" pageId="47" pageNumber="132">D.H. Kavanaugh</collectorName>
&amp;
<collectorName id="26F3DFCCFFF0FF9AFC8CFCD7A2E7FCC1" box="[872,993,791,815]" pageId="47" pageNumber="132">H.B. Liang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF0FF9AFB8BFCD7A588FC61" ID-GBIF-Occurrence="4899905308" collectingDate="2005-08-08" collectionCode="CAS" collectorName="H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="North Fork Yamu He" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" pageId="47" pageNumber="132" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF0FF9AFB8BFCD7A1C2FCA1" count="1" pageId="47" pageNumber="132" type="female">one female</specimenCount>
(
<collectionCode id="ED1722DFFFF0FF9AFF30FCF8A00EFCA1" box="[212,264,823,847]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="132" type="Museum">CAS</collectionCode>
), “
<collectingCountry id="F311FA8AFFF0FF9AFEC8FCF8A086FCA1" box="[300,384,823,847]" name="China" pageId="47" pageNumber="132">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF0FF9AFE6AFCF7A0D9FCA1" box="[398,479,824,847]" country="China" name="Yunnan" pageId="47" pageNumber="132">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF0FF9AFE0AFCF7A393FCA1" box="[494,661,823,848]" pageId="47" pageNumber="132">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BDD2060FFF0FF9AFD47FCF8A280FCA1" box="[675,902,823,847]" pageId="47" pageNumber="132">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A0BA338FFF0FF9AFC71FCF8A1C1FC81" location="Shibali on Shibali" pageId="47" pageNumber="132">
<quantity id="4CFE17FFFFF0FF9AFC71FCF8A2E6FCA1" box="[917,992,823,847]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="47" pageNumber="132" unit="km" value="8.5">8.5 km</quantity>
W of Shibali on Shibali
</locationDeviation>
road, south bank of
<location id="8ED9ECC1FFF0FF9AFE4CFC98A38BFC81" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFF0FF9AFE4CFC98A38BFC81" box="[424,653,855,879]" country="China" county="Fugong County" latitude="27.18326" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" name="North Fork Yamu He" pageId="47" pageNumber="132" stateProvince="Yunnan">North Fork Yamu He</location>
,
<geoCoordinate id="EE32DCDDFFF0FF9AFD78FC97A21DFC81" box="[668,795,855,879]" degrees="27. 18326" direction="north" orientation="latitude" pageId="47" pageNumber="132" precision="1" value="27.18326">N27.18326º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF0FF9AFCC7FC97A2AAFC81" box="[803,940,855,879]" degrees="098. 72002" direction="east" orientation="longitude" pageId="47" pageNumber="132" precision="1" value="98.72002">E098.72002º</geoCoordinate>
” / “
<quantity id="4CFE17FFFFF0FF9AFC38FC98A520FC81" box="[988,1062,855,879]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">
<elevation id="002B5D29FFF0FF9AFC38FC98A520FC81" box="[988,1062,855,879]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="47" pageNumber="132" unit="m" value="3100.0">3100m</elevation>
</quantity>
,
<date id="FFB89CDAFFF0FF9AFBD0FC98A1B6FC61" pageId="47" pageNumber="132" value="2005-08-08">
<collectingDate id="EFFC6532FFF0FF9AFBD0FC98A1B6FC61" pageId="47" pageNumber="132" value="2005-08-08">8 August 2005</collectingDate>
</date>
, Stop #DHK2005-067B, D.H. Kavanaugh &amp;
<collectorName id="26F3DFCCFFF0FF9AFD76FCB7A20EFC61" box="[658,776,887,911]" pageId="47" pageNumber="132">H.B. Liang</collectorName>
,
<collectorName id="26F3DFCCFFF0FF9AFCF0FCB7A280FC7E" box="[788,902,888,912]" pageId="47" pageNumber="132">D.Z. Dong</collectorName>
&amp;
<collectorName id="26F3DFCCFFF0FF9AFC42FCB7A513FC61" box="[934,1045,887,911]" pageId="47" pageNumber="132">J.F. Zhang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B6EB047FFF0FF9AFF9FFC57A1D3FBE1" ID-GBIF-Occurrence="4899905486" collectingDate="2004-05-06" collectionCode="CAS" collectorName="D. H. Kavanaugh &amp; C. E. Griswold &amp; Liang H. - B. &amp; Li X. - Y. &amp; Zhu B. - X." country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Lishadi Township" longLatPrecision="1" longitude="98.71763" pageId="47" pageNumber="132" specimenCode="CASENT1023565" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D007193FFF0FF9AFF9FFC57A1DBFC41" box="[123,221,919,943]" count="1" pageId="47" pageNumber="132" type="male">one male</specimenCount>
(
<collectionCode id="ED1722DFFFF0FF9AFF09FC58A027FC41" box="[237,289,919,943]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="132" type="Museum">CAS</collectionCode>
) labeled: “
<specimenCode id="DBA01261FFF0FF9AFE7DFC58A35BFC41" box="[409,605,919,943]" pageId="47" pageNumber="132">CASENT1023565</specimenCode>
”/ “
<collectingCountry id="F311FA8AFFF0FF9AFD60FC58A3DEFC41" box="[644,728,919,943]" name="China" pageId="47" pageNumber="132">CHINA</collectingCountry>
,
<collectingRegion id="49C274F8FFF0FF9AFD02FC57A231FC41" box="[742,823,920,943]" country="China" name="Yunnan" pageId="47" pageNumber="132">Yunnan</collectingRegion>
,
<collectingCounty id="62D8C296FFF0FF9AFCA1FC57A2EDFC41" box="[837,1003,919,944]" pageId="47" pageNumber="132">Fugong County</collectingCounty>
,
<location id="8ED9ECC1FFF0FF9AFC1DFC58A1A1FC21" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFF0FF9AFC1DFC58A1A1FC21" country="China" county="Fugong County" latitude="27.20676" longLatPrecision="1" longitude="98.71763" name="Lishadi Township" pageId="47" pageNumber="132" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A0BA338FFF0FF9AFF5CFC78A32CFC21" box="[184,554,951,975]" location="Shibali on Shibali" pageId="47" pageNumber="132">
<quantity id="4CFE17FFFFF0FF9AFF5CFC78A017FC21" box="[184,273,951,975]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="47" pageNumber="132" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Shibali
</locationDeviation>
road,
<geoCoordinate id="EE32DCDDFFF0FF9AFD97FC77A3F5FC21" box="[627,755,951,975]" degrees="27. 20676" direction="north" orientation="latitude" pageId="47" pageNumber="132" precision="1" value="27.20676">N27.20676º</geoCoordinate>
/
<geoCoordinate id="EE32DCDDFFF0FF9AFD1EFC77A282FC21" box="[762,900,951,975]" degrees="098. 71763" direction="east" orientation="longitude" pageId="47" pageNumber="132" precision="1" value="98.71763">E098.71763º</geoCoordinate>
”/ “
<quantity id="4CFE17FFFFF0FF9AFC49FC78A2F1FC21" box="[941,1015,951,975]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="47" pageNumber="132" unit="m" value="3290.0">
<elevation id="002B5D29FFF0FF9AFC49FC78A2F1FC21" box="[941,1015,951,975]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="47" pageNumber="132" unit="m" value="3290.0">3290m</elevation>
</quantity>
,
<date id="FFB89CDAFFF0FF9AFBECFC78A588FC21" box="[1032,1166,951,976]" pageId="47" pageNumber="132" value="2004-05-06">
<collectingDate id="EFFC6532FFF0FF9AFBECFC78A588FC21" box="[1032,1166,951,976]" pageId="47" pageNumber="132" value="2004-05-06">6 May 2004</collectingDate>
</date>
, Stop #DHK-2004-036,
<collectorName id="26F3DFCCFFF0FF9AFE9CFC17A32AFC01" box="[376,556,983,1007]" pageId="47" pageNumber="132">D.H. Kavanaugh</collectorName>
,
<collectorName id="26F3DFCCFFF0FF9AFDDFFC18A3DAFC01" box="[571,732,983,1007]" pageId="47" pageNumber="132">C. E. Griswold</collectorName>
,
<collectorName id="26F3DFCCFFF0FF9AFD0FFC18A26AFC01" box="[747,876,983,1007]" pageId="47" pageNumber="132">Liang H.-B.</collectorName>
,
<collectorName id="26F3DFCCFFF0FF9AFC9EFC18A2D5FC01" box="[890,979,983,1007]" pageId="47" pageNumber="132">Li X.-Y.</collectorName>
&amp;
<collectorName id="26F3DFCCFFF0FF9AFC13FC18A563FC01" box="[1015,1125,983,1007]" pageId="47" pageNumber="132">Zhu B.-X.</collectorName>
collectors”.
</materialsCitation>
All paratypes also bear the following label: “PARATYPE
<taxonomicName id="4C06C199FFF0FF9AFCD8FC37A1D3FBC1" authority="Kavanaugh &amp; Liang" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="parvum" status="sp. nov.">
<emphasis id="B9726608FFF0FF9AFCD8FC37A510FBE1" box="[828,1046,1016,1039]" italics="true" pageId="47" pageNumber="132">Broscosoma parvum</emphasis>
Kavanaugh &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A241DB73FFF0FF9AFF07FBD7A030FBC1" box="[227,310,1048,1071]" pageId="47" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
designated 2021” [yellow label].
</paragraph>
<paragraph id="8BB9BA1AFFF0FF9AFF4FFBF8A25FFB81" blockId="47.[123,1174,439,1712]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FFBF8A044FBA1" bold="true" box="[171,322,1079,1103]" pageId="47" pageNumber="132">Type locality.</emphasis>
China, Yunnan, Fugong County, Lumadeng Township, 8.5 km W of Shibali on Shibali road, North Fork of Yamu He, 27.18326º/98.72002º, 3100m.
</paragraph>
<paragraph id="8BB9BA1AFFF0FF9AFF4FFBB8A01CFB21" blockId="47.[123,1174,439,1712]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FFBB8A0ECFB7E" bold="true" box="[171,490,1143,1168]" pageId="47" pageNumber="132">Derivation of species name.</emphasis>
The species epithet,
<taxonomicName id="4C06C199FFF0FF9AFD31FBB7A220FB61" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[725,806,1144,1167]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="132" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFF0FF9AFD31FBB7A220FB61" box="[725,806,1144,1167]" italics="true" pageId="47" pageNumber="132">parvum</emphasis>
</taxonomicName>
, is an adjective derived from the Latin word,
<emphasis id="B9726608FFF0FF9AFF1BFB57A04EFB41" box="[255,328,1176,1199]" italics="true" pageId="47" pageNumber="132">parvus</emphasis>
, meaning small. The name refers to the relatively small body size of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF0FF9AFF4FFB18A19AF9E1" pageId="47" pageNumber="132" type="diagnosis">
<paragraph id="8BB9BA1AFFF0FF9AFF4FFB18A19AF9E1" blockId="47.[123,1174,439,1712]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FFB18A019FB01" bold="true" box="[171,287,1239,1263]" pageId="47" pageNumber="132">Diagnosis.</emphasis>
Adults of this species (
<figureCitation id="133DA69FFFF0FF9AFDC5FB18A37CFB01" box="[545,634,1239,1263]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="47" pageNumber="132">Fig. 26a</figureCitation>
) can be distinguished from those of other species in the region by the following combination of character states: size small, BL = 7.5 to 8.
<quantity id="4CFE17FFFFF0FF9AFBFAFB38A55DFAE1" box="[1054,1115,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="132" unit="mm" value="5.0">5 mm</quantity>
; dorsum with distinct metallic reflection, green on head, green or blue-green on pronotum, blue-green on elytra; antennomeres 3 and 4 glabrous except for apical whorl of setae; eyes (
<figureCitation id="133DA69FFFF0FF9AFC3BFAF8A521FAA1" box="[991,1063,1335,1359]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="47" pageNumber="132">Fig. 6f</figureCitation>
) less convex, slightly flattened in some specimens; pronotum with basolateral setae absent; elytral silhouette ovoid, with greatest width at or near midlength, with humeri rounded yet slightly evident, two or more elytral striae impressed, all striae punctate with punctures diminishing in size and depth laterally and posteriorly, medial two or more elytral intervals very slightly to moderately convex; male protarsi with tarsomeres 1 to 3 with ventral pads of adhesive setae; male genitalia as in
<figureCitation id="133DA69FFFF0FF9AFB8FFA18A190F9E1" captionStart="FIGURE" captionStartId="49.[171,183,963,982]" captionTargetBox="[144,1168,235,949]" captionTargetId="figure-1710@49.[120,1176,216,957]" captionTargetPageId="49" captionText="FIGURE 27. Male genitalia, Broscosoma parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067409" httpUri="https://zenodo.org/record/11067409/files/figure.png" pageId="47" pageNumber="132">Fig. 27</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFF0FF87FF4FF9D8A0E3FCD9" lastPageId="50" lastPageNumber="135" pageId="47" pageNumber="132" type="description">
<paragraph id="8BB9BA1AFFF0FF9AFF4FF9D8A2D0F941" blockId="47.[123,1174,439,1712]" pageId="47" pageNumber="132">
<emphasis id="B9726608FFF0FF9AFF4FF9D8A030F9C1" bold="true" box="[171,310,1559,1583]" pageId="47" pageNumber="132">Description.</emphasis>
<figureCitation id="133DA69FFFF0FF9AFED9F9D8A093F9C1" box="[317,405,1559,1583]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="47" pageNumber="132">Fig. 26a</figureCitation>
. Size small, BL male = 7.5-7.
<quantity id="4CFE17FFFFF0FF9AFD3DF9D7A211F9C1" box="[729,791,1560,1583]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="47" pageNumber="132" unit="mm" value="7.0">7 mm</quantity>
, female = 7.9-
<quantity id="4CFE17FFFFF0FF9AFC52F9D8A50EF9C1" box="[950,1032,1559,1583]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="47" pageNumber="132" unit="mm" value="8.5">8.5 mm</quantity>
, ratio EL/PL male 2.2-2.4, female 2.3. Body color black or piceous, last visible addominal sternite dark brown or rufous brown, antennae with all antennomeres rufous brown, femora black or dark piceous, tibiae and tarsi rufous brown. Dorsum with distinct metallic reflection, green on head, green or blue-green on pronotum, and blue-green on elytra, venter without metallic reflection.
</paragraph>
<caption id="DF79EA92FFEFFF85FF4FF9FCA3E1F997" ID-DOI="http://doi.org/10.5281/zenodo.11067407" ID-Zenodo-Dep="11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="48" pageNumber="133" startId="48.[171,183,1587,1606]" targetBox="[132,1176,216,1571]" targetPageId="48" targetType="figure">
<paragraph id="8BB9BA1AFFEFFF85FF4FF9FCA3E1F997" blockId="48.[123,1173,1586,1657]" pageId="48" pageNumber="133">
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26.
<taxonomicName id="4C06C199FFEFFF85FEEBF9FCA0B9F9AB" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[271,447,1587,1605]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="133" phylum="Arthropoda" rank="species" species="parvum" status="sp. nov.">
<emphasis id="B9726608FFEFFF85FEEBF9FCA0B9F9AB" box="[271,447,1587,1605]" italics="true" pageId="48" pageNumber="133">Broscosoma parvum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFEFFF85FE22F9FCA30FF9A8" box="[454,521,1587,1606]" pageId="48" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<caption id="DF79EA92FFEEFF84FF4FFC0CA3D1FBE4" ID-DOI="http://doi.org/10.5281/zenodo.11067409" ID-Zenodo-Dep="11067409" httpUri="https://zenodo.org/record/11067409/files/figure.png" pageId="49" pageNumber="134" startId="49.[171,183,963,982]" targetBox="[144,1168,235,949]" targetPageId="49" targetType="figure">
<paragraph id="8BB9BA1AFFEEFF84FF4FFC0CA3D1FBE4" blockId="49.[123,1173,963,1034]" pageId="49" pageNumber="134">
<smallCapsWord id="8D5F2CC6FFEEFF84FF4FFC0CA1EEFC3B" baselines="977,977" box="[171,232,963,982]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="49" pageNumber="134">FIGURE</smallCapsWord>
27. Male genitalia,
<taxonomicName id="4C06C199FFEEFF84FE73FC0BA341FC38" authority="Kavanaugh &amp; Liang, 2021" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[407,583,964,982]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="134" phylum="Arthropoda" rank="species" species="parvum" status="sp. nov.">
<emphasis id="B9726608FFEEFF84FE73FC0BA341FC38" box="[407,583,964,982]" italics="true" pageId="49" pageNumber="134">Broscosoma parvum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241DB73FFEEFF84FDAAFC0CA397FC38" box="[590,657,963,982]" pageId="49" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
(CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB9BA1AFFEEFF84FF4FFBFDA582FA87" blockId="49.[123,1174,1074,1706]" pageId="49" pageNumber="134">
Head.
<figureCitation id="133DA69FFFEEFF84FF15FBFDA03FFBA4" box="[241,313,1074,1098]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="49" pageNumber="134">
Fig.
<specimenCount id="9D007193FFEEFF84FEC7FBFDA046FBA7" box="[291,320,1074,1098]" count="6" pageId="49" pageNumber="134" type="generic">6f.</specimenCount>
</figureCitation>
Eyes only slightly convex, slightly flattened, small, their diameter slightly less than length of tempora. Frontal furrows deeply impressed, narrow, linear, slightly divergent posteriorly, medium length, extended posteriorly to middle of eyes, smooth or faintly rugulose, impunctate. Vertex with post-temporal transverse sulcus deep, sharply defined, moderately punctate. Tempora straight, oblique. Genal ridges absent or restricted to head region distinctly posterior to post-temporal constriction. Clypeus with one pair of setae. Supraorbital setae present, one pair. Eustipes of maxilla with two setae, the dorsobasal seta distinctly more than half as long as ventrobasal seta.
<collectingRegion id="49C274F8FFEEFF84FF9FFADDA1D3FAC7" box="[123,213,1298,1321]" country="Myanmar" name="Chin" pageId="49" pageNumber="134">Mentum</collectingRegion>
with tooth present, simple, paramedial region deeply foveate, one pair of mental setae present. Glosal sclerite (ligula) with one pair of setae. Submentum with two or three pairs of setae. Gula without transverse grooves. Antennomeres 3 and 4 without pubescence, ratio A3/A5 = 1.2.
</paragraph>
<paragraph id="8BB9BA1AFFEEFF84FF4FFABDA336F987" blockId="49.[123,1174,1074,1706]" pageId="49" pageNumber="134">
Pronotum.
<figureCitation id="133DA69FFFEEFF84FEC7FABDA06AFA64" box="[291,364,1394,1418]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="49" pageNumber="134">
Fig.
<specimenCount id="9D007193FFEEFF84FEB2FABDA075FA67" box="[342,371,1394,1418]" count="6" pageId="49" pageNumber="134" type="generic">6f.</specimenCount>
</figureCitation>
Disc anterior to sub-basal constriction slightly narrow, slightly longitudinally ovoid in dorsal view with base distinctly pedunculate, widest at middle of discal region, ratio PL/PW = 1.2–1.3, disk markedly convex; apical, lateral, and basal margination absent; anterior transverse impression shallow, broad and vaguely delineated, impunctate or sparsely and coarsely punctate; median longitudinal impression moderately impressed; posterior transverse impression deep, sharply delineated; anterior region smooth; pronotal base coarsely and sparsely to densely punctate; one pair of midlateral pronotal setae inserted slightly anterior to mid-length of discal region; basolateral pronotal setae absent.
</paragraph>
<paragraph id="8BB9BA1AFFEEFF87FF4FF9BDA289FDBB" blockId="49.[123,1174,1074,1706]" lastBlockId="50.[123,1174,219,1726]" lastPageId="50" lastPageNumber="135" pageId="49" pageNumber="134">Elytra. Elytral silhouette subovoid, slightly narrowed in basal half, widest at or near middle, ratio EL/EW male and female = 1.6, greatest elevation above lateral margin (in lateral view) posterior to middle; humeri faintly present, roundly obtuse; lateral margins moderately arcuate and nearly straight in anterior half sub-basally; parascutellar striole present, short, distally merged in a distinct jog with free base of stria 1. Eight striae present; stria 1 moderately deeply and sharply impressed, striae 2 to 3, 4, or 5 shallowly impressed in basal two-thirds of striae and successively more shallowly impressed to indistinct in apical third, striae 3 to 8 successively more shallowly impressed to indistinct or obliterated laterally and apically, striae 8 merged with lateral groove throughout its length or nearly so; stria 1 coarsely and moderately densely punctate at least in basal half, striae 2 to 3, 4, or 5 moderately punctate in basal two-thirds and successively more shallowly and sparsely punctate in apical third, striae 3 to 8 successively more shallowly and sparsely punctate laterally and apically. Elytral microsculpture comprised of very faintly impressed, nearly effaced isodiametric sculpticells. Parascutellar seta present at base of stria 2, discal setae absent, umbilicate series comprised of one post-humeral and two preapical setae.</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFD91A3D8FD9B" blockId="50.[123,1174,219,1726]" box="[171,734,606,630]" pageId="50" pageNumber="135">Hindwings. Vestigial, incapable of supporting flight.</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFDB1A213FD7B" blockId="50.[123,1174,219,1726]" box="[171,789,638,662]" pageId="50" pageNumber="135">Thoracic venter. Metepisternum ratio ML/MW = 1.8–2.2.</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFD50A2E8FD18" blockId="50.[123,1174,219,1726]" pageId="50" pageNumber="135">Legs. Hind trochanters asetose. Tarsomeres without dorsal pubescence; tarsomere 5 with two or more pairs of setae ventrally; male protarsi with pads of adhesive setae on tarsomeres 1 to 3 ventrally, male mesotarsi with pads of adhesive setae on tarsomeres 1 and 2 ventrally.</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFD30A555FCF8" blockId="50.[123,1174,219,1726]" box="[171,1107,767,791]" pageId="50" pageNumber="135">Abdomen. Sternite VI of both male and female with one pair of apicoparamedial setae.</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFCD0A0E3FCD9" blockId="50.[123,1174,219,1726]" box="[171,485,799,823]" pageId="50" pageNumber="135">
Male genitalia. As in
<figureCitation id="133DA69FFFEDFF87FE76FCD0A0D9FCD9" box="[402,479,799,823]" captionStart="FIGURE" captionStartId="49.[171,183,963,982]" captionTargetBox="[144,1168,235,949]" captionTargetId="figure-1710@49.[120,1176,216,957]" captionTargetPageId="49" captionText="FIGURE 27. Male genitalia, Broscosoma parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067409" httpUri="https://zenodo.org/record/11067409/files/figure.png" pageId="50" pageNumber="135">Fig. 27</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CE991FFEDFF86FF4FFCF0A586FD97" lastPageId="51" lastPageNumber="136" pageId="50" pageNumber="135" type="discussion">
<paragraph id="8BB9BA1AFFEDFF87FF4FFCF0A1C2FA95" blockId="50.[123,1174,219,1726]" pageId="50" pageNumber="135">
<emphasis id="B9726608FFEDFF87FF4FFCF0A02BFCB9" bold="true" box="[171,301,831,855]" pageId="50" pageNumber="135">Comments.</emphasis>
Only four species in the study area have the full dorsum of the body with green metallic reflection. Members of
<taxonomicName id="4C06C199FFEDFF87FE3CFCAFA36EFC99" box="[472,616,863,887]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="danzhuens">
<emphasis id="B9726608FFEDFF87FE3CFCAFA36EFC99" box="[472,616,863,887]" italics="true" pageId="50" pageNumber="135">B. danzhuens</emphasis>
</taxonomicName>
e are larger in size, have eyes that are more convex (
<figureCitation id="133DA69FFFEDFF87FF60FC4FA1C8FC79" box="[132,206,896,920]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="50" pageNumber="135">Fig. 6e</figureCitation>
), the elytral silhouette slightly broader across the humeri and their greatest width distinctly posterior to midlength (at or very near midlength in
<taxonomicName id="4C06C199FFEDFF87FD21FC6EA232FC56" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[709,820,929,952]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD21FC6EA232FC56" box="[709,820,929,952]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), elytra striae that are more finely punctate, and elytral intervals that are flat (at least some intervals slightly convex in
<taxonomicName id="4C06C199FFEDFF87FBF3FC0EA583FC36" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[1047,1157,961,984]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FBF3FC0EA583FC36" box="[1047,1157,961,984]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
). The
<typeStatus id="54BD04B8FFEDFF87FF4FFC2FA00EFC16" box="[171,264,992,1016]" pageId="50" pageNumber="135" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C06C199FFEDFF87FEC8FC2EA0A5FC16" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[300,419,992,1016]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="resbecqi">
<emphasis id="B9726608FFEDFF87FEC8FC2EA0A5FC16" box="[300,419,992,1016]" italics="true" pageId="50" pageNumber="135">B. resbecqi</emphasis>
</taxonomicName>
is larger and has the pronotum with the discal portion relatively shorter and more globose (
<figureCitation id="133DA69FFFEDFF87FE85FBCEA0ADFBF7" box="[353,427,1025,1049]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="50" pageNumber="135">Fig. 6d</figureCitation>
) and lateral margination present and nearly complete (
<figureCitation id="133DA69FFFEDFF87FC0BFBCEA53FFBF7" box="[1007,1081,1025,1049]" captionStart="FIGURE" captionStartId="12.[171,183,1195,1214]" captionTargetBox="[120,1176,216,1175]" captionTargetId="figure-55@12.[120,1176,216,1175]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax, Broscosoma species, oblique left lateral aspect. a. a. B. holomarginatum sp. nov. (CASENT1016746; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. purpureum sp. nov. (CASENT1015871; Moqie Wang, Dulongjiang Township, Gongshan Yunnan, China). White arrows denote end points of lateral margination." figureDoi="http://doi.org/10.5281/zenodo.11067369" httpUri="https://zenodo.org/record/11067369/files/figure.png" pageId="50" pageNumber="135">Fig. 9b</figureCitation>
) (absent from
<taxonomicName id="4C06C199FFEDFF87FF53FBEDA021FBD7" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[183,295,1058,1081]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FF53FBEDA021FBD7" box="[183,295,1058,1081]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), the femora pale rufous (darker in
<taxonomicName id="4C06C199FFEDFF87FD4AFBEDA218FBD7" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[686,798,1058,1081]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD4AFBEDA218FBD7" box="[686,798,1058,1081]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), and the median lobe of the male genitalia much thicker (
<figureCitation id="133DA69FFFEDFF87FE93FB8EA0C8FBB7" box="[375,462,1089,1113]" captionStart="FIGURE" captionStartId="58.[171,183,951,970]" captionTargetBox="[133,1161,228,923]" captionTargetId="figure-1524@58.[120,1176,216,932]" captionTargetPageId="58" captionText="FIGURE 31. Male genitalia, Broscosoma resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China). a. Right paramere, lateral aspect; b. Left paramere, lateral aspect; c. Median lobe, left lateral aspect; d. Median lobe, ventral aspect; scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067419" httpUri="https://zenodo.org/record/11067419/files/figure.png" pageId="50" pageNumber="135">Fig. 31c</figureCitation>
) and with the apical lamella short and tapered to a narrowly rounded tip in lateral aspect (median lobe much thinner and the apical lamella slightly hooked dorsally in
<taxonomicName id="4C06C199FFEDFF87FF73FB4DA000FB77" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[151,262,1154,1177]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FF73FB4DA000FB77" box="[151,262,1154,1177]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
). Members of
<taxonomicName id="4C06C199FFEDFF87FE4AFB4DA30BFB74" authorityName="Putzeys. We" authorityYear="1877" box="[430,525,1154,1178]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="ribbei">
<emphasis id="B9726608FFEDFF87FE4AFB4DA30BFB74" box="[430,525,1154,1178]" italics="true" pageId="50" pageNumber="135">B. ribbei</emphasis>
</taxonomicName>
are generally much larger and have the eyes distinctly more convex, the pronotum with the discal portion relatively shorter and more globose (
<figureCitation id="133DA69FFFEDFF87FC0EFB6DA533FB54" box="[1002,1077,1186,1210]" captionStart="FIGURE" captionStartId="9.[171,183,1467,1486]" captionTargetBox="[120,1176,216,1447]" captionTargetId="figure-66@9.[120,1176,216,1447]" captionTargetPageId="9" captionText="FIGURE 6. Forebody, Broscosoma species, dorsal aspect. a. B. holomarginatum sp. nov. (CASENT1015162; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); b. B. ribbei Putzeys (CASENT1025337; 0.5 km WSW of Maku village, Dulongjiang Township, Gongshan County, Yunnan, China); c. B. purpureum sp. nov. (CASENT1015338; 0.5 km N of Kongdang, Dulongjiang Township, Gongshan County, Yunnan, China); d. B. resbecqi sp. nov. (Holotype; Siran Wang, Dulongjiang Township, Gongshan County, Yunnan, China); e. B. danzhuense sp. nov. (Holotype; Danzhum He drainage, Cikai Township, Gongshan County, Yunnan, China); f. B. parvum sp. nov. (CASENT1023565; 11.5 km above Shibali, Lishadi Township, Fugong County, Yunnan, China)." figureDoi="http://doi.org/10.5281/zenodo.11067363" httpUri="https://zenodo.org/record/11067363/files/figure.png" pageId="50" pageNumber="135">Fig. 6b</figureCitation>
) and lateral margination incomplete but present in most specimens, at least in the vicinity of the midlateral setae and more extensively in some specimens, elytral microsculpture distinct, deeply to moderately impressed (very faintly impressed or effaced in
<taxonomicName id="4C06C199FFEDFF87FD2DFACCA23DFAF4" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[713,827,1283,1306]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD2DFACCA23DFAF4" box="[713,827,1283,1306]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), the metatrochanters unisetose (unilaterally asetose in a few specimens) (asetose in
<taxonomicName id="4C06C199FFEDFF87FD56FAECA224FAD4" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[690,802,1315,1338]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD56FAECA224FAD4" box="[690,802,1315,1338]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), and the median lobe of the male genitalia much longer, thinner, and with the apical lamella more symmetrically rounded in lateral aspect.
</paragraph>
<paragraph id="8BB9BA1AFFEDFF87FF4FFA4BA1DAF953" blockId="50.[123,1174,219,1726]" pageId="50" pageNumber="135">
As noted in the Comments section for
<taxonomicName id="4C06C199FFEDFF87FDB2FA4BA3F4FA75" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[598,754,1411,1435]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="danzhuense">
<emphasis id="B9726608FFEDFF87FDB2FA4BA3F4FA75" box="[598,754,1411,1435]" italics="true" pageId="50" pageNumber="135">B. danzhuense</emphasis>
</taxonomicName>
, only two
<taxonomicName id="4C06C199FFEDFF87FC8FFA4BA542FA75" box="[875,1092,1411,1435]" pageId="50" pageNumber="135">
<emphasis id="B9726608FFEDFF87FC8FFA4BA2E8FA75" box="[875,1006,1412,1435]" italics="true" pageId="50" pageNumber="135">Broscosoma</emphasis>
species
</taxonomicName>
known from outside the study area have members in which the elytral humeri are at least faintly evident (although in both they are less evident than in
<taxonomicName id="4C06C199FFEDFF87FD95FA0AA3E6FA32" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[625,736,1477,1500]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD95FA0AA3E6FA32" box="[625,736,1477,1500]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
) and the entire dorsum exhibits metallic green or bluish-green reflection. In members of
<taxonomicName id="4C06C199FFEDFF87FD60FA2AA216FA12" authorityName="Deuve" authorityYear="2006" box="[644,784,1508,1532]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="montreuili">
<emphasis id="B9726608FFEDFF87FD60FA2AA216FA12" box="[644,784,1508,1532]" italics="true" pageId="50" pageNumber="135">B. montreuili</emphasis>
</taxonomicName>
, the forebody is a darker, less vivid metallic green than the elytra (both parts equally vivid or forebody lighter and more vivid in
<taxonomicName id="4C06C199FFEDFF87FB9AF9CAA1CAF9D2" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FB9AF9CAA1CAF9D2" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), and in both
<taxonomicName id="4C06C199FFEDFF87FE86F9EAA0E8F9D2" authorityName="Deuve" authorityYear="2006" box="[354,494,1572,1596]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="montreuili">
<emphasis id="B9726608FFEDFF87FE86F9EAA0E8F9D2" box="[354,494,1572,1596]" italics="true" pageId="50" pageNumber="135">B. montreuili</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06C199FFEDFF87FDC7F9EAA374F9D2" authorityName="Deuve" authorityYear="2006" box="[547,626,1573,1596]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="tiani">
<emphasis id="B9726608FFEDFF87FDC7F9EAA374F9D2" box="[547,626,1573,1596]" italics="true" pageId="50" pageNumber="135">B. tiani</emphasis>
</taxonomicName>
body size is larger, the discal portion of the pronotum is more globose (narrower and more elongate in
<taxonomicName id="4C06C199FFEDFF87FD55F98AA226F9B2" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[689,800,1605,1628]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FD55F98AA226F9B2" box="[689,800,1605,1628]" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
), the elytra have striae 2 to 8 more faintly defined and finely punctate, and the median lobe of the male genitalia is longer, less arcuate and with the apical lamella either rounded apically or hooked ventrally (hooked dorsally in
<taxonomicName id="4C06C199FFEDFF87FB9AF949A1CAF953" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="135" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B9726608FFEDFF87FB9AF949A1CAF953" italics="true" pageId="50" pageNumber="135">B. parvum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB9BA1AFFECFF86FF4FFF14A024FE9B" blockId="51.[123,1174,219,634]" pageId="51" pageNumber="136">
<emphasis id="B9726608FFECFF86FF4FFF14A09EFF1D" bold="true" box="[171,408,219,243]" pageId="51" pageNumber="136">Habitat distribution.</emphasis>
Members of this species have been found under stones along small to medium-sized streams on moist organic substrate. Half of the specimens collected were found along a small snowmelt stream on a steep, north-facing slope (
<figureCitation id="133DA69FFFECFF86FCC1FED3A278FEDA" box="[805,894,284,308]" captionStart="FIGURE" captionStartId="85.[171,183,1590,1609]" captionTargetBox="[148,1148,216,1572]" captionTargetId="figure-67@85.[148,1148,216,1572]" captionTargetPageId="85" captionText="FIGURE 45. Photographs of habitats for Broscosoma species in the Gaoligong Shan region. a. Snowmelt stream on north-facing slope above North Fork of Yamu He, 8.5 km W of Shibali, 3100-3200 m, 27.18315º/098.71921º, Lumadeng Township, Fugong County, Yunnan (habitat for Broscosoma parvum sp. nov.); b. 0.6 km N of Dizhengdang village on Dulongjiang, 1880 m, 28.084º/098.327º, Dulongjiang Township, Gongshan County, Yunnan (habitat for Broscosoma purpureum sp. nov.). Photos by David H. Kavanaugh" figureDoi="http://doi.org/10.5281/zenodo.11067449" httpUri="https://zenodo.org/record/11067449/files/figure.png" pageId="51" pageNumber="136">Fig. 45a</figureCitation>
). This species is found at moderately high elevations, with our records documenting its occurrence in the
<quantity id="4CFE17FFFFECFF86FC03FEF2A590FEBA" box="[999,1174,317,341]" metricMagnitude="3" metricUnit="m" metricValue="3.195" metricValueMax="3.29" metricValueMin="3.1" pageId="51" pageNumber="136" unit="m" value="3195.0" valueMax="3290.0" valueMin="3100.0">3100 to 3290 m</quantity>
range (
<figureCitation id="133DA69FFFECFF86FF22FE92A015FE9B" box="[198,275,349,373]" captionStart="FIGURE" captionStartId="88.[184,196,1682,1701]" captionTargetBox="[130,1162,1063,1660]" captionTargetId="figure-332@88.[120,1176,1054,1668]" captionTargetPageId="88" captionText="FIGURE 49. Chart illustrating the altitudinal ranges of broscine species represented in the Gaoligong Shan region. Green bars mark the elevational range recorded for each species." figureDoi="http://doi.org/10.5281/zenodo.11067457" httpUri="https://zenodo.org/record/11067457/files/figure.png" pageId="51" pageNumber="136">Fig. 49</figureCitation>
).
</paragraph>
<paragraph id="8BB9BA1AFFECFF86FF4FFEB2A0F9FE19" blockId="51.[123,1174,219,634]" pageId="51" pageNumber="136">
<emphasis id="B9726608FFECFF86FF4FFEB2A218FE78" bold="true" box="[171,798,381,406]" pageId="51" pageNumber="136">
Geographical distribution within the Gaoligong
<materialsCitation id="3B6EB047FFECFF86FD39FEB2A590FE58" ID-GBIF-Occurrence="4899905312" location="We" pageId="51" pageNumber="136" specimenCount="14" specimenCount-female="3" specimenCount-male="4" stateProvince="Shan">
<collectingRegion id="49C274F8FFECFF86FD39FEB2A211FE7B" box="[733,791,381,405]" country="Myanmar" name="Shan" pageId="51" pageNumber="136">Shan</collectingRegion>
.
<figureCitation id="133DA69FFFECFF86FCC3FEB1A285FE78" box="[807,899,382,406]" captionStart="FIGURE" captionStartId="48.[171,183,1587,1606]" captionTargetBox="[132,1176,216,1571]" captionTargetId="figure-56@48.[120,1176,216,1571]" captionTargetPageId="48" captionText="FIGURE 26. Broscosoma parvum sp. nov. a. Habitus (CASENT1020001; first cirque S of Shibali Yakou, Lumadeng Township, Fugong County, Yunnan, China), dorsal aspect, scale line = 1.0 mm; b. Map showing locality records (blue circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067407" httpUri="https://zenodo.org/record/11067407/files/figure.png" pageId="51" pageNumber="136">Fig. 26b</figureCitation>
.
<location id="8ED9ECC1FFECFF86FC77FEB1A2B0FE7B" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFECFF86FC77FEB1A2B0FE7B" box="[915,950,382,405]" name="We" pageId="51" pageNumber="136" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D007193FFECFF86FF9FFE50A029FE58" box="[123,303,414,438]" count="7" pageId="51" pageNumber="136" type="generic">seven specimens</specimenCount>
(
<specimenCount id="9D007193FFECFF86FEA5FE51A0B2FE58" box="[321,436,414,438]" count="4" pageId="51" pageNumber="136" type="male">four males</specimenCount>
and
<specimenCount id="9D007193FFECFF86FE09FE51A378FE58" box="[493,638,414,438]" count="3" pageId="51" pageNumber="136" type="female">three females</specimenCount>
), all from the northcentral part of the Gaoligong
</materialsCitation>
</emphasis>
<materialsCitation id="3B6EB047FFECFF86FF9FFE70A0FEFE16" ID-GBIF-Occurrence="4899905539" county="Fugong County" location="Core Area" pageId="51" pageNumber="136" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49C274F8FFECFF86FF9FFE70A1B6FE39" box="[123,176,447,471]" country="Myanmar" name="Shan" pageId="51" pageNumber="136">Shan</collectingRegion>
in
<collectingCounty id="62D8C296FFECFF86FF30FE70A07AFE39" box="[212,380,447,471]" pageId="51" pageNumber="136">Fugong County</collectingCounty>
on the eastern side of the range in
<location id="8ED9ECC1FFECFF86FD1DFE70A261FE39" LSID="urn:lsid:plazi:treatment:03AF0B0CFFF0FF86FF9FFEA1A586FD97:8ED9ECC1FFECFF86FD1DFE70A261FE39" box="[761,871,447,471]" county="Fugong County" name="Core Area" pageId="51" pageNumber="136" stateProvince="Shan">Core Area</location>
3 (
<figureCitation id="133DA69FFFECFF86FC69FE70A2DCFE39" box="[909,986,447,471]" captionStart="FIGURE" captionStartId="88.[171,183,995,1014]" captionTargetBox="[130,1164,233,969]" captionTargetId="figure-57@88.[120,1176,215,989]" captionTargetPageId="88" captionText="FIGURE 48. Chart showing the representation of broscine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." figureDoi="http://doi.org/10.5281/zenodo.11067455" httpUri="https://zenodo.org/record/11067455/files/figure.png" pageId="51" pageNumber="136">Fig. 48</figureCitation>
) (see
<typeStatus id="54BD04B8FFECFF86FBF8FE70A557FE39" box="[1052,1105,447,471]" pageId="51" pageNumber="136">Type</typeStatus>
material above for exact collection data)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9BA1AFFECFF86FF4FFDCFA25FFDD6" blockId="51.[123,1174,219,634]" pageId="51" pageNumber="136">
<emphasis id="B9726608FFECFF86FF4FFDCFA336FDF6" bold="true" box="[171,560,512,536]" pageId="51" pageNumber="136">Overall geographical distribution.</emphasis>
This species currently is known only from the northcentral part of the Gaoligong Shan, in western
<collectingRegion id="49C274F8FFECFF86FDA8FDEEA202FDD6" box="[588,772,544,568]" country="China" name="Yunnan" pageId="51" pageNumber="136">Yunnan Province</collectingRegion>
,
<collectingCountry id="F311FA8AFFECFF86FCF6FDEFA254FDD6" box="[786,850,544,568]" name="China" pageId="51" pageNumber="136">China</collectingCountry>
.
</paragraph>
<paragraph id="8BB9BA1AFFECFF86FF4FFD8EA586FD97" blockId="51.[123,1174,219,634]" pageId="51" pageNumber="136">
<emphasis id="B9726608FFECFF86FF4FFD8EA24DFDB7" bold="true" box="[171,843,577,601]" pageId="51" pageNumber="136">
Geographical relationships with other
<taxonomicName id="4C06C199FFECFF86FD83FD8EA24DFDB7" box="[615,843,577,601]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="136" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9726608FFECFF86FD83FD8EA3E8FDB6" bold="true" box="[615,750,577,600]" italics="true" pageId="51" pageNumber="136">Broscosoma</emphasis>
species.
</taxonomicName>
</emphasis>
Members of this species have been found together (syntopic) only with those of
<taxonomicName id="4C06C199FFECFF86FD73FDADA23FFD94" authorityName="Kavanaugh &amp; Liang" authorityYear="2021" box="[663,825,610,634]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="136" phylum="Arthropoda" rank="species" species="viridicollare">
<emphasis id="B9726608FFECFF86FD73FDADA23FFD94" box="[663,825,610,634]" italics="true" pageId="51" pageNumber="136">B. viridicollare</emphasis>
</taxonomicName>
at one or more sites (
<figureCitation id="133DA69FFFECFF86FBC0FDADA577FD97" box="[1060,1137,609,634]" captionStart="FIGURE" captionStartId="89.[171,183,1461,1480]" captionTargetBox="[140,1154,249,1428]" captionTargetId="figure-68@89.[120,1176,216,1448]" captionTargetPageId="89" captionText="FIGURE 50. Chart illustrating the co-occurence (syntopy) of broscine species in samples from the same habitats and at the same sites in the Gaoligong Shan region. Incidents of syntopy marked in black represent confirmed co-occurrence, those marked in grey not confirmed by records but likely." figureDoi="http://doi.org/10.5281/zenodo.11067459" httpUri="https://zenodo.org/record/11067459/files/figure.png" pageId="51" pageNumber="136">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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