treatments-xml/data/03/92/9D/03929D142A547617FF3A84902F9CFE21.xml

<document id="B90617747C3F01A53CD3EAFB5A7F3865" ID-DOI="10.5281/zenodo.178383" ID-GBIF-Dataset="ff588456-e455-4555-bfe9-bb1f4cd80fbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892246009" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Ahyong, Shane T." docDate="2007" docId="03929D142A547617FF3A84902F9CFE21" docLanguage="en" docName="zt01563p030.pdf" docOrigin="Zootaxa 1563" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Athanas sydneyensis Anker &amp; Ahyong, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFABE56C2A557611FFAD803A2E26FFEB" masterDocTitle="Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea)" masterLastPageNumber="30" masterPageNumber="17" pageNumber="18" updateTime="1698228108777" updateUser="plazi">
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<mods:title id="024D37A114B3B891EFBC50AD45585851">Description of two species in the alpheid shrimp genus Athanas Leach, 1814, with remarks on A. amazone Holthuis, 1951 (Decapoda, Caridea)</mods:title>
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<mods:namePart id="F72C668AA7FBE6C209BDB5D60991A7BE">Anker, Arthur</mods:namePart>
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<mods:date id="DC07B4F5FDACD29913A4BC81635F9D00">2007</mods:date>
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<treatment id="03929D142A547617FF3A84902F9CFE21" ID-DOI="http://doi.org/10.5281/zenodo.5685974" ID-GBIF-Taxon="119372135" ID-Zenodo-Dep="5685974" LSID="urn:lsid:plazi:treatment:03929D142A547617FF3A84902F9CFE21" httpUri="http://treatment.plazi.org/id/03929D142A547617FF3A84902F9CFE21" lastPageId="6" lastPageNumber="23" pageId="1" pageNumber="18">
<subSubSection id="C3217F892A547610FF3A84902F3EFB0C" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="8B842C022A547610FF3A84902FFCFB2F" blockId="1.[151,474,1194,1255]" box="[151,474,1194,1220]" pageId="1" pageNumber="18">
<heading id="D0CC9B6E2A547610FF3A84902FFCFB2F" bold="true" box="[151,474,1194,1220]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
<emphasis id="B94FF0102A547610FF3A84902FFCFB2F" bold="true" box="[151,474,1194,1220]" pageId="1" pageNumber="18">
<taxonomicName id="4C3B57812A547610FF3A84902FB6FB2F" ID-CoL="85MD5" box="[151,400,1194,1220]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A547610FF3A84902FB6FB2F" bold="true" box="[151,400,1194,1220]" italics="true" pageId="1" pageNumber="18">Athanas sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A547610FE3584902FFCFB2F" box="[408,474,1194,1220]" pageId="1" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B842C022A547610FF3A84F72F3EFB0C" blockId="1.[151,474,1194,1255]" box="[151,280,1229,1255]" pageId="1" pageNumber="18">
(
<figureCitation id="130030872A547610FF3284F72ED7FB0C" box="[159,241,1229,1255]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="1" pageNumber="18">Figs. 1</figureCitation>
, 
<figureCitation id="130030872A547610FF5284F72F29FB0C" box="[255,271,1229,1255]" captionStart="FIGURE 2" captionStartId="3.[151,255,1982,2006]" captionTargetBox="[173,1388,1144,1951]" captionTargetId="figure@3.[151,1436,1128,1964]" captionTargetPageId="3" captionText="FIGURE 2. Athanas sydneyensis n. sp., female, paratype, CL 3.2 mm, AM P 62762, Hawkesbury River, New South Wales, Australia. A – left cheliped, lateral view; B – right cheliped, lateral view. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/178385/files/figure.png" pageId="1" pageNumber="18">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A547610FF3A85272C4BF92C" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="8B842C022A547610FF3A85272C4BF92C" blockId="1.[151,1437,1309,1975]" pageId="1" pageNumber="18">
<emphasis id="B94FF0102A547610FF3A85272F60FADC" bold="true" box="[151,326,1309,1335]" pageId="1" pageNumber="18">
<typeStatus id="548092A02A547610FF3A85272EF4FADC" box="[151,210,1309,1335]" pageId="1" pageNumber="18">Type</typeStatus>
material
</emphasis>
: 
<typeStatus id="548092A02A547610FEF785272FEAFADC" box="[346,460,1309,1335]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D3DE78B2A547610FE7485272C62FADC" box="[473,580,1309,1335]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
, CL 
<quantity id="4CC381E72A547610FD2F85272CC4FADC" box="[642,738,1309,1335]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="18" unit="mm" value="4.8">4.8 mm</quantity>
, TL ca. 12.0 mm, AM P62764, mid-stream between Juno Head and Hungry Beach, Hawkesbury River, New South 
<collectingCountry id="F32C6C922A547610FC93857F2DA3FAB4" box="[830,901,1349,1375]" name="United Kingdom" pageId="1" pageNumber="18">Wales</collectingCountry>
(
<collectionCode id="ED2AB4C72A547610FC38857F2DFAFAB4" LSID="urn:lsid:biocol.org:col:15627" box="[917,988,1349,1375]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="1" pageNumber="18">NSW</collectionCode>
), 
<collectingCountry id="F32C6C922A547610FC5C857F2A46FAB4" box="[1009,1120,1349,1375]" name="Australia" pageId="1" pageNumber="18">Australia</collectingCountry>
, 
<geoCoordinate id="EE0F4AC52A547610FBC6857F2AE8FAB4" box="[1131,1230,1349,1375]" direction="south" orientation="latitude" pageId="1" pageNumber="18" precision="925" value="-33.566666">33°34’S</geoCoordinate>
, 
<geoCoordinate id="EE0F4AC52A547610FB76857F2B68FAB4" box="[1243,1358,1349,1375]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="925" value="151.26666">151°16’E</geoCoordinate>
, depth 
<quantity id="4CC381E72A547610FF3A85572EF1FA6C" box="[151,215,1389,1415]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="18" unit="m" value="10.0">10 m</quantity>
, sandy mud, coll. A. Jones 
<emphasis id="B94FF0102A547610FD8985572C47FA6D" box="[548,609,1389,1414]" italics="true" pageId="1" pageNumber="18">et al.</emphasis>
, 
<date id="FF850AC22A547610FDDF85572D22FA6C" box="[626,772,1389,1415]" pageId="1" pageNumber="18" value="1982-11-09">9 Nov 1982</date>
. 
<typeStatus id="548092A02A547610FCBC85572DA1FA6C" box="[785,903,1389,1415]" pageId="1" pageNumber="18" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D3DE78B2A547610FC3885572DD9FA6C" box="[917,1023,1389,1415]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
, CL 
<quantity id="4CC381E72A547610FB9185572ABDFA6C" box="[1084,1179,1389,1415]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="18" unit="mm" value="2.7">2.7 mm</quantity>
, TL ca. 
<quantity id="4CC381E72A547610FAAD85572B79FA6C" box="[1280,1375,1389,1415]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="18" unit="mm" value="7.5">7.5 mm</quantity>
, AM P62760, near Hungry Beach, Hawkesbury River, 
<collectionCode id="ED2AB4C72A547610FD4B85AF2D0EFA44" LSID="urn:lsid:biocol.org:col:15627" box="[742,808,1429,1455]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="1" pageNumber="18">NSW</collectionCode>
, 
<collectingCountry id="F32C6C922A547610FC9885AF2D82FA44" box="[821,932,1429,1455]" name="Australia" pageId="1" pageNumber="18">Australia</collectingCountry>
, 
<geoCoordinate id="EE0F4AC52A547610FC1C85AF2A0CFA44" box="[945,1066,1429,1455]" direction="south" orientation="latitude" pageId="1" pageNumber="18" precision="92" value="-33.575">33°34.5’S</geoCoordinate>
, 
<geoCoordinate id="EE0F4AC52A547610FB9A85AF2AE7FA44" box="[1079,1217,1429,1455]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="92" value="151.275">151°16.5’E</geoCoordinate>
, depth 
<quantity id="4CC381E72A547610FAB585AF2B6CFA44" box="[1304,1354,1429,1455]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="18" unit="m" value="4.0">4 m</quantity>
, sandy mud, Smith MacIntyre grab, coll. A. Jones 
<emphasis id="B94FF0102A547610FD3185872CFFFA3D" box="[668,729,1469,1494]" italics="true" pageId="1" pageNumber="18">et al.</emphasis>
, 
<date id="FF850AC22A547610FD4485872DADFA3C" box="[745,907,1469,1495]" pageId="1" pageNumber="18" value="1982-05-17">17 May 1982</date>
; 
<specimenCount id="9D3DE78B2A547610FC3485872A25FA3C" box="[921,1027,1469,1495]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
, CL 
<quantity id="4CC381E72A547610FB9285872ABBFA3C" box="[1087,1181,1469,1495]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="18" unit="mm" value="2.2">2.2 mm</quantity>
, TL ca. 6.0 mm, AM P 
<quantity id="4CC381E72A547610FF0585DF2ED2FA14" box="[168,244,1509,1535]" metricMagnitude="7" metricUnit="m" metricValue="6.27613" pageId="1" pageNumber="18" unit="m" value="6.27613E7">62761</quantity>
, 300 m NE of Green Point, Hawkesbury River, 
<collectionCode id="ED2AB4C72A547610FC9985DF2D50FA14" LSID="urn:lsid:biocol.org:col:15627" box="[820,886,1509,1535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="1" pageNumber="18">NSW</collectionCode>
, 
<collectingCountry id="F32C6C922A547610FC2F85DF2DD6FA14" box="[898,1008,1509,1535]" name="Australia" pageId="1" pageNumber="18">Australia</collectingCountry>
, 
<geoCoordinate id="EE0F4AC52A547610FC5385DF2A51FA14" box="[1022,1143,1509,1535]" direction="south" orientation="latitude" pageId="1" pageNumber="18" precision="92" value="-33.566666">33°34.0’S</geoCoordinate>
, 
<geoCoordinate id="EE0F4AC52A547610FB2885DF2B28FA14" box="[1157,1294,1509,1535]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="92" value="151.225">151°13.5’E</geoCoordinate>
, depth 
<quantity id="4CC381E72A547610FACB85DF2BBEFA14" box="[1382,1432,1509,1535]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="18" unit="m" value="5.0">5 m</quantity>
, sandy mud, coll. A. Jones &amp; A. Murray, 
<date id="FF850AC22A547610FDD286372D04F9CC" box="[639,802,1549,1575]" pageId="1" pageNumber="18" value="1982-05-17">17 May 1982</date>
; 
<specimenCount id="9D3DE78B2A547610FC9D86372DBCF9CC" box="[816,922,1549,1575]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
, CL 
<quantity id="4CC381E72A547610FC7886372A12F9CC" box="[981,1076,1549,1575]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="18" unit="mm" value="3.2">3.2 mm</quantity>
, TL ca. 9.0 mm, AM P62762, between Juno Head and Hungry Beach, Hawkesbury River, 
<collectionCode id="ED2AB4C72A547610FCFB860F2DBEF9A4" LSID="urn:lsid:biocol.org:col:15627" box="[854,920,1589,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="1" pageNumber="18">NSW</collectionCode>
, 
<collectingCountry id="F32C6C922A547610FC0F860F2A36F9A4" box="[930,1040,1589,1615]" name="Australia" pageId="1" pageNumber="18">Australia</collectingCountry>
, 
<geoCoordinate id="EE0F4AC52A547610FBB1860F2AB3F9A4" box="[1052,1173,1589,1615]" direction="south" orientation="latitude" pageId="1" pageNumber="18" precision="92" value="-33.575">33°34.5’S</geoCoordinate>
, 
<geoCoordinate id="EE0F4AC52A547610FB0C860F2B35F9A4" box="[1185,1299,1589,1615]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="925" value="151.26666">151°16’E</geoCoordinate>
, depth 
<quantity id="4CC381E72A547610FAC5860F2BBEF9A4" box="[1384,1432,1589,1615]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="18" unit="m" value="4.0">4 m</quantity>
, sandy mud, coll. A. Jones 
<emphasis id="B94FF0102A547610FE7C86672C2BF99D" box="[465,525,1629,1654]" italics="true" pageId="1" pageNumber="18">et al.</emphasis>
, 
<date id="FF850AC22A547610FDB086672C98F99C" box="[541,702,1629,1655]" pageId="1" pageNumber="18" value="1981-05-26">26 May 1981</date>
; 
<specimenCount id="9D3DE78B2A547610FD6686672D13F99C" box="[715,821,1629,1655]" pageId="1" pageNumber="18" type="female">1 female</specimenCount>
, CL 
<quantity id="4CC381E72A547610FCC286672DEBF99C" box="[879,973,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="18" unit="mm" value="2.3">2.3 mm</quantity>
, TL ca. 
<quantity id="4CC381E72A547610FB8286672AABF99C" box="[1071,1165,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="18" unit="mm" value="6.5">6.5 mm</quantity>
, AM P62763, between Juno Head and Hungry Beach, Hawkesbury River, 
<collectionCode id="ED2AB4C72A547610FD5686BF2D1BF974" LSID="urn:lsid:biocol.org:col:15627" box="[763,829,1669,1695]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="1" pageNumber="18">NSW</collectionCode>
, 
<collectingCountry id="F32C6C922A547610FCE786BF2D9EF974" box="[842,952,1669,1695]" name="Australia" pageId="1" pageNumber="18">Australia</collectingCountry>
, 
<geoCoordinate id="EE0F4AC52A547610FC6B86BF2A19F974" box="[966,1087,1669,1695]" direction="south" orientation="latitude" pageId="1" pageNumber="18" precision="92" value="-33.575">33°34.5’S</geoCoordinate>
, 
<geoCoordinate id="EE0F4AC52A547610FBE086BF2AE6F974" box="[1101,1216,1669,1695]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="925" value="151.26666">151°16’E</geoCoordinate>
, depth 
<quantity id="4CC381E72A547610FAB586BF2B6CF974" box="[1304,1354,1669,1695]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="18" unit="m" value="4.0">4 m</quantity>
, sandy mud, coll. A. Jones 
<emphasis id="B94FF0102A547610FE2D86972F9DF92D" box="[384,443,1709,1734]" italics="true" pageId="1" pageNumber="18">et al.</emphasis>
, 
<date id="FF850AC22A547610FE6786972C4FF92C" box="[458,617,1709,1735]" pageId="1" pageNumber="18" value="1983-05-27">27 May 1983</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A547615FF6B86EF2DCFFE19" lastPageId="4" lastPageNumber="21" pageId="1" pageNumber="18" type="description">
<paragraph id="8B842C022A547610FF6B86EF2D1EF85C" blockId="1.[151,1437,1309,1975]" pageId="1" pageNumber="18">
<emphasis id="B94FF0102A547610FF6B86EF2F71F904" bold="true" box="[198,343,1749,1775]" pageId="1" pageNumber="18">Description</emphasis>
: Body elongate, slender. Carapace glabrous, not setose. Rostrum laterally compressed, slightly descending, with subacute tip, not reaching mid-length of second segment of antennular peduncle (
<figureCitation id="130030872A547610FF32871F2EC5F8D4" box="[159,227,1829,1855]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
A), proximally broadened; rostral carina slight, posteriorly not reaching rostral base (
<figureCitation id="130030872A547610FB73871F2B04F8D4" box="[1246,1314,1829,1855]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
B). Extracorneal spine acute, triangular, not reaching anterior margin of cornea (
<figureCitation id="130030872A547610FC4B87772A0FF88C" box="[998,1065,1869,1895]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
A, B); infra-corneal and supracorneal spine absent. Eyes largely exposed in dorsal and lateral views; cornea well pigmented (
<figureCitation id="130030872A547610FAA7874F2B6BF864" box="[1290,1357,1909,1935]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
A, B). Pterygostomial region rounded, not protruding anteriorly.
</paragraph>
<caption id="DF447C8A2A577613FF3A86F42B38F89B" httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="2" pageNumber="19" targetBox="[185,1378,207,1704]" targetPageId="2">
<paragraph id="8B842C022A577613FF3A86F42B38F89B" blockId="2.[151,1436,1742,1904]" pageId="2" pageNumber="19">
<emphasis id="B94FF0102A577613FF3A86F42F33F90D" bold="true" box="[151,277,1742,1766]" pageId="2" pageNumber="19">FIGURE 1</emphasis>
. 
<taxonomicName id="4C3B57812A577613FE8886F52C22F90D" box="[293,516,1743,1766]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A577613FE8886F52C22F90D" box="[293,516,1743,1766]" italics="true" pageId="2" pageNumber="19">Athanas sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A577613FDA386F42C6AF90E" box="[526,588,1742,1765]" pageId="2" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
, female, holotype, CL 4.8 mm, AM P62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D–H, J, K = 1.0 mm; C, H = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B842C022A567612FF6A80A22CF7FE74" blockId="3.[151,1437,152,1088]" pageId="3" pageNumber="20">
Antennular peduncle with second segment subequal to dorsally visible portion of first segment and equal to third segment (
<figureCitation id="130030872A567612FEC480852F8BFF32" box="[361,429,191,217]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
B, C); stylocerite acute, reaching mid-length of second segment; ventromesial carina with well developed, acute spine; lateral flagellum biramous, fused portion composed of about five segments (
<figureCitation id="130030872A567612FF3281342ECFFEC3" box="[159,233,270,296]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
B). Antenna with basicerite bearing acute ventrolateral spine; scaphocerite exceeding antennular peduncle (
<figureCitation id="130030872A567612FEBC810C2F73FEBB" box="[273,341,310,336]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
B, C), oval, lateral margin straight, anterior margin broadly convex, reaching beyond distlateral spine (
<figureCitation id="130030872A567612FEB781642F78FE93" box="[282,350,350,376]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
C); carpocerite exceeding scaphocerite by 1/5 of scaphocerite length (
<figureCitation id="130030872A567612FB3381642AC7FE93" box="[1182,1249,350,376]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
A, C), reaching far beyond distal margin of antennular peduncle.
</paragraph>
<paragraph id="8B842C022A567612FF6A81972C2FFDFD" blockId="3.[151,1437,152,1088]" pageId="3" pageNumber="20">Mouthparts without specific features, typical for genus. Third maxilliped slender, ultimate segment 1.5 times as long as penultimate segment, about half as long as antepenultimate segment; tip of ultimate segment setose, without spiniform setae.</paragraph>
<paragraph id="8B842C022A567612FF6A821E2BB1FCB8" blockId="3.[151,1437,152,1088]" pageId="3" pageNumber="20">
Female chelipeds subsymmetrical in shape, subequal in size (
<figureCitation id="130030872A567612FC15821E2A21FDD5" box="[952,1031,548,574]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Figs. 1</figureCitation>
D–G, 2 A, B); ischium elongate, almost as long as merus, ventral margin irregularly toothed, distally projecting, dorsal margin proximally with group of three to five slender spiniform setae (
<figureCitation id="130030872A567612FD1882492CD0FD66" box="[693,758,627,653]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
D–G); merus elongate, ventromesially slightly depressed, ventrolateral margin irregularly toothed; ventral surface furnished with numerous setae; carpus subequal to merus, distally widening, unarmed, setose ventrally; chela laterally compressed, ventral margin of palm concave at pollex base; fingers slightly shorter than palm, more or less gaping, tips crossing, cutting edges armed proximally with broadly triangular teeth, distally unarmed or armed with inconspicuous, irregular teeth, one chela of larger individuals with pronounced teeth on dactylus and pollex (
<figureCitation id="130030872A567612FC5B83032A11FCB8" box="[1014,1079,825,851]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
D). Male chelipeds unknown.
</paragraph>
<paragraph id="8B842C022A567612FF6A835B2D39FBAB" blockId="3.[151,1437,152,1088]" pageId="3" pageNumber="20">
Second pereiopod slender (
<figureCitation id="130030872A567612FDA9835B2C63FC90" box="[516,581,865,891]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
H); ischium slightly shorter than merus; carpus five-segmented, first article longer than four others combined; ratio of carpal segments (from proximal to distal) approximately: 8: 1: 1: 1: 3; chela simple, slightly longer than distal carpal article, fingers longer than palm. Third pereiopod slen- der (
<figureCitation id="130030872A567612FF6083ED2F29FC1A" box="[205,271,983,1009]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
G); ischium with two ventrolateral spiniform setae, dorsal margin with small distal spiniform seta; merus, carpus and propodus unarmed, merus about eight to ten times as long as broad; dactylus about 0.6 times propodus length, simple, slender, slightly curved.
</paragraph>
<caption id="DF447C8A2A567612FF3A87842AEAF81E" httpUri="https://zenodo.org/record/178385/files/figure.png" pageId="3" pageNumber="20" targetBox="[173,1388,1144,1951]" targetPageId="3">
<paragraph id="8B842C022A567612FF3A87842AEAF81E" blockId="3.[151,1436,1982,2037]" pageId="3" pageNumber="20">
<emphasis id="B94FF0102A567612FF3A87842F33F83D" bold="true" box="[151,277,1982,2006]" pageId="3" pageNumber="20">FIGURE 2</emphasis>
. 
<taxonomicName id="4C3B57812A567612FE8B87842C23F83E" box="[294,517,1982,2005]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A567612FE8B87842C23F83E" box="[294,517,1982,2005]" italics="true" pageId="3" pageNumber="20">Athanas sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A567612FDA387842C6AF83E" box="[526,588,1982,2005]" pageId="3" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
, female, paratype, CL 3.2 mm, AM P62762, Hawkesbury River, New South Wales, Australia. A – left cheliped, lateral view; B – right cheliped, lateral view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B842C022A517615FF6A80A22A5EFEB9" blockId="4.[151,1436,152,818]" pageId="4" pageNumber="21">
Pleura of first to fifth abdominal somites with rounded posteroventral angles; sixth somite with articulated triangular flap posteroventrally (
<figureCitation id="130030872A517615FDB780FA2C79FF31" box="[538,607,192,218]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="4" pageNumber="21">Fig. 1</figureCitation>
K). Uropod with protopod (sympodite) bearing distally two subacute lobes (
<figureCitation id="130030872A517615FF4B80D22F0CFEE9" box="[230,298,232,258]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="4" pageNumber="21">Fig. 1</figureCitation>
J); diaeresis straight, distolateral spiniform seta small. Telson (
<figureCitation id="130030872A517615FBBE80D22A70FEE9" box="[1043,1110,232,258]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="4" pageNumber="21">Fig. 1</figureCitation>
J) widest at proximal third, distally tapering, with two pairs of slender dorsal spiniform setae; posterior margin medially convex, with two pairs of slender posterolateral spiniform setae, mesial almost twice as long as lateral.
</paragraph>
<paragraph id="8B842C022A517615FF6A815A2C81FE21" blockId="4.[151,1436,152,818]" pageId="4" pageNumber="21">Gill formula typical for genus: pleurobranchs above P1–P5; podobranch and arthrobranch absent; straplike epipods (mastigobranchs) on coxae of Mxp3 and P1–P3; setobranchs on coxae of P1–P4; exopods on Mxp1–Mxp3, rudimentary exopods on P1–2.</paragraph>
<paragraph id="8B842C022A517615FF6A81E22DCFFE19" blockId="4.[151,1436,152,818]" box="[199,1001,472,498]" pageId="4" pageNumber="21">
Colour in life unknown. Size: CL 
<quantity id="4CC381E72A517615FDFB81E22CC0FE19" box="[598,742,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.8" metricValueMin="2.2" pageId="4" pageNumber="21" unit="mm" value="3.5" valueMax="4.8" valueMin="2.2">2.2–4.8 mm</quantity>
, TL ca. 6.0–12.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A517615FF6A81C52BB0FDF1" box="[199,1430,511,538]" pageId="4" pageNumber="21" type="etymology">
<paragraph id="8B842C022A517615FF6A81C52BB0FDF1" blockId="4.[151,1436,152,818]" box="[199,1430,511,538]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF6A81C52F68FDF2" bold="true" box="[199,334,511,537]" pageId="4" pageNumber="21">Etymology</emphasis>
: The new species is named after the Australian city of Sydney, situated near the 
<typeStatus id="548092A02A517615FB52823A2B14FDF1" box="[1279,1330,512,538]" pageId="4" pageNumber="21">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A517615FF6A821D2B32FD81" pageId="4" pageNumber="21" type="materials_examined">
<paragraph id="8B842C022A517615FF6A821D2A3EFDA9" blockId="4.[151,1436,152,818]" box="[199,1048,551,578]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF6A821D2F41FDAA" bold="true" box="[199,359,551,577]" pageId="4" pageNumber="21">
<typeStatus id="548092A02A517615FF6A821D2F22FDAA" box="[199,260,551,577]" pageId="4" pageNumber="21">Type</typeStatus>
locality
</emphasis>
: Hawkesbury River, Sydney, New South 
<collectingCountry id="F32C6C922A517615FCE282122DBFFDA9" box="[847,921,552,578]" name="United Kingdom" pageId="4" pageNumber="21">Wales</collectingCountry>
, 
<collectingCountry id="F32C6C922A517615FC0982122A35FDA9" box="[932,1043,552,578]" name="Australia" pageId="4" pageNumber="21">Australia</collectingCountry>
.
</paragraph>
<paragraph id="8B842C022A517615FF6A82752B32FD81" blockId="4.[151,1436,152,818]" box="[199,1300,591,618]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF6A82752F01FD82" bold="true" box="[199,295,591,617]" pageId="4" pageNumber="21">Habitat</emphasis>
: All specimens were dredged from sand–mud bottoms at depths between 4 and 
<specimenCount id="9D3DE78B2A517615FB7C826A2B32FD81" box="[1233,1300,592,618]" pageId="4" pageNumber="21" type="generic">
<quantity id="4CC381E72A517615FB7C826A2B36FD81" box="[1233,1296,592,618]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="21" unit="m" value="10.0">10 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A517615FF6A824D2AAAFD79" box="[199,1164,631,658]" pageId="4" pageNumber="21" type="distribution">
<paragraph id="8B842C022A517615FF6A824D2AAAFD79" blockId="4.[151,1436,152,818]" box="[199,1164,631,658]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF6A824D2F46FD7A" bold="true" box="[199,352,631,657]" pageId="4" pageNumber="21">Distribution</emphasis>
: Presently known only from the 
<typeStatus id="548092A02A517615FD4F82422D33FD79" box="[738,789,632,658]" pageId="4" pageNumber="21">type</typeStatus>
locality near Sydney, 
<collectingCountry id="F32C6C922A517615FBB482422AAFFD79" box="[1049,1161,632,658]" name="Australia" pageId="4" pageNumber="21">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A517614FF6A82A52AA0FE91" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21" type="discussion">
<paragraph id="8B842C022A517615FF6A82A52DD4FCD9" blockId="4.[151,1436,152,818]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF6A82A52F1FFD52" bold="true" box="[199,313,671,697]" pageId="4" pageNumber="21">Remarks</emphasis>
: 
<taxonomicName id="4C3B57812A517615FEE4829A2C1AFD52" box="[329,572,672,697]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FEE4829A2C1AFD52" box="[329,572,672,697]" italics="true" pageId="4" pageNumber="21">Athanas sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FDE8829A2CA0FD51" box="[581,646,672,698]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
, is most closely related to 
<taxonomicName id="4C3B57812A517615FC68829A2BBAFD51" authority="Banner &amp; Banner, 1983" authorityName="Banner &amp; Banner" authorityYear="1983" box="[965,1436,672,698]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FC68829A2A5FFD52" box="[965,1145,672,697]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
<bibRefCitation id="EFAA51F32A517615FB2F829A2BBAFD51" author="Banner" box="[1154,1436,672,698]" pageId="4" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner, 1983</bibRefCitation>
</taxonomicName>
presently known only from the 
<typeStatus id="548092A02A517615FDA682F22C1BFD09" box="[523,573,712,738]" pageId="4" pageNumber="21">type</typeStatus>
series of 
<specimenCount id="9D3DE78B2A517615FD1C82F22D3CFD09" box="[689,794,712,738]" pageId="4" pageNumber="21" type="male">13 males</specimenCount>
and two ovigerous females, dredged from 
<quantity id="4CC381E72A517615FABA82F22BBAFD09" box="[1303,1436,712,738]" metricMagnitude="2" metricUnit="m" metricValue="3.975" metricValueMax="4.5" metricValueMin="3.45" pageId="4" pageNumber="21" unit="m" value="397.5" valueMax="450.0" valueMin="345.0">345–450 m</quantity>
off 
<collectingCountry id="F32C6C922A517615FF6D82CA2F6CFCE1" box="[192,330,752,778]" name="Réunion" pageId="4" pageNumber="21">La Réunion</collectingCountry>
in the southwestern Indian Ocean (
<bibRefCitation id="EFAA51F32A517615FD4782CA2A24FCE1" author="Banner" box="[746,1026,752,778]" pageId="4" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
). However, the new species differs in several characteristics from 
<taxonomicName id="4C3B57812A517615FE5383222C97FCDA" box="[510,689,792,817]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FE5383222C97FCDA" box="[510,689,792,817]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
, as summarized in 
<tableCitation id="C6B919B92A517615FC3883222DCAFCD9" box="[917,1004,792,818]" captionStart="TABLE 1" captionStartId="4.[151,239,871,895]" captionTargetBox="[159,1408,928,1524]" captionTargetPageId="4" captionText="TABLE 1. Differences between Athanas phyllocheles Banner &amp; Banner, 1983 and A. sydneyensis n. sp." httpUri="http://table.plazi.org/id/DF447C8A2A517615FF3A835D2ACFFC94" pageId="4" pageNumber="21" tableUuid="DF447C8A2A517615FF3A835D2ACFFC94">Table 1</tableCitation>
.
</paragraph>
<caption id="DF447C8A2A517615FF3A835D2ACFFC94" ID-Table-UUID="DF447C8A2A517615FF3A835D2ACFFC94" box="[151,1257,871,895]" httpUri="http://table.plazi.org/id/DF447C8A2A517615FF3A835D2ACFFC94" pageId="4" pageNumber="21" targetBox="[159,1408,928,1524]" targetIsTable="true" targetPageId="4">
<paragraph id="8B842C022A517615FF3A835D2ACFFC94" blockId="4.[151,1257,871,895]" box="[151,1257,871,895]" pageId="4" pageNumber="21">
<emphasis id="B94FF0102A517615FF3A835D2F24FC94" bold="true" box="[151,258,871,895]" pageId="4" pageNumber="21">TABLE 1</emphasis>
. Differences between 
<taxonomicName id="4C3B57812A517615FE5F83522DFCFC94" authority="Banner &amp; Banner, 1983" authorityName="Banner &amp; Banner" authorityYear="1983" box="[498,986,872,895]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FE5F83522CF3FC94" box="[498,725,872,895]" italics="true" pageId="4" pageNumber="21">Athanas phyllocheles</emphasis>
Banner &amp; Banner, 1983
</taxonomicName>
and 
<taxonomicName id="4C3B57812A517615FBA383522A8CFC94" box="[1038,1194,872,895]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FBA383522A8CFC94" box="[1038,1194,872,895]" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FB1D83522ACFFC94" box="[1200,1257,872,895]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B842C022A517615FF32839A2A10FA1F" pageId="4" pageNumber="21">
<table id="F93BDEA22A5189EEFF32839A2BA6FA1F" box="[159,1408,928,1524]" gridcols="3" gridrows="12" pageId="4" pageNumber="21">
<tr id="350B2E402A5189EEFF32839A2BA6FC5C" box="[159,1408,928,951]" gridrow="0" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF32839A2FD6FC5C" box="[159,496,928,951]" gridcol="0" gridrow="0" pageId="4" pageNumber="21">Features / Species</th>
<th id="76DA473C2A5189EEFDB2839A2DEDFC5C" box="[543,971,928,951]" gridcol="1" gridrow="0" pageId="4" pageNumber="21">
<taxonomicName id="4C3B57812A517615FDB2839A2CE7FC5C" box="[543,705,928,951]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FDB2839A2CE7FC5C" box="[543,705,928,951]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
</th>
<th id="76DA473C2A5189EEFC48839A2BA6FC5C" box="[997,1408,928,951]" gridcol="2" gridrow="0" pageId="4" pageNumber="21">
<taxonomicName id="4C3B57812A517615FC4B839A2AA4FC5C" box="[998,1154,928,951]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FC4B839A2AA4FC5C" box="[998,1154,928,951]" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FB25839A2AE7FC5C" box="[1160,1217,928,951]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr id="350B2E402A5189EEFF3283F02BA6FC0A" box="[159,1408,970,993]" gridrow="1" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3283F02FD6FC0A" box="[159,496,970,993]" gridcol="0" gridrow="1" pageId="4" pageNumber="21">1. Rostrum lateral view</th>
<td id="76DA473C2A5189EEFDB283F02DEDFC0A" box="[543,971,970,993]" gridcol="1" gridrow="1" pageId="4" pageNumber="21">slender, with tip slightly ascending</td>
<td id="76DA473C2A5189EEFC4883F02BA6FC0A" box="[997,1408,970,993]" gridcol="2" gridrow="1" pageId="4" pageNumber="21">more robust, tip slightly descending</td>
</tr>
<tr id="350B2E402A5189EEFF3283CF2BA6FBE7" box="[159,1408,1013,1036]" gridrow="2" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3283CF2FD6FBE7" box="[159,496,1013,1036]" gridcol="0" gridrow="2" pageId="4" pageNumber="21">2. Rostrum, dorsal view</th>
<td id="76DA473C2A5189EEFDB283CF2DEDFBE7" box="[543,971,1013,1036]" gridcol="1" gridrow="2" pageId="4" pageNumber="21">proximally not broadened</td>
<td id="76DA473C2A5189EEFC4883CF2BA6FBE7" box="[997,1408,1013,1036]" gridcol="2" gridrow="2" pageId="4" pageNumber="21">proximally broadened, convex</td>
</tr>
<tr id="350B2E402A5189EEFF32841A2BA6FBDC" box="[159,1408,1056,1079]" gridrow="3" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF32841A2FD6FBDC" box="[159,496,1056,1079]" gridcol="0" gridrow="3" pageId="4" pageNumber="21">3. Extra-corneal spines</th>
<td id="76DA473C2A5189EEFDB2841A2DEDFBDC" box="[543,971,1056,1079]" gridcol="1" gridrow="3" pageId="4" pageNumber="21">reaching anterior margin of eyes</td>
<td id="76DA473C2A5189EEFC48841A2BA6FBDC" box="[997,1408,1056,1079]" gridcol="2" gridrow="3" pageId="4" pageNumber="21">not reaching anterior margin of eyes</td>
</tr>
<tr id="350B2E402A5189EEFF3284702BA6FB6F" box="[159,1408,1098,1156]" gridrow="4" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3284702FD6FB6F" box="[159,496,1098,1156]" gridcol="0" gridrow="4" pageId="4" pageNumber="21">4. Anterolateral margin of cara- pace (antennal region)</th>
<td id="76DA473C2A5189EEFDB284702DEDFB6F" box="[543,971,1098,1156]" gridcol="1" gridrow="4" pageId="4" pageNumber="21">concave</td>
<td id="76DA473C2A5189EEFC4884702BA6FB6F" box="[997,1408,1098,1156]" gridcol="2" gridrow="4" pageId="4" pageNumber="21">convex</td>
</tr>
<tr id="350B2E402A5189EEFF3284A22BA6FB3A" box="[159,1408,1176,1233]" gridrow="5" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3284A22FD6FB3A" box="[159,496,1176,1233]" gridcol="0" gridrow="5" pageId="4" pageNumber="21">5. Stylocerite</th>
<td id="76DA473C2A5189EEFDB284A22DEDFB3A" box="[543,971,1176,1233]" gridcol="1" gridrow="5" pageId="4" pageNumber="21">slightly exceeding distal margin of first segment of antennular peduncle</td>
<td id="76DA473C2A5189EEFC4884A22BA6FB3A" box="[997,1408,1176,1233]" gridcol="2" gridrow="5" pageId="4" pageNumber="21">reaching almost middle of second seg- ment of antennular peduncle</td>
</tr>
<tr id="350B2E402A5189EEFF3284DF2BA6FB17" box="[159,1408,1253,1276]" gridrow="6" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3284DF2FD6FB17" box="[159,496,1253,1276]" gridcol="0" gridrow="6" pageId="4" pageNumber="21">6. Scaphocerite</th>
<td id="76DA473C2A5189EEFDB284DF2DEDFB17" box="[543,971,1253,1276]" gridcol="1" gridrow="6" pageId="4" pageNumber="21">not reaching end of antennular peduncle</td>
<td id="76DA473C2A5189EEFC4884DF2BA6FB17" box="[997,1408,1253,1276]" gridcol="2" gridrow="6" pageId="4" pageNumber="21">clearly exceeding antennular peduncle</td>
</tr>
<tr id="350B2E402A5189EEFF32852A2BA6FACC" box="[159,1408,1296,1319]" gridrow="7" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF32852A2FD6FACC" box="[159,496,1296,1319]" gridcol="0" gridrow="7" pageId="4" pageNumber="21">7. Major cheliped, carpus</th>
<td id="76DA473C2A5189EEFDB2852A2DEDFACC" box="[543,971,1296,1319]" gridcol="1" gridrow="7" pageId="4" pageNumber="21">shorter than merus</td>
<td id="76DA473C2A5189EEFC48852A2BA6FACC" box="[997,1408,1296,1319]" gridcol="2" gridrow="7" pageId="4" pageNumber="21">subequal to merus</td>
</tr>
<tr id="350B2E402A5189EEFF3285002BA6FABA" box="[159,1408,1338,1361]" gridrow="8" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3285002FD6FABA" box="[159,496,1338,1361]" gridcol="0" gridrow="8" pageId="4" pageNumber="21">8. Third pereiopod, propodus</th>
<td id="76DA473C2A5189EEFDB285002DEDFABA" box="[543,971,1338,1361]" gridcol="1" gridrow="8" pageId="4" pageNumber="21">almost 15 times as long as broad</td>
<td id="76DA473C2A5189EEFC4885002BA6FABA" box="[997,1408,1338,1361]" gridcol="2" gridrow="8" pageId="4" pageNumber="21">8–10 times as long as broad</td>
</tr>
<tr id="350B2E402A5189EEFF32855F2BA6FA97" box="[159,1408,1381,1404]" gridrow="9" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF32855F2FD6FA97" box="[159,496,1381,1404]" gridcol="0" gridrow="9" pageId="4" pageNumber="21">9. Third pereiopod</th>
<td id="76DA473C2A5189EEFDB2855F2DEDFA97" box="[543,971,1381,1404]" gridcol="1" gridrow="9" pageId="4" pageNumber="21">dactylus / propodus ratio 1/3</td>
<td id="76DA473C2A5189EEFC48855F2BA6FA97" box="[997,1408,1381,1404]" gridcol="2" gridrow="9" pageId="4" pageNumber="21">dactylus / propodus ratio 1/2</td>
</tr>
<tr id="350B2E402A5189EEFF3285AA2BA6FA22" box="[159,1408,1424,1481]" gridrow="10" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3285AA2FD6FA22" box="[159,496,1424,1481]" gridcol="0" gridrow="10" pageId="4" pageNumber="21">10. Third maxilliped, dactylus: propodus ratio</th>
<td id="76DA473C2A5189EEFDB285AA2DEDFA22" box="[543,971,1424,1481]" gridcol="1" gridrow="10" pageId="4" pageNumber="21">~1.3</td>
<td id="76DA473C2A5189EEFC4885AA2BA6FA22" box="[997,1408,1424,1481]" gridcol="2" gridrow="10" pageId="4" pageNumber="21">~1.5</td>
</tr>
<tr id="350B2E402A5189EEFF3285E72BA6FA1F" box="[159,1408,1501,1524]" gridrow="11" pageId="4" pageNumber="21">
<th id="76DA473C2A5189EEFF3285E72FD6FA1F" box="[159,496,1501,1524]" gridcol="0" gridrow="11" pageId="4" pageNumber="21">10. Bathymetry</th>
<td id="76DA473C2A5189EEFDB285E72DEDFA1F" box="[543,971,1501,1524]" gridcol="1" gridrow="11" pageId="4" pageNumber="21">345–450 m</td>
<td id="76DA473C2A5189EEFC4885E72BA6FA1F" box="[997,1408,1501,1524]" gridcol="2" gridrow="11" pageId="4" pageNumber="21">6–10 m</td>
</tr>
</table>
</paragraph>
<paragraph id="8B842C022A517615FF6A86272C99F804" blockId="4.[151,1436,1565,2031]" pageId="4" pageNumber="21">
We believe that some of these differences, e.g., the proportion of the articles in the third pereiopod (8, 9), the length of the scaphocerite compared to that of the antennular peduncle (6), the carpus/merus ratio in the chelipeds (7), and the relative lengths of the dactylus and propodus of the third maxilliped (10) are taxonomically reliable. In view of the intraspecific polymorphism of the chelipeds reported in 
<taxonomicName id="4C3B57812A517615FB2F86AF2F7FF93C" authority="Banner &amp; Banner 1983" authorityName="Banner &amp; Banner" authorityYear="1983" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FB2F86AF2B13F945" box="[1154,1333,1685,1710]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
(
<bibRefCitation id="EFAA51F32A517615FAE886AF2F69F93C" author="Banner" pageId="4" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
)
</taxonomicName>
, other differences on these appendages appear to be less significant. The chelipeds of 
<taxonomicName id="4C3B57812A517615FAE786872EDDF915" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FAE786872EDDF915" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FEAE86DF2F63F914" box="[259,325,1765,1791]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
, are somewhat “intermediate” between the chelipeds of the smaller male 
<typeStatus id="548092A02A517615FB1D86DF2B33F914" box="[1200,1301,1765,1791]" pageId="4" pageNumber="21" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C3B57812A517615FA9386DF2ED1F8CD" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FA9386DF2ED1F8CD" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
with TL 
<quantity id="4CC381E72A517615FECE87372F9AF8CC" box="[355,444,1805,1831]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="4" pageNumber="21" unit="mm" value="8.7">8.7 mm</quantity>
(
<bibRefCitation id="EFAA51F32A517615FE6787372CF8F8CC" author="Banner" box="[458,734,1805,1831]" pageId="4" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
, fig. 14 l, m) and the juvenile 
<typeStatus id="548092A02A517615FB9687372A86F8CC" box="[1083,1184,1805,1831]" pageId="4" pageNumber="21" type="paratype">paratype</typeStatus>
of unknown sex with TL 
<quantity id="4CC381E72A517615FF6E870F2F38F8A4" box="[195,286,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="21" unit="mm" value="6.5">6.5 mm</quantity>
(
<emphasis id="B94FF0102A517615FE9D870F2F4FF8A5" box="[304,361,1845,1870]" italics="true" pageId="4" pageNumber="21">idem</emphasis>
., fig. 14 n, o). At TL 12.0 mm, however, the 
<typeStatus id="548092A02A517615FC2A870F2DCBF8A4" box="[903,1005,1845,1871]" pageId="4" pageNumber="21" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C3B57812A517615FBBB870F2AE4F8A5" box="[1046,1218,1845,1870]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FBBB870F2AE4F8A5" box="[1046,1218,1845,1870]" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FB66870F2B2DF8A4" box="[1227,1291,1845,1871]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
is considerably larger than the aforementioned specimens of 
<taxonomicName id="4C3B57812A517615FD4E87672DB0F89D" box="[739,918,1885,1910]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FD4E87672DB0F89D" box="[739,918,1885,1910]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
. Although both 
<taxonomicName id="4C3B57812A517615FBFA87672B24F89D" box="[1111,1282,1885,1910]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FBFA87672B24F89D" box="[1111,1282,1885,1910]" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FAA787672B6FF89C" box="[1290,1353,1885,1911]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
and 
<taxonomicName id="4C3B57812A517615FA2E87672F0CF875" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FA2E87672F0CF875" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
live on soft bottoms, the bathymetry is very different: 
<taxonomicName id="4C3B57812A517615FC0787BF2A73F875" box="[938,1109,1925,1950]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A517615FC0787BF2A73F875" box="[938,1109,1925,1950]" italics="true" pageId="4" pageNumber="21">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A517615FBF187BF2ABDF874" box="[1116,1179,1925,1951]" pageId="4" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
was collected in relatively shallow water (
<quantity id="4CC381E72A517615FE3587972FD2F82C" box="[408,500,1965,1991]" metricMagnitude="0" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="4" pageNumber="21" unit="m" value="8.0" valueMax="10.0" valueMin="6.0">6–10 m</quantity>
) of an estuary, whereas 
<taxonomicName id="4C3B57812A517615FCB987972DEFF82D" box="[788,969,1965,1990]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A517615FCB987972DEFF82D" box="[788,969,1965,1990]" italics="true" pageId="4" pageNumber="21">A. phyllocheles</emphasis>
</taxonomicName>
was collected from much deeper water (
<quantity id="4CC381E72A517615FF3287EF2F00F804" box="[159,294,2005,2031]" metricMagnitude="2" metricUnit="m" metricValue="3.975" metricValueMax="4.5" metricValueMin="3.45" pageId="4" pageNumber="21" unit="m" value="397.5" valueMax="450.0" valueMin="345.0">345–450 m</quantity>
) off the coast of an oceanic island.
</paragraph>
<paragraph id="8B842C022A507614FF6A80A22AA0FE91" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="22">
Several other species of 
<taxonomicName id="4C3B57812A507614FE5980A22C72FF5A" box="[500,596,152,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B94FF0102A507614FE5980A22C72FF5A" box="[500,596,152,177]" italics="true" pageId="5" pageNumber="22">Athanas</emphasis>
</taxonomicName>
are characterized by extremely slender, scythe-shaped dactyli on the third to fifth pereiopods. However, only three of them appear to be more closely related to 
<taxonomicName id="4C3B57812A507614FB7C80FA2B5BFF32" box="[1233,1405,192,217]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FB7C80FA2B5BFF32" box="[1233,1405,192,217]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FA2B80FA2E9DFEE9" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
: 
<taxonomicName id="4C3B57812A507614FF6480D22CA0FEE9" authority="Banner &amp; Banner, 1978" authorityName="Banner &amp; Banner" authorityYear="1978" box="[201,646,232,258]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FF6480D22F46FEEA" box="[201,352,232,257]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
<bibRefCitation id="EFAA51F32A507614FEC480D22CA0FEE9" author="Banner" box="[361,646,232,258]" pageId="5" pageNumber="29" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner, 1978</bibRefCitation>
</taxonomicName>
, dredged from 
<quantity id="4CC381E72A507614FC9180D22D9BFEE9" box="[828,957,232,258]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="3.85" metricValueMin="3.65" pageId="5" pageNumber="22" unit="m" value="375.0" valueMax="385.0" valueMin="365.0">365-385 m</quantity>
in the South 
<collectingCountry id="F32C6C922A507614FBF380D22A82FEE9" box="[1118,1188,232,258]" name="China" pageId="5" pageNumber="22">China</collectingCountry>
Sea, off 
<collectingCountry id="F32C6C922A507614FABC80D22BBAFEE9" box="[1297,1436,232,258]" name="Hong Kong" pageId="5" pageNumber="22">Hong Kong</collectingCountry>
(
<bibRefCitation id="EFAA51F32A507614FF32812A2F9AFEC1" author="Banner" box="[159,444,272,298]" pageId="5" pageNumber="29" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
); 
<taxonomicName id="4C3B57812A507614FE7E812A2C7EFEC2" box="[467,600,272,297]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FE7E812A2C7EFEC2" box="[467,600,272,297]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
de Man, 1910, collected from 
<quantity id="4CC381E72A507614FC62812A2A2AFEC1" box="[975,1036,272,298]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="22" unit="m" value="72.0">72 m</quantity>
off Sulawesi, 
<collectingCountry id="F32C6C922A507614FB11812A2B08FEC1" box="[1212,1326,272,298]" name="Indonesia" pageId="5" pageNumber="22">Indonesia</collectingCountry>
(de 
<bibRefCitation id="EFAA51F32A507614FACB812A2EF3FEB9" author="Man" pageId="5" pageNumber="30" refString="Man, J. G de, (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1, 133 - 465; 1915, Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden." type="journal article" year="1911">Man 1911</bibRefCitation>
); and 
<taxonomicName id="4C3B57812A507614FE8C81022C4FFEB9" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" box="[289,617,312,338]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FE8C81022F8BFEBA" box="[289,429,312,337]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
<bibRefCitation id="EFAA51F32A507614FE1A81022C4FFEB9" author="Holthuis" box="[439,617,312,338]" pageId="5" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
</taxonomicName>
known from several localities in the eastern Atlantic, including the Mediterranean Sea (
<bibRefCitation id="EFAA51F32A507614FE2F815A2C08FE91" author="Holthuis" box="[386,558,352,378]" pageId="5" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
; 
<bibRefCitation id="EFAA51F32A507614FD97815A2D75FE91" author="Crosnier" box="[570,851,352,378]" pageId="5" pageNumber="29" refString="Crosnier, A. &amp; Forest, J. 1973. Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris. 409 pp." type="book" year="1973">Crosnier &amp; Forest 1973</bibRefCitation>
; 
<bibRefCitation id="EFAA51F32A507614FCF2815A2A5FFE91" author="Froglia" box="[863,1145,352,378]" pageId="5" pageNumber="29" refString="Froglia, C &amp; Argenti, L. (1993) Athanas amazone Holthuis, 1951, first record for the Italian seas (Crustacea, Decapoda, Alpheidae). Bios (Macedonia, Greece), Scientific Annals of the School of Biology, 1, 201 - 205." type="journal article" year="1993">Froglia &amp; Argenti 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A507617FF6A81B22F24FF31" lastPageId="6" lastPageNumber="23" pageId="5" pageNumber="22" type="reference_group">
<paragraph id="8B842C022A507614FF6A81B22AF8FC41" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="22">
<taxonomicName id="4C3B57812A507614FF6A81B22F81FE4A" ID-CoL="85MBX" box="[199,423,392,417]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FF6A81B22F81FE4A" box="[199,423,392,417]" italics="true" pageId="5" pageNumber="22">Athanas gracilipes</emphasis>
</taxonomicName>
was described on the basis of two fragmentary specimens, both without chelipeds. Although 
<bibRefCitation id="EFAA51F32A507614FEA3818A2C11FE21" author="Banner" box="[270,567,432,458]" pageId="5" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
did not contrast 
<taxonomicName id="4C3B57812A507614FCAD818A2D93FE22" ID-CoL="85MCS" box="[768,949,432,457]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FCAD818A2D93FE22" box="[768,949,432,457]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
with 
<emphasis id="B94FF0102A507614FC57818A2ABFFE22" box="[1018,1177,432,457]" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C3B57812A507614FC57818A2AB2FE22" ID-CoL="85MBX" box="[1018,1172,432,457]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">A. gracilipes</taxonomicName>
,
</emphasis>
these two species are possibly closely related, and both are similar to 
<taxonomicName id="4C3B57812A507614FD6A81E22D57FE1A" ID-CoL="85MD5" box="[711,881,472,497]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FD6A81E22D57FE1A" box="[711,881,472,497]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FCD481E22D91FE19" box="[889,951,472,498]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
The only significant differences between 
<taxonomicName id="4C3B57812A507614FF3A823A2F0BFDF2" ID-CoL="85MBX" box="[151,301,512,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FF3A823A2F0BFDF2" box="[151,301,512,537]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
and the other two species are: (1) the slightly shorter rostrum, not reaching distal margin of the first segment of the antennular peduncle; (2) the presence of small, triangular infra-corneal spines; (3) the more slender second pereiopod; and (4) the different proportional lengths of articles in the third pereiopod, e.g., the dactylus being only half length of the propodus (instead of 3/ 
<quantity id="4CC381E72A507614FC6382422DDDFD79" box="[974,1019,632,658]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="22" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C3B57812A507614FBAE82422A88FD7A" ID-CoL="85MD5" box="[1027,1198,632,657]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FBAE82422A88FD7A" box="[1027,1198,632,657]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FB1B82422AD1FD79" box="[1206,1271,632,658]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
, and 1/ 
<quantity id="4CC381E72A507614FAE082422B5DFD79" box="[1357,1403,632,658]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="22" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C3B57812A507614FA2E82422F0CFD52" ID-CoL="85MCS" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FA2E82422F0CFD52" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
). In this last feature, and generally in the proportions of the articles of the third pereiopod, 
<taxonomicName id="4C3B57812A507614FAE7829A2EC2FD0A" ID-CoL="85MBX" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FAE7829A2EC2FD0A" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
is closer to 
<taxonomicName id="4C3B57812A507614FEDB82F22C04FD0A" ID-CoL="85MD5" box="[374,546,712,737]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FEDB82F22C04FD0A" box="[374,546,712,737]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C4D6B2A507614FD9F82F22C52FD09" box="[562,628,712,738]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
, differing from it mainly in (1) the presence of infra-corneal spines; (2) the shorter stylocerite, only slightly exceeding the distal margin of the first segment of the antennular peduncle; (3) the more slender antennular peduncle, with the second segment 1.5 times as long as broad (vs. 1.2 times in 
<taxonomicName id="4C3B57812A507614FE84837A2FF3FCB2" ID-CoL="85MD5" box="[297,469,832,857]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FE84837A2FF3FCB2" box="[297,469,832,857]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FE70837A2C38FCB1" box="[477,542,832,858]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
); (4) the shorter scaphocerite, not exceeding distal margin of the antennular peduncle; (5) the much more slender second pereiopod; (6) differently shaped telson, with lateral margins straight, not convex, as in 
<taxonomicName id="4C3B57812A507614FE6683AA2C50FC42" ID-CoL="85MD5" box="[459,630,912,937]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FE6683AA2C50FC42" box="[459,630,912,937]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FDD083AA2C9AFC41" box="[637,700,912,938]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
(cf. 
<figureCitation id="130030872A507614FD5E83AA2D11FC41" box="[755,823,912,938]" captionStart="FIGURE 1" captionStartId="2.[151,255,1742,1766]" captionTargetBox="[185,1378,207,1704]" captionTargetId="figure@2.[151,1436,197,1716]" captionTargetPageId="2" captionText="FIGURE 1. Athanas sydneyensis n. sp., female, holotype, CL 4.8 mm, AM P 62764, Hawkesbury River, New South Wales, Australia. A – frontal region, lateral view; B – same, dorsal view; C – right antennule and antenna, dorsal view; D – right cheliped, lateral view; E – merus and ischium of right cheliped, mesial view; F – left cheliped, lateral; G – merus and ischium of left cheliped, mesial view; H – right second pereiopod, lateral view; I – right third pereiopod, lateral view; J – tail fan, dorsal view; K – tailfan, right lateral view. Scale bar: A, B, D – H, J, K = 1.0 mm; C, H = 0.5 mm." httpUri="https://zenodo.org/record/178384/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
and 
<bibRefCitation id="EFAA51F32A507614FCDD83AA2AA5FC41" author="Banner" box="[880,1155,912,938]" pageId="5" pageNumber="29" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
, fig. 3).
</paragraph>
<paragraph id="8B842C022A507614FF6A83822CBCFA61" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="22">
<taxonomicName id="4C3B57812A507614FF6A83822FB7FC3A" ID-CoL="85MD6" box="[199,401,952,977]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FF6A83822FB7FC3A" box="[199,401,952,977]" italics="true" pageId="5" pageNumber="22">Athanas tenuipes</emphasis>
</taxonomicName>
is presently known only from the possibly juvenile 
<typeStatus id="548092A02A507614FC5183822A43FC39" box="[1020,1125,952,978]" pageId="5" pageNumber="22" type="holotype">holotype</typeStatus>
, which, like 
<emphasis id="B94FF0102A507614FB5383822BBAFC3A" box="[1278,1436,952,977]" italics="true" pageId="5" pageNumber="22">
<taxonomicName id="4C3B57812A507614FB5383822BB1FC3A" ID-CoL="85MBX" box="[1278,1431,952,977]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">A. gracilipes</taxonomicName>
,
</emphasis>
lacks both chelipeds. A complete specimen, possibly assignable to 
<taxonomicName id="4C3B57812A507614FC7F83DA2A7EFC12" ID-CoL="85MD6" box="[978,1112,992,1017]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FC7F83DA2A7EFC12" box="[978,1112,992,1017]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
, was recently collected in southern 
<collectingCountry id="F32C6C922A507614FEAF84322F43FBC9" box="[258,357,1032,1058]" name="Vietnam" pageId="5" pageNumber="22">Vietnam</collectingCountry>
(Anker &amp; Marin unpubl.). 
<taxonomicName id="4C3B57812A507614FD0F84322D4DFBCA" ID-CoL="85MD6" box="[674,875,1032,1057]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FD0F84322D4DFBCA" box="[674,875,1032,1057]" italics="true" pageId="5" pageNumber="22">Athanas tenuipes</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B57812A507614FBA984322A88FBCA" ID-CoL="85MD5" box="[1028,1198,1032,1057]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FBA984322A88FBCA" box="[1028,1198,1032,1057]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FB1884322AD3FBC9" box="[1205,1269,1032,1058]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
, but also from 
<taxonomicName id="4C3B57812A507614FF3A840A2F0BFBA2" ID-CoL="85MBX" box="[151,301,1072,1097]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FF3A840A2F0BFBA2" box="[151,301,1072,1097]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B57812A507614FEC5840A2C3AFBA2" ID-CoL="85MCS" box="[360,540,1072,1097]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FEC5840A2C3AFBA2" box="[360,540,1072,1097]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
by (1) the carpocerite being much shorter than the scaphocerite; (2) the lateral spine of the scaphocerite exceeding the anterior margin of the blade; (3) the telson being more slender, with a slightly emarginated posterior margin (cf. de 
<bibRefCitation id="EFAA51F32A507614FCAA84BA2DA4FB71" author="Man" box="[775,898,1152,1178]" pageId="5" pageNumber="30" refString="Man, J. G de, (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1, 133 - 465; 1915, Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden." type="journal article" year="1911">Man 1911</bibRefCitation>
, 1915); and possibly also several features on the chelipeds (A. Anker pers. obs.). Further, 
<taxonomicName id="4C3B57812A507614FD0484922D0AFB2A" ID-CoL="85MD6" box="[681,812,1192,1217]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FD0484922D0AFB2A" box="[681,812,1192,1217]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B57812A507614FC6684922A44FB2A" ID-CoL="85MBX" box="[971,1122,1192,1217]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FC6684922A44FB2A" box="[971,1122,1192,1217]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
by the much longer stylocerite, exceeding mid-length of the second segment of the antennular peduncle, and from 
<taxonomicName id="4C3B57812A507614FB6F84EA2B50FB02" ID-CoL="85MCS" box="[1218,1398,1232,1257]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FB6F84EA2B50FB02" box="[1218,1398,1232,1257]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
by the presence of small infra-corneal spines. 
<bibRefCitation id="EFAA51F32A507614FD2684C22D89FAF9" author="Banner" box="[651,943,1272,1298]" pageId="5" pageNumber="29" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner (1978)</bibRefCitation>
assumed that the 
<typeStatus id="548092A02A507614FBD284C22A94FAF9" box="[1151,1202,1272,1298]" pageId="5" pageNumber="22">type</typeStatus>
of 
<taxonomicName id="4C3B57812A507614FB7A84C22B7FFAFA" ID-CoL="85MD6" box="[1239,1369,1272,1297]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FB7A84C22B7FFAFA" box="[1239,1369,1272,1297]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
, with TL 
<quantity id="4CC381E72A507614FF6F851A2F2EFAD1" box="[194,264,1312,1338]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="22" unit="mm" value="7.0">7 mm</quantity>
, is immature, and that this may account for the slight differences between 
<taxonomicName id="4C3B57812A507614FBDD851A2AD4FAD2" ID-CoL="85MD6" box="[1136,1266,1312,1337]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FBDD851A2AD4FAD2" box="[1136,1266,1312,1337]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B57812A507614FA81851A2EE0FA8A" ID-CoL="85MBX" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FA81851A2EE0FA8A" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
. However, we feel that both 
<taxonomicName id="4C3B57812A507614FD8885722C9BFA8A" ID-CoL="85MBX" box="[549,701,1352,1377]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FD8885722C9BFA8A" box="[549,701,1352,1377]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B57812A507614FD5785722D58FA8A" ID-CoL="85MD6" box="[762,894,1352,1377]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B94FF0102A507614FD5785722D58FA8A" box="[762,894,1352,1377]" italics="true" pageId="5" pageNumber="22">A. tenuipes</emphasis>
</taxonomicName>
should be treated as valid species until more material of 
<taxonomicName id="4C3B57812A507614FEB3854A2F95FA62" ID-CoL="85MBX" box="[286,435,1392,1417]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B94FF0102A507614FEB3854A2F95FA62" box="[286,435,1392,1417]" italics="true" pageId="5" pageNumber="22">A. gracilipes</emphasis>
</taxonomicName>
becomes available.
</paragraph>
<paragraph id="8B842C022A507617FF6A85A22F24FF31" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,458]" lastPageId="6" lastPageNumber="23" pageId="5" pageNumber="22">
The eastern Atlantic 
<taxonomicName id="4C3B57812A507614FE6F85A22D21FA59" ID-CoL="85MBL" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" box="[450,775,1432,1458]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FE6F85A22C6AFA5A" box="[450,588,1432,1457]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
<bibRefCitation id="EFAA51F32A507614FDFB85A22D21FA59" author="Holthuis" box="[598,775,1432,1458]" pageId="5" pageNumber="29" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis, 1951</bibRefCitation>
</taxonomicName>
(
<figureCitation id="130030872A507614FCB485A22D47FA59" box="[793,865,1432,1458]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
) has many affinities with 
<taxonomicName id="4C3B57812A507614FB3085A22B6CFA5A" ID-CoL="85MD5" box="[1181,1354,1432,1457]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FB3085A22B6CFA5A" box="[1181,1354,1432,1457]" italics="true" pageId="5" pageNumber="22">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FAF985A22BB0FA59" box="[1364,1430,1432,1458]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
, and is also closely related to 
<taxonomicName id="4C3B57812A507614FE4585FA2CBDFA32" ID-CoL="85MCS" box="[488,667,1472,1497]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FE4585FA2CBDFA32" box="[488,667,1472,1497]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
, to such degree that in the description of the latter species 
<bibRefCitation id="EFAA51F32A507614FAEA85FA2F45F9E9" author="Banner" pageId="5" pageNumber="29" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
added that &quot;when enough mature specimens from the tropical Atlantic and Mediterranean are studied, it may be found that their forms show such variation that 
<taxonomicName id="4C3B57812A507614FC68862A2A5EF9C2" ID-CoL="85MCS" box="[965,1144,1552,1577]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FC68862A2A5EF9C2" box="[965,1144,1552,1577]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
may be considered to be a synonym&quot;. However, the chelipeds of 
<taxonomicName id="4C3B57812A507614FDD186022D14F9BA" ID-CoL="85MCS" box="[636,818,1592,1617]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FDD186022D14F9BA" box="[636,818,1592,1617]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
differ in many ways from those of 
<taxonomicName id="4C3B57812A507614FB4B86022B54F9BA" ID-CoL="85MBL" box="[1254,1394,1592,1617]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FB4B86022B54F9BA" box="[1254,1394,1592,1617]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
. In adult males of 
<taxonomicName id="4C3B57812A507614FEE4865A2FD8F992" ID-CoL="85MCS" box="[329,510,1632,1657]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FEE4865A2FD8F992" box="[329,510,1632,1657]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
the chelipeds are more or less asymmetrical in shape, and subequal in shape, the chelae are rounded to oval-shaped and laterally compressed in both sexes (at least in adults), whereas in adult males of 
<taxonomicName id="4C3B57812A507614FEE1868A2FF1F922" ID-CoL="85MBL" box="[332,471,1712,1737]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FEE1868A2FF1F922" box="[332,471,1712,1737]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
the chelipeds are both strongly asymmetrical in shape and unequal in size (
<figureCitation id="130030872A507614FAC0868A2E83F919" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
A), the chelae are sub-rectangular, not oval, flattened only on “mesial” side (
<figureCitation id="130030872A507614FB8086E22A55F919" box="[1069,1139,1752,1778]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
C, E). In 
<taxonomicName id="4C3B57812A507614FB4C86E22BB2F91A" ID-CoL="85MCS" box="[1249,1428,1752,1777]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FB4C86E22BB2F91A" box="[1249,1428,1752,1777]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
, the merus is not expanded and relatively slender, whereas in 
<taxonomicName id="4C3B57812A507614FC08873A2A13F8F2" ID-CoL="85MBL" box="[933,1077,1792,1817]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FC08873A2A13F8F2" box="[933,1077,1792,1817]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
, the merus is conspicuously expanded and stouter in both sexes, especially in males (
<figureCitation id="130030872A507614FC8387122D52F8A9" box="[814,884,1832,1858]" captionStart="FIGURE 5" captionStartId="9.[151,255,1620,1644]" captionTargetBox="[169,1427,213,1585]" captionTargetId="figure@9.[151,1436,194,1597]" captionTargetPageId="9" captionText="FIGURE 5. Athanas amazone Holthuis, 1951: male, CL 4.9 mm, MNHN-Na 13748, Banyuls-sur-Mer, southern France: A – habitus; B – major cheliped, chela and distal carpus, mesial view; male, CL 3.2 mm, MNHN-Na 8806, Pointe Noire, Congo: C – major cheliped, chela and distal carpus, ventral view; D – same, ventromesial view; E – same, dorsomesial view; F – same, basis and ischium, ventromesial view; G – same as B, fingers in slightly different angle (entire cheliped illustrated by Crosnier &amp; Forest 1973, fig. 48); ovigerous female, CL not measured, MNHN-Na 1745, Cap l'Abeille, Banyuls-sur-Mer, southern France, H - minor cheliped, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/178387/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
A). The major chelipeds of 
<taxonomicName id="4C3B57812A507614FB1A87122B4CF8AA" ID-CoL="85MCS" box="[1207,1386,1832,1857]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FB1A87122B4CF8AA" box="[1207,1386,1832,1857]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B57812A507614FF3A876A2F39F882" ID-CoL="85MBL" box="[151,287,1872,1897]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FF3A876A2F39F882" box="[151,287,1872,1897]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
also differ in the relative proportions of the ischium to the merus, and the merus to the carpus; for instance, in 
<taxonomicName id="4C3B57812A507614FE8987422FF0F87A" ID-CoL="85MCS" box="[292,470,1912,1937]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FE8987422FF0F87A" box="[292,470,1912,1937]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
the carpus is much shorter than the merus (at least in adults), while in 
<taxonomicName id="4C3B57812A507614FABE87422BBAF87A" ID-CoL="85MBL" box="[1299,1436,1912,1937]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FABE87422BBAF87A" box="[1299,1436,1912,1937]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
these articles are subequal. In our opinion, these differences are sufficient to reject Banner’s tentative assumption of the synonymy of 
<taxonomicName id="4C3B57812A507614FE1187F22C56F80A" ID-CoL="85MCS" box="[444,624,1992,2017]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="phyllocheles">
<emphasis id="B94FF0102A507614FE1187F22C56F80A" box="[444,624,1992,2017]" italics="true" pageId="5" pageNumber="22">A. phyllocheles</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C3B57812A507614FD1987F22D18F80A" ID-CoL="85MBL" box="[692,830,1992,2017]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A507614FD1987F22D18F80A" box="[692,830,1992,2017]" italics="true" pageId="5" pageNumber="22">A. amazone</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C3B57812A507614FCE087F22A67F80A" ID-CoL="85MD5" box="[845,1089,1992,2017]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A507614FCE087F22A67F80A" box="[845,1089,1992,2017]" italics="true" pageId="5" pageNumber="22">Athanas sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A507614FBE587F22AACF809" box="[1096,1162,1992,2018]" pageId="5" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
, can be separated from 
<taxonomicName id="4C3B57812A537617FF3A80A22F07FF5A" ID-CoL="85MBL" box="[151,289,152,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="amazone">
<emphasis id="B94FF0102A537617FF3A80A22F07FF5A" box="[151,289,152,177]" italics="true" pageId="6" pageNumber="23">A. amazone</emphasis>
</taxonomicName>
by (1) the absence of the infra-corneal spines; and (2) the more elongated ischium of the major cheliped.
</paragraph>
</subSubSection>
<subSubSection id="C3217F892A537617FF6A80D22F9CFE21" pageId="6" pageNumber="23" type="description">
<paragraph id="8B842C022A537617FF6A80D22F9CFE21" blockId="6.[151,1437,152,458]" pageId="6" pageNumber="23">
All other species of 
<taxonomicName id="4C3B57812A537617FE1F80D22C34FEEA" box="[434,530,232,257]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B94FF0102A537617FE1F80D22C34FEEA" box="[434,530,232,257]" italics="true" pageId="6" pageNumber="23">Athanas</emphasis>
</taxonomicName>
bearing slender, scythe-shaped dactyli on the walking legs are more distantly related to 
<taxonomicName id="4C3B57812A537617FEA6812A2F90FEC2" box="[267,438,272,297]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="sydneyensis" status="sp. nov.">
<emphasis id="B94FF0102A537617FEA6812A2F90FEC2" box="[267,438,272,297]" italics="true" pageId="6" pageNumber="23">A. sydneyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C4D6B2A537617FE10812A2FDDFEC1" box="[445,507,272,298]" pageId="6" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
For instance, 
<taxonomicName id="4C3B57812A537617FD0D812A2A22FEC1" authority="Bruce, 1990" authorityName="Bruce" authorityYear="1990" box="[672,1028,272,298]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="B94FF0102A537617FD0D812A2D48FEC2" box="[672,878,272,297]" italics="true" pageId="6" pageNumber="23">A. hongkongensis</emphasis>
<bibRefCitation id="EFAA51F32A537617FCD8812A2A22FEC1" author="Bruce" box="[885,1028,272,298]" pageId="6" pageNumber="29" refString="Bruce, A. J. (1990) Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, 2, pp. 611 - 648. Hong Kong University Press, Hong Kong." type="journal article" year="1990">Bruce, 1990</bibRefCitation>
</taxonomicName>
can be distinguished from the new species by the proportions of the dactylus to the propodus on the third to fifth pereiopods; 
<taxonomicName id="4C3B57812A537617FB5F81022FF7FE91" authority="Anker, Jeng &amp; Chan, 2001" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="B94FF0102A537617FB5F81022BBAFEBA" box="[1266,1436,312,337]" italics="true" pageId="6" pageNumber="23">A. dentirostris</emphasis>
<bibRefCitation id="EFAA51F32A537617FF3A815A2FF7FE91" author="Anker" box="[151,465,352,378]" pageId="6" pageNumber="29" refString="Anker, A., Jeng, M. S., &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061." type="journal article" year="2001">Anker, Jeng &amp; Chan, 2001</bibRefCitation>
</taxonomicName>
by the dorsally dentate rostrum; 
<taxonomicName id="4C3B57812A537617FCF8815A2A98FE91" authority="Hayashi, 2002" authorityName="Hayashi" authorityYear="2002" box="[853,1214,352,378]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="squillophilus">
<emphasis id="B94FF0102A537617FCF8815A2A2BFE92" box="[853,1037,352,377]" italics="true" pageId="6" pageNumber="23">A. squillophilus</emphasis>
<bibRefCitation id="EFAA51F32A537617FBB9815A2A98FE91" author="Hayashi" box="[1044,1214,352,378]" pageId="6" pageNumber="29" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi, 2002</bibRefCitation>
</taxonomicName>
by the presence of a strong pterygostomial spine; also, all three species have differently shaped chelipeds (
<bibRefCitation id="EFAA51F32A537617FB3181B22B0FFE49" author="Bruce" box="[1180,1321,392,418]" pageId="6" pageNumber="29" refString="Bruce, A. J. (1990) Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, 2, pp. 611 - 648. Hong Kong University Press, Hong Kong." type="journal article" year="1990">Bruce 1990</bibRefCitation>
; Anker 
<emphasis id="B94FF0102A537617FA2A81B22E93FE22" italics="true" pageId="6" pageNumber="23">et al.</emphasis>
2001; 
<bibRefCitation id="EFAA51F32A537617FEAB818A2F88FE21" author="Hayashi" box="[262,430,432,458]" pageId="6" pageNumber="29" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403." type="journal article" year="2002">Hayashi 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>