313 lines
39 KiB
XML
313 lines
39 KiB
XML
<document id="17BA52A521B94BAF33B3E8BBE07D1CE9" ID-CLB-Dataset="30691" ID-DOI="10.11646/zootaxa.4382.2.3" ID-GBIF-Dataset="6dc55986-74fa-4ac0-bca6-b301f3831180" ID-ISSN="1175-5326" ID-Zenodo-Dep="1182167" ID-ZooBank="8E248E6F-FDA5-4829-9DD7-4CC0EFDB08B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519199964901" checkinUser="plazi" docAuthor="Okamoto, Makoto & Gon, Ofer" docDate="2018" docId="03A2BF386E604066FF18580B14A49EE7" docLanguage="en" docName="zootaxa.4382.2.3.pdf" docOrigin="Zootaxa 4382 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigonus idai Okamoto & Gon 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="276" masterDocId="FF9BC7406E6D4069FF8F5C1A170D9F12" masterDocTitle="A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species" masterLastPageNumber="291" masterPageNumber="261" pageNumber="274" updateTime="1698523467852" updateUser="ExternalLinkService">
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<mods:mods id="6804A5981E8347AEBBD4CBD9A133A534" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FB03B7FC463FB058D62806CA5985030B">
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<mods:title id="9F2E20257BE66D406D1650C1D805C5D2">A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species</mods:title>
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<mods:name id="30AADF751B5418B2DD4DD16F50C14210" type="personal">
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<mods:roleTerm id="19C7A06575DE4E5D06C19A2CC324A2BB">Author</mods:roleTerm>
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<mods:namePart id="A262F90E285F8E1F497A81C82EF29641">Okamoto, Makoto</mods:namePart>
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<mods:name id="FF9DA2C54CCFEB6A74FD127E589687CF" type="personal">
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<mods:roleTerm id="4DE1349554B2BE9806A1FA66188CAADA">Author</mods:roleTerm>
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<mods:namePart id="8B771D105892D0499D220B026270DA19">Gon, Ofer</mods:namePart>
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<mods:typeOfResource id="28F17E20DA4E031954F0B80BBAF8E0C3">text</mods:typeOfResource>
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<mods:title id="C9D65F457F4751EC0835BA4A40483B36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2340370D0C4A8A6EA33941EE4D4F3B3B">
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<mods:date id="4F7A3D6FEE9A5FA8261528D910D4565E">2018</mods:date>
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<mods:number id="7E1CC324521D3FA09DAE12287BEDC47D">2018-02-21</mods:number>
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<mods:detail id="CC501AB1878E08FB0B599F73750E57C6" type="volume">
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<mods:number id="E57793C7668C867B986407D305BE877B">4382</mods:number>
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<mods:detail id="F7D901BA0FA04D28E497678288B1F62B" type="issue">
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<mods:classification id="45D876F30456CFF0AA9F7D3193EA8D24">journal article</mods:classification>
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<mods:identifier id="D7F445DDC049F49F7BAA83F279BFEF9E" type="DOI">10.11646/zootaxa.4382.2.3</mods:identifier>
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<treatment id="03A2BF386E604066FF18580B14A49EE7" ID-DOI="http://doi.org/10.5281/zenodo.5998245" ID-GBIF-Taxon="141118069" ID-Zenodo-Dep="5998245" LSID="urn:lsid:plazi:treatment:03A2BF386E604066FF18580B14A49EE7" httpUri="http://treatment.plazi.org/id/03A2BF386E604066FF18580B14A49EE7" lastPageId="15" lastPageNumber="276" pageId="13" pageNumber="274">
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<subSubSection id="C3115DA56E604064FF18580B15099B62" pageId="13" pageNumber="274" type="nomenclature">
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<paragraph id="8BB40E2E6E604064FF18580B16849B39" blockId="13.[151,725,1041,1136]" box="[151,393,1041,1067]" pageId="13" pageNumber="274">
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<heading id="D0FCB9426E604064FF18580B16849B39" bold="true" box="[151,393,1041,1067]" fontSize="11" level="1" pageId="13" pageNumber="274" reason="1">
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<emphasis id="B97FD23C6E604064FF18580B16849B39" bold="true" box="[151,393,1041,1067]" pageId="13" pageNumber="274">
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<taxonomicName id="4C0B75AD6E604064FF18580B164D9B39" authority="Okamoto & Gon, 2018" authorityName="Okamoto & Gon" authorityYear="2018" box="[151,320,1041,1067]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="274" phylum="Chordata" rank="species" species="idai" status="sp. nov.">
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<emphasis id="B97FD23C6E604064FF18580B164D9B39" bold="true" box="[151,320,1041,1067]" italics="true" pageId="13" pageNumber="274">Epigonus idai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24C6F476E604064FEC8580B16849B39" box="[327,393,1041,1067]" pageId="13" pageNumber="274" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB40E2E6E604064FF18582E15099B62" blockId="13.[151,725,1041,1136]" pageId="13" pageNumber="274">
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(New English name: Ida’s Deepwater Cardinalfish) (
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<figureCitation id="133012AB6E604064FF10584D17E09B62" box="[159,237,1111,1136]" captionStart="FIGURE 3" captionStartId="2.[151,250,1684,1706]" captionTargetBox="[341,1239,999,1649]" captionTargetId="figure@2.[302,1284,987,1662]" captionTargetPageId="2" captionText="FIGURE 3. Distribution of seven species of the Epigonus constanciae group inthe Western Indian Ocean." httpUri="https://zenodo.org/record/1182173/files/figure.png" pageId="13" pageNumber="274">Figs. 3</figureCitation>
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,
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<figureCitation id="133012AB6E604064FF75584D16109B62" box="[250,285,1111,1136]" captionStart="FIGURE6" captionStartId="7.[151,250,1874,1896]" captionTargetBox="[226,1344,1400,1852]" captionTargetId="figure@7.[226,1360,1400,1852]" captionTargetPageId="7" captionText="FIGURE6. Ventral viewof symphysis of lowerjaw.A:Epigonus bispinosusn. sp., SAIAB35606, 1of 5 paratypes,141.7 mm SL; B: Epigonus idai n. sp., SAIAB 204434, paratype, 132.3 mm SL." httpUri="https://zenodo.org/record/1182179/files/figure.png" pageId="13" pageNumber="274">6B</figureCitation>
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,
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<figureCitation id="133012AB6E604064FEA6584216419B7D" box="[297,332,1112,1135]" captionStart="FIGURE 7" captionStartId="9.[151,250,1902,1924]" captionTargetBox="[188,1359,892,1847]" captionTargetId="figure@9.[188,1361,892,1848]" captionTargetPageId="9" captionText="FIGURE 7. Pored lateral-line scales. A: Epigonus bispinosus n. sp., SAIAB 189476, holotype, 165.5 mm SL; B: E. idai, n. sp., SAIAB 204434, paratype, 132.3 mm SL." httpUri="https://zenodo.org/record/1182181/files/figure.png" pageId="13" pageNumber="274">7B</figureCitation>
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,
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<figureCitation id="133012AB6E604064FED75842167B9B62" box="[344,374,1112,1136]" captionStart="FIGURE 11" captionStartId="14.[151,250,1968,1990]" captionTargetBox="[188,1398,697,1947]" captionTargetId="figure@14.[188,1398,697,1947]" captionTargetPageId="14" captionText="FIGURE 11. Epigonus idai n. sp., SAIAB 82171, holotype, 136.8 mm SL; A: fresh specimen; B: preserved specimen; C: radiograph." httpUri="https://zenodo.org/record/1182191/files/figure.png" pageId="13" pageNumber="274">11</figureCitation>
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; Tables 1–3)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3115DA56E604064FF18588512019A12" pageId="13" pageNumber="274" type="materials_examined">
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<paragraph id="8BB40E2E6E604064FF18588512289BCE" blockId="13.[151,1437,1183,2037]" pageId="13" pageNumber="274">
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<materialsCitation id="3B6304736E604064FF18588512289BCE" ID-GBIF-Occurrence="1820369969" collectingDate="2007-10-19" collectionCode="SAIAB" collectorName="P. C. Heemstra & Heemstra." country="Mozambique" elevation="715" latitude="-20.593334" location="South" longLatPrecision="126" longitude="35.97167" pageId="13" pageNumber="274" specimenCode="SAIAB 82171, 136.8" specimenCount="1" stateProvince="South" typeStatus="holotype">
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<emphasis id="B97FD23C6E604064FF18588516069BAA" bold="true" box="[151,267,1183,1208]" pageId="13" pageNumber="274">
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<typeStatus id="54B0B08C6E604064FF185885160B9BAA" box="[151,262,1183,1208]" pageId="13" pageNumber="274" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBADA6556E604064FE9958BA16B39BAA" box="[278,446,1184,1208]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="13" pageNumber="274">
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SAIAB
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<quantity id="4CF3A3CB6E604064FEFA58BA16B39BAA" box="[373,446,1184,1208]" metricMagnitude="4" metricUnit="m" metricValue="8.21711368" pageId="13" pageNumber="274" unit="mm" value="8.21711368E7">82171</quantity>
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</specimenCode>
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||
,
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<specimenCode id="DBADA6556E604064FE42588515019BAA" box="[461,524,1183,1208]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="13" pageNumber="274">136.8</specimenCode>
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mm SL, male,
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<geoCoordinate id="EE3F68E96E604064FD46588514339BAA" box="[713,830,1183,1208]" direction="south" orientation="latitude" pageId="13" pageNumber="274" precision="92" value="-20.593334">20°35.6 S</geoCoordinate>
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,
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<geoCoordinate id="EE3F68E96E604064FCC1588514C89BA5" box="[846,965,1183,1208]" direction="east" orientation="longitude" pageId="13" pageNumber="274" precision="92" value="35.97167">35°58.3 E</geoCoordinate>
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, off
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<collectingCountry id="F31C4EBE6E604064FB8F5885139A9BAA" box="[1024,1175,1183,1208]" name="Mozambique" pageId="13" pageNumber="274">Mozambique</collectingCountry>
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, western
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<collectingRegion id="49CFC0CC6E604064FA87588512449BAA" box="[1288,1353,1183,1208]" country="Tanzania" name="Kusini Unguja" pageId="13" pageNumber="274">South</collectingRegion>
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Indian Ocean,
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<elevation id="0026E91D6E604064FF6358DE16669BCE" box="[236,363,1220,1244]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.209999999999999" metricValueMin="7.09" pageId="13" pageNumber="274" unit="m" value="715.0" valueMax="721.0" valueMin="709.0">
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<quantity id="4CF3A3CB6E604064FF6358DE16669BCE" box="[236,363,1220,1244]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.209999999999999" metricValueMin="7.09" pageId="13" pageNumber="274" unit="m" value="715.0" valueMax="721.0" valueMin="709.0">709–721 m</quantity>
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</elevation>
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depth,
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<collectingDate id="EFF1D1066E604064FE3058DE15709BCE" box="[447,637,1220,1245]" pageId="13" pageNumber="274" value="2007-10-19">
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||
<date id="FFB528EE6E604064FE3058DE15709BCE" box="[447,637,1220,1245]" pageId="13" pageNumber="274" value="2007-10-19">19 October 2007</date>
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</collectingDate>
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, bottom trawl, collected by
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<collectorName id="26FE6BF86E604064FC3958DF13509BCE" box="[950,1117,1220,1244]" pageId="13" pageNumber="274">P. C. Heemstra</collectorName>
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and E.
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<collectorName id="26FE6BF86E604064FB3D58DF12289BCE" box="[1202,1317,1221,1244]" pageId="13" pageNumber="274">Heemstra.</collectorName>
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BB40E2E6E604064FF4858FD12019A12" blockId="13.[151,1437,1183,2037]" box="[199,1292,1255,1280]" pageId="13" pageNumber="274">
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<emphasis id="B97FD23C6E604064FF4858FD16369A12" bold="true" box="[199,315,1255,1280]" pageId="13" pageNumber="274">
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<typeStatus id="54B0B08C6E604064FF4858FD163B9A12" box="[199,310,1255,1280]" pageId="13" pageNumber="274" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED1A96EB6E604064FECD58F2169B9A12" box="[322,406,1256,1280]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="13" pageNumber="274">SAIAB</collectionCode>
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204434 (formerly
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<collectionCode id="ED1A96EB6E604064FDE558F215B39A12" box="[618,702,1256,1280]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="13" pageNumber="274">SAIAB</collectionCode>
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82171),
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<quantity id="4CF3A3CB6E604064FCAD58F214999BED" box="[802,916,1256,1280]" metricMagnitude="-1" metricUnit="m" metricValue="1.323" pageId="13" pageNumber="274" unit="mm" value="132.3">132.3 mm</quantity>
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SL, male, same data as
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<typeStatus id="54B0B08C6E604064FB2D58FD12059A12" box="[1186,1288,1255,1280]" pageId="13" pageNumber="274" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3115DA56E604064FF48591616A59AA7" pageId="13" pageNumber="274" type="diagnosis">
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<paragraph id="8BB40E2E6E604064FF48591616A59AA7" blockId="13.[151,1437,1183,2037]" pageId="13" pageNumber="274">
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||
<emphasis id="B97FD23C6E604064FF485916164C9A37" bold="true" box="[199,321,1292,1317]" pageId="13" pageNumber="274">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="4C0B75AD6E604064FE665917155E9A36" authorityName="Rafinesque" authorityYear="1810" box="[489,595,1293,1316]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="274" phylum="Chordata" rank="genus">
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<emphasis id="B97FD23C6E604064FE665917155E9A36" box="[489,595,1293,1316]" italics="true" pageId="13" pageNumber="274">Epigonus</emphasis>
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</taxonomicName>
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with the following combination of characters: dorsal-fin rays VII-I, 10; pectoral-fin rays 18–19; total gill rakers 28–29; vertebrae 10 + 15; pyloric caeca 10–12; pored lateral-line scales 48–49 + 3; scales strongly ctenoid; scales below lateral line 10; pungent opercular spine present; maxillary mustache-like process absent; ribs on last abdominal vertebra present; and 4–5 tiny projections present on symphysis of lower jaw.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3115DA56E604067FF4859A5138D9EC2" lastPageId="14" lastPageNumber="275" pageId="13" pageNumber="274" type="description">
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||
<paragraph id="8BB40E2E6E604067FF4859A513249E52" blockId="13.[151,1437,1183,2037]" lastBlockId="14.[151,1437,151,645]" lastPageId="14" lastPageNumber="275" pageId="13" pageNumber="274">
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<emphasis id="B97FD23C6E604064FF4859A516549ACA" bold="true" box="[199,345,1471,1496]" pageId="13" pageNumber="274">Description.</emphasis>
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Counts and measurements of
|
||
<typeStatus id="54B0B08C6E604064FD3659A514169ACA" box="[697,795,1471,1496]" pageId="13" pageNumber="274" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="54B0B08C6E604064FCD759DA14B59ACA" box="[856,952,1472,1496]" pageId="13" pageNumber="274" type="paratype">paratype</typeStatus>
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are shown in Tables 1–3. Body elongate, laterally compressed, deepest at pectoral-fin base; nape not humped. Head large, broad. Maxillary mustache-like processes absent. Snout short and round, length slightly longer than interorbital width; nostrils closely set at level of upper edge of pupil, anterior nostril without membranous tube, posterior nostril elliptical without dermal flap. Eye large, round, orbital diameter greater than postorbital length; bony rim of orbit slightly raised above dorsal profile; interorbital region broad, concave at center of posterior half. Mouth large, terminal; gape oblique; posterior margin of maxilla not extending to below center of pupil; lower jaw not projecting when mouth closed; tiny 4–5 projections present on symphysis of lower jaw (
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<figureCitation id="133012AB6E604064FD335AA6141B99C7" box="[700,790,1724,1749]" captionStart="FIGURE6" captionStartId="7.[151,250,1874,1896]" captionTargetBox="[226,1344,1400,1852]" captionTargetId="figure@7.[226,1360,1400,1852]" captionTargetPageId="7" captionText="FIGURE6. Ventral viewof symphysis of lowerjaw.A:Epigonus bispinosusn. sp., SAIAB35606, 1of 5 paratypes,141.7 mm SL; B: Epigonus idai n. sp., SAIAB 204434, paratype, 132.3 mm SL." httpUri="https://zenodo.org/record/1182179/files/figure.png" pageId="13" pageNumber="274">Fig. 6B</figureCitation>
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). Teeth minute, arranged in 2–3 rows on anterior half of premaxilla and dentary, a single row on posterior half of both bones. Small number of minute teeth present on vomer and palatine. Basihyal toothless. Opercular spine strong, pungent, forming ridge; posterior corner of preopercle weakly serrated. Origin of first dorsal fin above anterior part of pectoral fin; first dorsal-fin spine short; third dorsal-fin spine longest; no isolated dorsal-fin spine between first and second dorsal fins. Spine of second dorsal fin long, slightly thicker than those of first dorsal fin. Origin of anal fin below posterior part of second dorsal-fin base; first anal-fin spine minute; second anal-fin spine long, but its length slightly shorter than second dorsal-fin spine. Proximal radial of first anal-fin pterygiophore slender. Pectoral and pelvic fins short, posterior tip of both fins not reaching vertical drawn from anus; upper margin of pectoral-fin base lower than level of horizontal line through center of pupil. Caudal fin deeply forked. Anus located slightly anterior to vertical drawn through origin of second dorsal fin. Ribs present on last abdominal vertebra. Supraneural bones three (0+0/0+2/1+1/1/). Scales large, deciduous, strongly ctenoid (
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<figureCitation id="133012AB6E634067FDE35CC515CF9FEA" box="[620,706,223,248]" captionStart="FIGURE 7" captionStartId="9.[151,250,1902,1924]" captionTargetBox="[188,1359,892,1847]" captionTargetId="figure@9.[188,1361,892,1848]" captionTargetPageId="9" captionText="FIGURE 7. Pored lateral-line scales. A: Epigonus bispinosus n. sp., SAIAB 189476, holotype, 165.5 mm SL; B: E. idai, n. sp., SAIAB 204434, paratype, 132.3 mm SL." httpUri="https://zenodo.org/record/1182181/files/figure.png" pageId="14" pageNumber="275">Fig. 7B</figureCitation>
|
||
), covering whole body except for area anterior to rim of orbit and surfaces of jaws; small scales also present on bases of second dorsal, anal, and caudal fins; series of pored lateralline scales complete, 3 pored scales on caudal fin. Ventral luminous organ absent.
|
||
</paragraph>
|
||
<paragraph id="8BB40E2E6E634067FF485D5613A29E9A" blockId="14.[151,1437,151,645]" pageId="14" pageNumber="275">
|
||
<emphasis id="B97FD23C6E634067FF485D5616849E76" box="[199,393,332,356]" italics="true" pageId="14" pageNumber="275">Color when fresh</emphasis>
|
||
(based on
|
||
<figureCitation id="133012AB6E634067FD8B5D5615659E76" box="[516,616,332,357]" captionStart="FIGURE 11" captionStartId="14.[151,250,1968,1990]" captionTargetBox="[188,1398,697,1947]" captionTargetId="figure@14.[188,1398,697,1947]" captionTargetPageId="14" captionText="FIGURE 11. Epigonus idai n. sp., SAIAB 82171, holotype, 136.8 mm SL; A: fresh specimen; B: preserved specimen; C: radiograph." httpUri="https://zenodo.org/record/1182191/files/figure.png" pageId="14" pageNumber="275">Fig. 11A</figureCitation>
|
||
). Body uniformly tan; pectoral, second dorsal, anal, and caudal fins light brown; first dorsal and pelvic fins black. Head light brown and opercle with bluish reflection.
|
||
</paragraph>
|
||
<paragraph id="8BB40E2E6E634067FF485D8E138D9EC2" blockId="14.[151,1437,151,645]" pageId="14" pageNumber="275">
|
||
<emphasis id="B97FD23C6E634067FF485D8E168B9EBE" box="[199,390,404,428]" italics="true" pageId="14" pageNumber="275">Color in alcohol</emphasis>
|
||
. Body, pectoral and caudal fins, head (except gill cover which is dark-brown), and mouth cavity with tongue light brown. First and second dorsal, pelvic, and anal fins dark brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3115DA56E634067FF485DC616D89D05" pageId="14" pageNumber="275" type="distribution">
|
||
<paragraph id="8BB40E2E6E634067FF485DC616D89D05" blockId="14.[151,1437,151,645]" pageId="14" pageNumber="275">
|
||
<emphasis id="B97FD23C6E634067FF485DC6166D9EE7" bold="true" box="[199,352,476,501]" pageId="14" pageNumber="275">Distribution.</emphasis>
|
||
The
|
||
<typeStatus id="54B0B08C6E634067FE125DC616F29EE7" box="[413,511,476,501]" pageId="14" pageNumber="275" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54B0B08C6E634067FDB65DC715979EE7" box="[569,666,477,501]" pageId="14" pageNumber="275" type="paratype">paratype</typeStatus>
|
||
were collected off
|
||
<collectingCountry id="F31C4EBE6E634067FCF75DC613029EE7" box="[888,1039,476,501]" name="Mozambique" pageId="14" pageNumber="275">Mozambique</collectingCountry>
|
||
, western
|
||
<collectingRegion id="49CFC0CC6E634067FBF35DC613B09EE7" box="[1148,1213,476,501]" country="Tanzania" name="Kusini Unguja" pageId="14" pageNumber="275">South</collectingRegion>
|
||
Indian Ocean (
|
||
<figureCitation id="133012AB6E634067FAE05DC617AB9D0A" captionStart="FIGURE 3" captionStartId="2.[151,250,1684,1706]" captionTargetBox="[341,1239,999,1649]" captionTargetId="figure@2.[302,1284,987,1662]" captionTargetPageId="2" captionText="FIGURE 3. Distribution of seven species of the Epigonus constanciae group inthe Western Indian Ocean." httpUri="https://zenodo.org/record/1182173/files/figure.png" pageId="14" pageNumber="275">Fig. 3</figureCitation>
|
||
), at a depth of
|
||
<quantity id="4CF3A3CB6E634067FEC15E1A16DD9D05" box="[334,464,512,536]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.209999999999999" metricValueMin="7.09" pageId="14" pageNumber="275" unit="m" value="715.0" valueMax="721.0" valueMin="709.0">
|
||
709–
|
||
<specimenCount id="9D0DC5A76E634067FE075E1A16DD9D05" box="[392,464,512,536]" pageId="14" pageNumber="275" type="generic">721 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3115DA56E634066FF485E3E14A49EE7" lastPageId="15" lastPageNumber="276" pageId="14" pageNumber="275" type="etymology">
|
||
<paragraph id="8BB40E2E6E634067FF485E3E16369D97" blockId="14.[151,1437,151,645]" pageId="14" pageNumber="275">
|
||
<emphasis id="B97FD23C6E634067FF485E3E16439D2F" bold="true" box="[199,334,548,573]" pageId="14" pageNumber="275">Etymology.</emphasis>
|
||
The specific name is in honor of Dr Hitoshi Ida (Professor Emeritus, Kitasato University, School of Marine Biosciences) for his contribution to
|
||
<taxonomicName id="4C0B75AD6E634067FD115E5214059D4D" authorityName="Rafinesque" authorityYear="1810" box="[670,776,584,607]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="275" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97FD23C6E634067FD115E5214059D4D" box="[670,776,584,607]" italics="true" pageId="14" pageNumber="275">Epigonus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9A73DF6E634067FCCD5E5D14E19D72" box="[834,1004,583,608]" pageId="14" pageNumber="275" refId="ref19644" refString="Ida, H., Okamoto, M. & Sakaue, J. (2007) Epigonus cavaticus (Teleostei: Perciformes), a new epigonid fish from Palau, western Central Pacific. Ichthyological Research, 54, 131 - 136. https: // doi. org / 10.1007 / s 10228 - 006 - 0384 - 9" type="journal article">
|
||
Ida
|
||
<emphasis id="B97FD23C6E634067FCE25E5314909D72" box="[877,925,584,608]" italics="true" pageId="14" pageNumber="275">et al</emphasis>
|
||
., 2007
|
||
</bibRefCitation>
|
||
) and other percoid studies in the Indo- Pacific region.
|
||
</paragraph>
|
||
<caption id="DF745EA66E634067FF185BAA160198F7" httpUri="https://zenodo.org/record/1182191/files/figure.png" pageId="14" pageNumber="275" startId="14.[151,250,1968,1990]" targetBox="[188,1398,697,1947]" targetPageId="14">
|
||
<paragraph id="8BB40E2E6E634067FF185BAA160198F7" blockId="14.[151,1436,1968,2021]" pageId="14" pageNumber="275">
|
||
<emphasis id="B97FD23C6E634067FF185BAA162C98D7" bold="true" box="[151,289,1968,1990]" pageId="14" pageNumber="275">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C0B75AD6E634067FEA55BAB16B498D4" authority="Okamoto & Gon, 2018" authorityName="Okamoto & Gon" authorityYear="2018" box="[298,441,1968,1990]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="275" phylum="Chordata" rank="species" species="idai" status="sp. nov.">
|
||
<emphasis id="B97FD23C6E634067FEA55BAB16B498D4" box="[298,441,1968,1990]" italics="true" pageId="14" pageNumber="275">Epigonus idai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C6F476E634067FE4C5BAB16F298D4" box="[451,511,1969,1990]" pageId="14" pageNumber="275" rank="species">
|
||
<emphasis id="B97FD23C6E634067FE4C5BAB16F298D4" bold="true" box="[451,511,1969,1990]" pageId="14" pageNumber="275">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, SAIAB 82171, holotype, 136.8 mm SL; A: fresh specimen; B: preserved specimen; C: radiograph.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB40E2E6E624066FF485C8D14A49EE7" blockId="15.[151,1436,151,501]" pageId="15" pageNumber="276">
|
||
<emphasis id="B97FD23C6E624066FF485C8D152F9FA2" bold="true" box="[199,546,151,176]" pageId="15" pageNumber="276">Comparisons and Remarks.</emphasis>
|
||
<taxonomicName id="4C0B75AD6E624066FDBF5C8215D99FA2" authorityName="Okamoto & Gon" authorityYear="2018" box="[560,724,152,176]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="idai">
|
||
<emphasis id="B97FD23C6E624066FDBF5C8215D99FA2" box="[560,724,152,176]" italics="true" pageId="15" pageNumber="276">Epigonus idai</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C0B75AD6E624066FC155C83134B9FBD" baseAuthorityName="Giglioli" baseAuthorityYear="1880" box="[922,1094,152,176]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="constanciae">
|
||
<emphasis id="B97FD23C6E624066FC155C83134B9FBD" box="[922,1094,152,176]" italics="true" pageId="15" pageNumber="276">E. constanciae</emphasis>
|
||
</taxonomicName>
|
||
group by having a pungent opercular spine (Okamoto 2012).
|
||
<taxonomicName id="4C0B75AD6E624066FD975CA715B89FC6" authorityName="Okamoto & Gon" authorityYear="2018" box="[536,693,188,212]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="idai">
|
||
<emphasis id="B97FD23C6E624066FD975CA715B89FC6" box="[536,693,188,212]" italics="true" pageId="15" pageNumber="276">Epigonus idai</emphasis>
|
||
</taxonomicName>
|
||
is unique in this species group in having 4–5 tiny projections on the symphysis of lower jaw (
|
||
<figureCitation id="133012AB6E624066FE565CC5153D9FEA" box="[473,560,223,248]" captionStart="FIGURE6" captionStartId="7.[151,250,1874,1896]" captionTargetBox="[226,1344,1400,1852]" captionTargetId="figure@7.[226,1360,1400,1852]" captionTargetPageId="7" captionText="FIGURE6. Ventral viewof symphysis of lowerjaw.A:Epigonus bispinosusn. sp., SAIAB35606, 1of 5 paratypes,141.7 mm SL; B: Epigonus idai n. sp., SAIAB 204434, paratype, 132.3 mm SL." httpUri="https://zenodo.org/record/1182179/files/figure.png" pageId="15" pageNumber="276">Fig. 6B</figureCitation>
|
||
). In addition to the diagnostic characters,
|
||
<taxonomicName id="4C0B75AD6E624066FB8B5CFB13429FEA" authorityName="Okamoto & Gon" authorityYear="2018" box="[1028,1103,224,248]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="idai">
|
||
<emphasis id="B97FD23C6E624066FB8B5CFB13429FEA" box="[1028,1103,224,248]" italics="true" pageId="15" pageNumber="276">E. idai</emphasis>
|
||
</taxonomicName>
|
||
differs from 20 species of the group in having 10 + 15 vertebrae (vs. 11 +
|
||
<quantity id="4CF3A3CB6E624066FD225D1E15E79E0F" box="[685,746,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="15" pageNumber="276" unit="in" value="14.0">14 in</quantity>
|
||
<taxonomicName id="4C0B75AD6E624066FD7A5D1F14779E0E" authorityName="Okamoto" authorityYear="2012" box="[757,890,260,284]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="chilensis">
|
||
<emphasis id="B97FD23C6E624066FD7A5D1F14779E0E" box="[757,890,260,284]" italics="true" pageId="15" pageNumber="276">E. chilensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FC035D1F134C9E0E" authorityName="de Buen" authorityYear="1959" box="[908,1089,260,284]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="crassicaudus">
|
||
<emphasis id="B97FD23C6E624066FC035D1F134C9E0E" box="[908,1089,260,284]" italics="true" pageId="15" pageNumber="276">E. crassicaudus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FBDB5D1F13E99E0E" authorityName="Parin & Abramov" authorityYear="1986" box="[1108,1252,260,284]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="heracleus">
|
||
<emphasis id="B97FD23C6E624066FBDB5D1F13E99E0E" box="[1108,1252,260,284]" italics="true" pageId="15" pageNumber="276">E. heracleus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FB795D1F127F9E0E" baseAuthorityName="Whitley" baseAuthorityYear="1935" box="[1270,1394,260,284]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="lenimen">
|
||
<emphasis id="B97FD23C6E624066FB795D1F127F9E0E" box="[1270,1394,260,284]" italics="true" pageId="15" pageNumber="276">E. lenimen</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FA0B5D1F160A9E52" authorityName="Okamoto" authorityYear="2012" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="machaera">
|
||
<emphasis id="B97FD23C6E624066FA0B5D1F160A9E52" italics="true" pageId="15" pageNumber="276">E. machaera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FE975D3316979E52" baseAuthorityName="Barnard" baseAuthorityYear="1927" box="[280,410,296,320]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="B97FD23C6E624066FE975D3316979E52" box="[280,410,296,320]" italics="true" pageId="15" pageNumber="276">E. robustus</emphasis>
|
||
</taxonomicName>
|
||
), 10 soft rays on the second dorsal fin (vs. 9 soft rays in
|
||
<taxonomicName id="4C0B75AD6E624066FBC35D3313B99E52" authorityName="Parin & Abramov" authorityYear="1986" box="[1100,1204,296,320]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="B97FD23C6E624066FBC35D3313B99E52" box="[1100,1204,296,320]" italics="true" pageId="15" pageNumber="276">E. affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FB495D3312339E52" authorityName="Parin & Abramov" authorityYear="1986" box="[1222,1342,296,320]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B97FD23C6E624066FB495D3312339E52" box="[1222,1342,296,320]" italics="true" pageId="15" pageNumber="276">E. elegans</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0B75AD6E624066FA0B5D33162F9E76" authorityName="Parin & Abramov" authorityYear="1986" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="waltersensis">
|
||
<emphasis id="B97FD23C6E624066FA0B5D33162F9E76" italics="true" pageId="15" pageNumber="276">E. waltersensis</emphasis>
|
||
</taxonomicName>
|
||
), 28–29 gill rakers (vs.
|
||
<quantity id="4CF3A3CB6E624066FDA45D5615839E77" box="[555,654,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.604" metricValueMin="5.08" pageId="15" pageNumber="276" unit="in" value="23.0" valueMax="26.0" valueMin="20.0">20–26 in</quantity>
|
||
<taxonomicName id="4C0B75AD6E624066FD1B5D5714329E76" baseAuthorityName="Gilbert" baseAuthorityYear="1905" box="[660,831,332,356]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="atherinoides">
|
||
<emphasis id="B97FD23C6E624066FD1B5D5714329E76" box="[660,831,332,356]" italics="true" pageId="15" pageNumber="276">E. atherinoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FCC35D5714D19E76" authorityName="Mochizuki and Shirakihara" authorityYear="1983" box="[844,988,332,356]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="ctenolepis">
|
||
<emphasis id="B97FD23C6E624066FCC35D5714D19E76" box="[844,988,332,356]" italics="true" pageId="15" pageNumber="276">E. ctenolepis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FC655D5713479E76" authorityName="Okamoto" authorityYear="2015" box="[1002,1098,332,356]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="draco">
|
||
<emphasis id="B97FD23C6E624066FC655D5713479E76" box="[1002,1098,332,356]" italics="true" pageId="15" pageNumber="276">E. draco</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FBD75D5713EC9E76" authorityName="Smith & Radcliffe" authorityYear="1912" box="[1112,1249,332,356]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="megalops">
|
||
<emphasis id="B97FD23C6E624066FBD75D5713EC9E76" box="[1112,1249,332,356]" italics="true" pageId="15" pageNumber="276">E. megalops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FB605D57127B9E76" authorityName="Fricke" authorityYear="2017" box="[1263,1398,332,356]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="okamotoi">
|
||
<emphasis id="B97FD23C6E624066FB605D57127B9E76" box="[1263,1398,332,356]" italics="true" pageId="15" pageNumber="276">E. okamotoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FA0B5D5716139E9A" authorityName="Goode & Bean" authorityYear="1896" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="occidentalis">
|
||
<emphasis id="B97FD23C6E624066FA0B5D5716139E9A" italics="true" pageId="15" pageNumber="276">E. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
), a pair of ribs on the last abdominal vertebra (vs. absent in
|
||
<taxonomicName id="4C0B75AD6E624066FC785D6B13C79E9A" authorityName="Parin & Abramov" authorityYear="1986" box="[1015,1226,368,392]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="marimonticolus">
|
||
<emphasis id="B97FD23C6E624066FC785D6B13C79E9A" box="[1015,1226,368,392]" italics="true" pageId="15" pageNumber="276">E. marimonticolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B75AD6E624066FA865D6B12559E9A" authorityName="Prokofiev & Bussarawit" authorityYear="2012" box="[1289,1368,368,392]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="thai">
|
||
<emphasis id="B97FD23C6E624066FA865D6B12559E9A" box="[1289,1368,368,392]" italics="true" pageId="15" pageNumber="276">E. thai</emphasis>
|
||
</taxonomicName>
|
||
), and lacking a pair of maxillary mustache-like processes (vs. a sharp-pointed maxillary mustache-like processes present in
|
||
<taxonomicName id="4C0B75AD6E624066FF395DA316509EDD" baseAuthorityName="Giglioli" baseAuthorityYear="1880" box="[182,349,440,464]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="constanciae">
|
||
<emphasis id="B97FD23C6E624066FF395DA316509EDD" box="[182,349,440,464]" italics="true" pageId="15" pageNumber="276">E. constanciae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B75AD6E624066FEE35DA316D69EDD" authorityName="Okamoto" authorityYear="2011" box="[364,475,440,464]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="mayeri">
|
||
<emphasis id="B97FD23C6E624066FEE35DA316D69EDD" box="[364,475,440,464]" italics="true" pageId="15" pageNumber="276">E. mayeri</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0B75AD6E624066FD925DA315A49EC2" authorityName="Mayer" authorityYear="1974" box="[541,681,440,464]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="pectinifer">
|
||
<emphasis id="B97FD23C6E624066FD925DA315A49EC2" box="[541,681,440,464]" italics="true" pageId="15" pageNumber="276">E. pectinifer</emphasis>
|
||
</taxonomicName>
|
||
). The diagnostic characters separating the two new species in the present study are described in the Comparison of
|
||
<taxonomicName id="4C0B75AD6E624066FD4D5DC7145A9EE6" authorityName="Okamoto & Gon" authorityYear="2018" box="[706,855,476,500]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="276" phylum="Chordata" rank="species" species="bispinosus">
|
||
<emphasis id="B97FD23C6E624066FD4D5DC7145A9EE6" box="[706,855,476,500]" italics="true" pageId="15" pageNumber="276">E. bispinosus</emphasis>
|
||
</taxonomicName>
|
||
above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |