285 lines
35 KiB
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285 lines
35 KiB
XML
<document id="4C358DFE0BB2ABE34F39D57A94CD7A4A" ID-CLB-Dataset="6323" ID-DOI="10.11646/phytotaxa.8.1.4" ID-GBIF-Dataset="cb03700a-fa68-425f-9546-f580382a091c" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894896" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621603841012" checkinUser="felipe" docAuthor="Turner, I. M." docDate="2010" docId="0392EA7CFFD6FF94FF38F9ACFA62FBAE" docLanguage="en" docName="Phytotaxa.8.41-45.pdf" docOrigin="Phytotaxa 8" docStyle="DocumentStyle:E765CC7F3D29AC31B7D5F874DD99C2DD.4:Phytotaxa.2009-2010.journal_article" docStyleId="E765CC7F3D29AC31B7D5F874DD99C2DD" docStyleName="Phytotaxa.2009-2010.journal_article" docStyleVersion="4" docTitle="Cleistopetalum H. Okada 1996" docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="FFAB9204FFD6FF95FFAAFFEAFFE2FFFC" masterDocTitle="A consideration of Cleistopetalum and a new combination in Polyalthia (Annonaceae)" masterLastPageNumber="45" masterPageNumber="41" pageNumber="41" updateTime="1698941252101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E5C716E4AED933495FDCF56369C881DA">
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<mods:title id="1D51F06D690DDBC4AE57A060CC6FB51D">A consideration of Cleistopetalum and a new combination in Polyalthia (Annonaceae)</mods:title>
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</mods:titleInfo>
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<mods:name id="39D47ADEDB8E561C641E2D1DAA25EA30" type="personal">
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<mods:role id="2D6E1BE0CB9881D690B0A0293226E3F1">
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<mods:namePart id="F18A6E736080A99C9606124EFA11B520">Turner, I. M.</mods:namePart>
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<mods:typeOfResource id="A7A8B00B6E3CBFB62A28EB172E3D1098">text</mods:typeOfResource>
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<mods:titleInfo id="3BD7642C66F032EAF1BFEC5FF10E8651">
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<mods:title id="23D457F4A0AD6F3385E2AC4881C2EEA4">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="72DB43D97199B75094451C0D5D9036BE">
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<mods:date id="1764133ED0D4EF8B16F8CFEA65E782C6">2010</mods:date>
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<mods:number id="479E5A2AF7C0169D47FB75ECCCA56342">2010-08-31</mods:number>
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<mods:detail id="32BEBE770D0BEFA444301385A592FE68" type="volume">
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<mods:number id="0972964C86B08942C786DC9794E30734">8</mods:number>
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</mods:detail>
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<mods:extent id="6523D2BD7CA44D7C06FB6649BE5B7F48" unit="page">
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<mods:start id="63A0B397D6566F5050C4FA3791024D39">41</mods:start>
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<mods:end id="EE97FCF7555C2F6192D93D083C307CDB">45</mods:end>
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<mods:classification id="92E82A4003586AC6E5EE872E7E3870B7">journal article</mods:classification>
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<mods:identifier id="5DD6DCA4D138910BA98F903C226DA0CC" type="CLB-Dataset">6323</mods:identifier>
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<mods:identifier id="2D6589C81F1CEEBAE93FF76EC17A1722" type="DOI">10.11646/phytotaxa.8.1.4</mods:identifier>
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<treatment id="0392EA7CFFD6FF94FF38F9ACFA62FBAE" ID-DOI="http://doi.org/10.5281/zenodo.5593441" ID-GBIF-Taxon="182486189" ID-Zenodo-Dep="5593441" LSID="urn:lsid:plazi:treatment:0392EA7CFFD6FF94FF38F9ACFA62FBAE" httpUri="http://treatment.plazi.org/id/0392EA7CFFD6FF94FF38F9ACFA62FBAE" lastPageId="1" lastPageNumber="42" pageId="0" pageNumber="41">
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<subSubSection id="C32108E1FFD6FF95FF38F9ACFE07F99E" box="[146,485,1606,1635]" pageId="0" pageNumber="41" type="nomenclature">
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<paragraph id="8B845B6AFFD6FF95FF38F9ACFE07F99E" blockId="0.[146,485,1606,1635]" box="[146,485,1606,1635]" pageId="0" pageNumber="41">
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<heading id="D0CCEC06FFD6FF95FF38F9ACFE07F99E" bold="true" box="[146,485,1606,1635]" fontSize="12" level="1" pageId="0" pageNumber="41" reason="1">
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<emphasis id="B94F8778FFD6FF95FF38F9ACFE07F99E" bold="true" box="[146,485,1606,1635]" pageId="0" pageNumber="41">
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Species of
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<taxonomicName id="4C3B20E9FFD6FF95FE8AF9ACFE07F99E" authorityName="H. Okada" authorityYear="1996" box="[288,485,1606,1634]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD6FF95FE8AF9ACFE07F99E" bold="true" box="[288,485,1606,1634]" italics="true" pageId="0" pageNumber="41">Cleistopetalum</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32108E1FFD6FF94FF38F978FA62FBAE" lastPageId="1" lastPageNumber="42" pageId="0" pageNumber="41" type="discussion">
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<paragraph id="8B845B6AFFD6FF95FF38F978FE23F8BB" blockId="0.[146,1431,1682,2018]" pageId="0" pageNumber="41">
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The species,
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<taxonomicName id="4C3B20E9FFD6FF95FE80F979FCC1F950" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[298,803,1682,1708]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="borneense">
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<emphasis id="B94F8778FFD6FF95FE80F979FDB9F950" box="[298,603,1683,1708]" italics="true" pageId="0" pageNumber="41">Cleistopetalum borneense</emphasis>
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<bibRefCitation id="EFAA269BFFD6FF95FDC9F978FCC1F950" author="Okada, H." box="[611,803,1682,1708]" pageId="0" pageNumber="41" pagination="1 - 9" refId="ref4221" refString="Okada, H. (1996) New genus and new species of the Annonaceae from the Malesian wet tropics. Acta Phytotaxonomica et Geobotanica 47: 1 - 9." type="journal article" year="1996">Okada (1996: 4)</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C3B20E9FFD6FF95FCF6F979FB3BF950" authority="Okada (1996: 5)" authorityName="Okada" authorityPageNumber="5" authorityYear="1996" box="[860,1241,1682,1708]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="sumatranum">
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<emphasis id="B94F8778FFD6FF95FCF6F979FBF2F950" box="[860,1040,1683,1708]" italics="true" pageId="0" pageNumber="41">C. sumatranum</emphasis>
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<bibRefCitation id="EFAA269BFFD6FF95FBB2F978FB3BF950" author="Okada, H." box="[1048,1241,1682,1708]" pageId="0" pageNumber="41" pagination="1 - 9" refId="ref4221" refString="Okada, H. (1996) New genus and new species of the Annonaceae from the Malesian wet tropics. Acta Phytotaxonomica et Geobotanica 47: 1 - 9." type="journal article" year="1996">Okada (1996: 5)</bibRefCitation>
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</taxonomicName>
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, were described as new and not transferred from other genera. Although it has not been possible for me to gain direct access to the type material of either species, by studying the original protologues,
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<typeStatus id="5480E5C8FFD6FF95FB92F90AFB46F906" box="[1080,1188,1760,1786]" pageId="0" pageNumber="41" type="paratype">paratype</typeStatus>
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material and other collections, and photographs of type specimens, I have been able to identify the two species of
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<taxonomicName id="4C3B20E9FFD6FF95FB4FF8EDFA74F8DC" authorityName="H. Okada" authorityYear="1996" box="[1253,1430,1799,1824]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD6FF95FB4FF8EDFA74F8DC" box="[1253,1430,1799,1824]" italics="true" pageId="0" pageNumber="41">Cleistopetalum</emphasis>
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</taxonomicName>
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as taxa already published.
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</paragraph>
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<paragraph id="8B845B6AFFD6FF94FF6BF8BEFF29FC96" blockId="0.[146,1431,1682,2018]" lastBlockId="1.[151,1436,152,1106]" lastPageId="1" lastPageNumber="42" pageId="0" pageNumber="41">
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<taxonomicName id="4C3B20E9FFD6FF95FF6BF8BEFE10F891" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[193,498,1876,1901]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="borneense">
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<emphasis id="B94F8778FFD6FF95FF6BF8BEFE10F891" box="[193,498,1876,1901]" italics="true" pageId="0" pageNumber="41">Cleistopetalum borneense</emphasis>
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</taxonomicName>
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was designated as the
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<typeStatus id="5480E5C8FFD6FF95FCABF8BEFCD1F892" box="[769,819,1876,1902]" pageId="0" pageNumber="41">type</typeStatus>
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species of the genus, so I will deal with it first. The collections reported as being
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<taxonomicName id="4C3B20E9FFD6FF95FE5BF891FD6CF868" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[497,654,1915,1940]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="borneense">
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<emphasis id="B94F8778FFD6FF95FE5BF891FD6CF868" box="[497,654,1915,1940]" italics="true" pageId="0" pageNumber="41">C. borneense</emphasis>
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</taxonomicName>
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, both from
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<collectingRegion id="49FF9588FFD6FF95FCB7F890FC07F868" box="[797,997,1914,1940]" country="Indonesia" name="Kalimantan Timur" pageId="0" pageNumber="41">East Kalimantan</collectingRegion>
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, are sufficiently similar to what was first described as
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<taxonomicName id="4C3B20E9FFD6FF95FEDCF84BFC8AF847" authority="King (1892: 60)" authorityName="King" authorityPageNumber="60" authorityYear="1892" box="[374,872,1953,1979]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="macropoda">
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<emphasis id="B94F8778FFD6FF95FEDCF84BFD72F846" box="[374,656,1953,1978]" italics="true" pageId="0" pageNumber="41">Polyalthia macropoda</emphasis>
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<bibRefCitation id="EFAA269BFFD6FF95FD31F84BFC8AF847" author="King, G." box="[667,872,1953,1979]" pageId="0" pageNumber="41" pagination="1 - 130" refId="ref3988" refString="King, G. (1892) Materials for a flora of the Malay Peninsula. No. 4. Journal of the Asiatic Society of Bengal, Part II Natural Science 61: 1 - 130." type="journal article" year="1892">King (1892: 60)</bibRefCitation>
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</taxonomicName>
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from the Malay Peninsula to be considered conspecific. Given that
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<taxonomicName id="4C3B20E9FFD6FF95FE1EF822FDB6F81D" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[436,596,1992,2017]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="species" species="borneense">
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<emphasis id="B94F8778FFD6FF95FE1EF822FDB6F81D" box="[436,596,1992,2017]" italics="true" pageId="0" pageNumber="41">C. borneense</emphasis>
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</taxonomicName>
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is the
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<typeStatus id="5480E5C8FFD6FF95FD0DF822FD39F81E" box="[679,731,1992,2018]" pageId="0" pageNumber="41">type</typeStatus>
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of
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<taxonomicName id="4C3B20E9FFD6FF95FCAFF822FC5EF81D" authorityName="H. Okada" authorityYear="1996" box="[773,956,1992,2017]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="41" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD6FF95FCAFF822FC5EF81D" box="[773,956,1992,2017]" italics="true" pageId="0" pageNumber="41">Cleistopetalum</emphasis>
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</taxonomicName>
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it is necessary to consider if the genus should be maintained or reduced to the synonymy of
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<taxonomicName id="4C3B20E9FFD7FF94FCB9FF72FB8FFF4E" authority="Blume (1830: 68)" authorityName="Blume" authorityPageNumber="68" authorityYear="1830" box="[787,1133,152,178]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD7FF94FCB9FF72FC6CFF4D" box="[787,910,152,177]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
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<bibRefCitation id="EFAA269BFFD7FF94FC3DFF72FB8FFF4E" author="Blume, C. L." box="[919,1133,152,178]" pageId="1" pageNumber="42" refId="ref3827" refString="Blume, C. L. (1830) Flora Javae. Anonaceae. J. Frank, Brussels, 56 pp." type="book" year="1830">Blume (1830: 68)</bibRefCitation>
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</taxonomicName>
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.
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<taxonomicName id="4C3B20E9FFD7FF94FBD0FF72FB17FF4D" authorityName="Blume" authorityYear="1830" box="[1146,1269,152,177]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD7FF94FBD0FF72FB17FF4D" box="[1146,1269,152,177]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
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</taxonomicName>
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is a large and distinctly unnatural genus of Old World
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<taxonomicName id="4C3B20E9FFD7FF94FDDDFF54FCEBFF24" box="[631,777,190,216]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
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that has been something of a dumping ground for nonclimbing species bearing relatively open flowers with little difference between the corolla whorls. Molecular analysis has provided strong evidence of the polyphyletic nature of
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<emphasis id="B94F8778FFD7FF94FC11FEE6FB54FED9" box="[955,1206,268,293]" italics="true" pageId="1" pageNumber="42">
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<taxonomicName id="4C3B20E9FFD7FF94FC11FEE6FBD7FED9" authorityName="Blume" authorityYear="1830" box="[955,1077,268,293]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus" sensu="lato">Polyalthia</taxonomicName>
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<taxonomicNameLabel id="A27C3A03FFD7FF94FB94FEE6FB54FED9" box="[1086,1206,268,293]" pageId="1" pageNumber="42" sensu="lato">sensu lato</taxonomicNameLabel>
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</emphasis>
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(
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<bibRefCitation id="EFAA269BFFD7FF94FB6CFEE6FA72FEDA" author="Mols, J. B. & Gravendeel, B. & Chatrou, L. W. & Pirie, M. D. & Bygrave, P. C. & Chase, M. W. & Kessler, P. J. A." box="[1222,1424,268,294]" pageId="1" pageNumber="42" pagination="590 - 600" refId="ref4126" refString="Mols, J. B., Gravendeel, B. Chatrou, L. W., Pirie, M. D., Bygrave, P. C., Chase, M. W. & Kessler, P. J. A. (2004) Identifying clades in Asian Annonaceae: monophyletic genera in the polyphyletic Miliuseae. American Journal of Botany 91: 590 - 600." type="journal article" year="2004">
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Mols
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<emphasis id="B94F8778FFD7FF94FAA6FEE6FAA2FED9" box="[1292,1344,268,293]" italics="true" pageId="1" pageNumber="42">et al</emphasis>
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. 2004
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</bibRefCitation>
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).
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<taxonomicName id="4C3B20E9FFD7FF94FF3DFED8FE03FEB0" authority="King" authorityName="King" authorityYear="1892" box="[151,481,306,332]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="macropoda">
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<emphasis id="B94F8778FFD7FF94FF3DFED8FE7FFEB7" box="[151,413,306,331]" italics="true" pageId="1" pageNumber="42">Polyalthia macropoda</emphasis>
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King
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</taxonomicName>
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, with its single-seeded monocarps, clearly belongs in
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<taxonomicName id="4C3B20E9FFD7FF94FBF7FED8FED9FE8F" authority="(Miq.) Benth." authorityName="Benth." baseAuthorityName="Miq." class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="section" section="Monoon">
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<emphasis id="B94F8778FFD7FF94FBF7FED8FB35FEB7" box="[1117,1239,306,331]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
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section
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<emphasis id="B94F8778FFD7FF94FA90FED8FA79FEB7" box="[1338,1435,306,331]" italics="true" pageId="1" pageNumber="42">Monoon</emphasis>
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(Miq.) Benth.
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</taxonomicName>
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, though this is itself something of a mixed bag of species and still remains to be typified. Other genera, such as
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<taxonomicName id="4C3B20E9FFD7FF94FEFEFE6AFDA5FE66" authority="Becc." authorityName="Becc." box="[340,583,384,410]" class="Magnoliopsida" family="Annonaceae" genus="Enicosanthum" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD7FF94FEFEFE6AFE1EFE65" box="[340,508,384,409]" italics="true" pageId="1" pageNumber="42">Enicosanthum</emphasis>
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Becc.
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</taxonomicName>
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and
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<taxonomicName id="4C3B20E9FFD7FF94FD28FE6AFCCFFE65" authority="Merr." authorityName="Merr." box="[642,813,384,409]" class="Magnoliopsida" family="Annonaceae" genus="Woodiellantha" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
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<emphasis id="B94F8778FFD7FF94FD28FE6AFCCFFE65" box="[642,813,384,409]" italics="true" pageId="1" pageNumber="42">Woodiellantha</emphasis>
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</taxonomicName>
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, need to be considered as well as
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<taxonomicName id="4C3B20E9FFD7FF94FB69FE6AFAC7FE65" authority="Miq" box="[1219,1317,384,409]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="section" section="Monoon">
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<emphasis id="B94F8778FFD7FF94FB69FE6AFAC7FE65" box="[1219,1317,384,409]" italics="true" pageId="1" pageNumber="42">Monoon</emphasis>
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</taxonomicName>
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Miq. The molecular analysis of
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<bibRefCitation id="EFAA269BFFD7FF94FE31FE4CFD92FE3C" author="Mols, J. B. & Gravendeel, B. & Chatrou, L. W. & Pirie, M. D. & Bygrave, P. C. & Chase, M. W. & Kessler, P. J. A." box="[411,624,422,448]" pageId="1" pageNumber="42" pagination="590 - 600" refId="ref4126" refString="Mols, J. B., Gravendeel, B. Chatrou, L. W., Pirie, M. D., Bygrave, P. C., Chase, M. W. & Kessler, P. J. A. (2004) Identifying clades in Asian Annonaceae: monophyletic genera in the polyphyletic Miliuseae. American Journal of Botany 91: 590 - 600." type="journal article" year="2004">
|
|
Mols
|
|
<emphasis id="B94F8778FFD7FF94FE75FE4DFDF1FE3C" box="[479,531,423,448]" italics="true" pageId="1" pageNumber="42">et al</emphasis>
|
|
. (2004)
|
|
</bibRefCitation>
|
|
distinguished clade F4 which included species of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FB68FE4DFA88FE3C" authority="Becc." authorityName="Becc." box="[1218,1386,423,448]" class="Magnoliopsida" family="Annonaceae" genus="Enicosanthum" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FB68FE4DFA88FE3C" box="[1218,1386,423,448]" italics="true" pageId="1" pageNumber="42">Enicosanthum</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C3B20E9FFD7FF94FF3DFE27FE02FE1A" authority="(Miq.) Benth." authorityName="Benth." baseAuthorityName="Miq." box="[151,480,461,487]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="section" section="Monoon">
|
|
<emphasis id="B94F8778FFD7FF94FF3DFE27FEF7FE1A" box="[151,277,461,486]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
|
|
section
|
|
<emphasis id="B94F8778FFD7FF94FED6FE27FE02FE1A" box="[380,480,461,486]" italics="true" pageId="1" pageNumber="42">Monoon</emphasis>
|
|
</taxonomicName>
|
|
(though species of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FD7BFE27FC6AFE1A" authorityName="H. Okada" authorityYear="1996" box="[721,904,461,486]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FD7BFE27FC6AFE1A" box="[721,904,461,486]" italics="true" pageId="1" pageNumber="42">Cleistopetalum</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC6EFE27FB91FE1A" authority="Rauschert" authorityName="Rauschert" box="[964,1139,461,486]" class="Magnoliopsida" family="Annonaceae" genus="Woodiellantha" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FC6EFE27FB91FE1A" box="[964,1139,461,486]" italics="true" pageId="1" pageNumber="42">Woodiellantha</emphasis>
|
|
</taxonomicName>
|
|
were not sampled). The characters emphasised by
|
|
<bibRefCitation id="EFAA269BFFD7FF94FE78FE1EFD95FDF2" author="Okada, H." box="[466,631,500,526]" pageId="1" pageNumber="42" pagination="1 - 9" refId="ref4221" refString="Okada, H. (1996) New genus and new species of the Annonaceae from the Malesian wet tropics. Acta Phytotaxonomica et Geobotanica 47: 1 - 9." type="journal article" year="1996">Okada (1996)</bibRefCitation>
|
|
in describing
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC89FE1EFC35FDF1" authorityName="H. Okada" authorityYear="1996" box="[803,983,500,525]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FC89FE1EFC35FDF1" box="[803,983,500,525]" italics="true" pageId="1" pageNumber="42">Cleistopetalum</emphasis>
|
|
</taxonomicName>
|
|
- cauliflory, relatively closed flowers with the petals remaining more or less erect at anthesis and the base of the inner petals pressed over the reproductive structures (more evident in
|
|
<taxonomicName id="4C3B20E9FFD7FF94FDD0FDABFCF7FDA6" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[634,789,577,602]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneense">
|
|
<emphasis id="B94F8778FFD7FF94FDD0FDABFCF7FDA6" box="[634,789,577,602]" italics="true" pageId="1" pageNumber="42">C. borneense</emphasis>
|
|
</taxonomicName>
|
|
than
|
|
<taxonomicName id="4C3B20E9FFD7FF94FCF3FDABFBEDFDA6" authority="Okada (1996: 5)" authorityName="Okada" authorityPageNumber="5" authorityYear="1996" box="[857,1039,577,602]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="sumatranum">
|
|
<emphasis id="B94F8778FFD7FF94FCF3FDABFBEDFDA6" box="[857,1039,577,602]" italics="true" pageId="1" pageNumber="42">C. sumatranum</emphasis>
|
|
</taxonomicName>
|
|
) - are common in
|
|
<taxonomicName id="4C3B20E9FFD7FF94FB46FDABFA76FDA6" authority="Becc." authorityName="Becc." box="[1260,1428,577,602]" class="Magnoliopsida" family="Annonaceae" genus="Enicosanthum" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FB46FDABFA76FDA6" box="[1260,1428,577,602]" italics="true" pageId="1" pageNumber="42">Enicosanthum</emphasis>
|
|
</taxonomicName>
|
|
, and present in
|
|
<taxonomicName id="4C3B20E9FFD7FF94FEECFD82FE13FD7D" authority="Rauschert" authorityName="Rauschert" box="[326,497,616,641]" class="Magnoliopsida" family="Annonaceae" genus="Woodiellantha" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FEECFD82FE13FD7D" box="[326,497,616,641]" italics="true" pageId="1" pageNumber="42">Woodiellantha</emphasis>
|
|
</taxonomicName>
|
|
(distinguished by its basally connate petal whorls), as is the single basal ovule per carpel. Other species of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FE42FD65FCC8FD54" authority="(Miq.) Benth." authorityName="Benth." baseAuthorityName="Miq." box="[488,810,654,680]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="section" section="Monoon">
|
|
<emphasis id="B94F8778FFD7FF94FE42FD65FD81FD54" box="[488,611,655,680]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
|
|
section
|
|
<emphasis id="B94F8778FFD7FF94FD62FD65FCC8FD54" box="[712,810,655,680]" italics="true" pageId="1" pageNumber="42">Monoon</emphasis>
|
|
</taxonomicName>
|
|
, such as
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC30FD65FF22FD33" authority="(Blume) Kurz (1874: 52)" authorityName="Kurz" authorityPageNumber="52" authorityYear="1874" baseAuthorityName="Blume" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="lateriflora">
|
|
<emphasis id="B94F8778FFD7FF94FC30FD65FB75FD54" box="[922,1175,655,680]" italics="true" pageId="1" pageNumber="42">Polyalthia lateriflora</emphasis>
|
|
(Blume)
|
|
<bibRefCitation id="EFAA269BFFD7FF94FAA1FD64FF22FD33" author="Kurz, W. S." pageId="1" pageNumber="42" pagination="39 - 141" refId="ref4026" refString="Kurz, W. S. (1874) Contributions towards a knowledge of the Burmese flora - Part I. Journal of the Asiatic Society of Bengal. Part II Natural Science 43: 39 - 141." type="journal article" year="1874">Kurz (1874: 52)</bibRefCitation>
|
|
</taxonomicName>
|
|
, have petals reflexed at anthesis. There is, as
|
|
<bibRefCitation id="EFAA269BFFD7FF94FD79FD5FFC44FD33" author="Mols, J. B. & Gravendeel, B. & Chatrou, L. W. & Pirie, M. D. & Bygrave, P. C. & Chase, M. W. & Kessler, P. J. A." box="[723,934,693,719]" pageId="1" pageNumber="42" pagination="590 - 600" refId="ref4126" refString="Mols, J. B., Gravendeel, B. Chatrou, L. W., Pirie, M. D., Bygrave, P. C., Chase, M. W. & Kessler, P. J. A. (2004) Identifying clades in Asian Annonaceae: monophyletic genera in the polyphyletic Miliuseae. American Journal of Botany 91: 590 - 600." type="journal article" year="2004">
|
|
Mols
|
|
<emphasis id="B94F8778FFD7FF94FCBCFD5FFCA8FD32" box="[790,842,693,718]" italics="true" pageId="1" pageNumber="42">et al</emphasis>
|
|
. (2004)
|
|
</bibRefCitation>
|
|
concluded, considerable variation in floral form in clade F4, but more consistent fruit morphology. Molecular analysis to ascertain the phylogenetic position of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FE8AFCE8FE36FCE7" authorityName="H. Okada" authorityYear="1996" box="[288,468,770,795]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FE8AFCE8FE36FCE7" box="[288,468,770,795]" italics="true" pageId="1" pageNumber="42">Cleistopetalum</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C3B20E9FFD7FF94FDA5FCE8FD59FCE7" authority="Rauschert" authorityName="Rauschert" box="[527,699,770,795]" class="Magnoliopsida" family="Annonaceae" genus="Woodiellantha" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FDA5FCE8FD59FCE7" box="[527,699,770,795]" italics="true" pageId="1" pageNumber="42">Woodiellantha</emphasis>
|
|
</taxonomicName>
|
|
is required in order to improve resolution of the membership and any subdivisions of clade F4. Meanwhile, I consider
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC9DFCC3FC0BFCBE" authority="Okada (1996: 4)" authorityName="H. Okada" authorityPageNumber="4" authorityYear="1996" box="[823,1001,809,834]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="41" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FC9DFCC3FC0BFCBE" box="[823,1001,809,834]" italics="true" pageId="1" pageNumber="42">Cleistopetalum</emphasis>
|
|
</taxonomicName>
|
|
to be a synonym of
|
|
<emphasis id="B94F8778FFD7FF94FB73FCC3FF27FC95" italics="true" pageId="1" pageNumber="42">
|
|
<taxonomicName id="4C3B20E9FFD7FF94FB73FCC3FAB1FCBE" authorityName="Blume" authorityYear="1830" box="[1241,1363,809,834]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus" sensu="lato">Polyalthia</taxonomicName>
|
|
<taxonomicNameLabel id="A27C3A03FFD7FF94FAF1FCC3FF27FC95" pageId="1" pageNumber="42" sensu="lato">sensu lato</taxonomicNameLabel>
|
|
</emphasis>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8B845B6AFFD7FF94FF6CFC9CFA62FBAE" blockId="1.[151,1436,152,1106]" pageId="1" pageNumber="42">
|
|
Unfortunately
|
|
<taxonomicName id="4C3B20E9FFD7FF94FEDCFC9DFD83FC6C" authority="King" authorityName="King" authorityYear="1892" box="[374,609,886,912]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="macropoda">
|
|
<emphasis id="B94F8778FFD7FF94FEDCFC9DFDF9FC6C" box="[374,539,887,912]" italics="true" pageId="1" pageNumber="42">P. macropoda</emphasis>
|
|
King
|
|
</taxonomicName>
|
|
is a later homonym of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC26FC9DFA9CFC6C" authority="(Miq.) Mueller (1877: 95)" authorityName="Mueller" authorityPageNumber="95" authorityYear="1877" baseAuthorityName="Miq." box="[908,1406,886,912]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="macropoda">
|
|
<emphasis id="B94F8778FFD7FF94FC26FC9DFBD3FC6C" box="[908,1073,887,912]" italics="true" pageId="1" pageNumber="42">P. macropoda</emphasis>
|
|
(Miq.)
|
|
<bibRefCitation id="EFAA269BFFD7FF94FB3EFC9CFA9CFC6C" author="Mueller, F." box="[1172,1406,886,912]" pageId="1" pageNumber="42" refId="ref4199" refString="Mueller, F., von (1877) Descriptive notes on Papuan plants: Appendix. George Skinner, Melbourne." type="book" year="1877">Mueller (1877: 95)</bibRefCitation>
|
|
</taxonomicName>
|
|
, a species from New
|
|
<collectingCountry id="F32C1BFAFFD7FF94FEDDFC77FE33FC4B" box="[375,465,925,951]" name="Guinea-Bissau" pageId="1" pageNumber="42">Guinea</collectingCountry>
|
|
. I therefore provided an avowed substitute
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC73FC77FF33FC22" authority="Turner (2007: 275)" authorityName="I. M. Turner" authorityPageNumber="275" authorityYear="2007" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="sinclairiana">
|
|
<emphasis id="B94F8778FFD7FF94FC73FC77FB0FFC4A" box="[985,1261,925,950]" italics="true" pageId="1" pageNumber="42">Polyalthia sinclairiana</emphasis>
|
|
<bibRefCitation id="EFAA269BFFD7FF94FB5CFC77FF33FC22" author="Turner, I. M." pageId="1" pageNumber="42" pagination="275 - 278" refId="ref4277" refString="Turner, I. M. (2007) Two new names and two new combinations in Malesian Annonaceae. Gardens' Bulletin, Singapore 58: 275 - 278." type="journal article" year="2007">Turner (2007: 275)</bibRefCitation>
|
|
</taxonomicName>
|
|
. I was not aware at that time of the identity of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FCAFFC2EFBD9FC21" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[773,1083,964,989]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneense">
|
|
<emphasis id="B94F8778FFD7FF94FCAFFC2EFBD9FC21" box="[773,1083,964,989]" italics="true" pageId="1" pageNumber="42">Cleistopetalum borneense</emphasis>
|
|
</taxonomicName>
|
|
. As a synonym of
|
|
<taxonomicName id="4C3B20E9FFD7FF94FA8AFC2EFE8CFBF8" authority="King" authorityName="King" authorityYear="1892" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="macropoda">
|
|
<emphasis id="B94F8778FFD7FF94FA8AFC2EFEC1FBFF" italics="true" pageId="1" pageNumber="42">Polyalthia macropoda</emphasis>
|
|
King
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C3B20E9FFD7FF94FED7FC00FDC1FBFF" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" box="[381,547,1002,1027]" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneense">
|
|
<emphasis id="B94F8778FFD7FF94FED7FC00FDC1FBFF" box="[381,547,1002,1027]" italics="true" pageId="1" pageNumber="42">C. borneense</emphasis>
|
|
</taxonomicName>
|
|
should provide the correct name for the species. However
|
|
<taxonomicName id="4C3B20E9FFD7FF94FAB3FC00FEFBFBD6" authorityName="Merrill" authorityYear="1929" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneensis">
|
|
<emphasis id="B94F8778FFD7FF94FAB3FC00FEFBFBD6" italics="true" pageId="1" pageNumber="42">Polyalthia borneensis</emphasis>
|
|
</taxonomicName>
|
|
is already occupied by a different species (
|
|
<taxonomicName id="4C3B20E9FFD7FF94FC86FBFBFB49FBD7" authority="Merrill, 1929: 65" authorityName="Merrill" authorityPageNumber="65" authorityYear="1929" box="[812,1195,1041,1067]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneensis">
|
|
<emphasis id="B94F8778FFD7FF94FC86FBFBFC2FFBD6" box="[812,973,1041,1066]" italics="true" pageId="1" pageNumber="42">P. borneensis</emphasis>
|
|
<bibRefCitation id="EFAA269BFFD7FF94FC7CFBFBFB49FBD7" author="Merrill, E. D." box="[982,1195,1041,1067]" pageId="1" pageNumber="42" pagination="1 - 316" refId="ref4065" refString="Merrill, E. D. (1929) Plantae elmerianae borneenses. University of California Publications in Botany 15: 1 - 316." type="journal article" year="1929">Merrill, 1929: 65</bibRefCitation>
|
|
</taxonomicName>
|
|
), so
|
|
<taxonomicName id="4C3B20E9FFD7FF94FB4FFBFBFEEDFBAD" authority="Okada (1996: 4)" authorityName="Okada" authorityPageNumber="4" authorityYear="1996" class="Magnoliopsida" family="Annonaceae" genus="Cleistopetalum" higherTaxonomySource="CoL" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="borneense">
|
|
<emphasis id="B94F8778FFD7FF94FB4FFBFBFEEDFBAD" italics="true" pageId="1" pageNumber="42">Cleistopetalum borneense</emphasis>
|
|
</taxonomicName>
|
|
cannot be transferred to
|
|
<taxonomicName id="4C3B20E9FFD7FF94FD99FBD2FD4FFBAD" authorityName="Blume" authorityYear="1830" box="[563,685,1080,1105]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B94F8778FFD7FF94FD99FBD2FD4FFBAD" box="[563,685,1080,1105]" italics="true" pageId="1" pageNumber="42">Polyalthia</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C3B20E9FFD7FF94FD4CFBD2FC14FBAD" authorityName="I. M. Turner" authorityYear="2007" box="[742,1014,1080,1105]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="species" species="sinclairiana">
|
|
<emphasis id="B94F8778FFD7FF94FD4CFBD2FC14FBAD" box="[742,1014,1080,1105]" italics="true" pageId="1" pageNumber="42">Polyalthia sinclairiana</emphasis>
|
|
</taxonomicName>
|
|
continues to be the correct name.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |