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<document id="BE35BFB907A72A50EADD5ED215EF0960" ID-CLB-Dataset="7606" ID-DOI="10.11646/zootaxa.4942.2.2" ID-GBIF-Dataset="01037cc6-51ee-42b2-8a7d-4ac91c6ab3e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4600424" ID-ZooBank="2E66E17F-621B-41B1-A806-38FA45F3112E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615534866783" checkinUser="plazi" docAuthor="Quéinnec, Eric &amp; Ollivier, Eric" docDate="2021" docId="F25E555CFFACC41FFF21F950FBFE2F68" docLanguage="en" docName="zootaxa.4942.2.2.pdf" docOrigin="Zootaxa 4942 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Duvalius (Neoduvalius) lohaji Queinnec and Ollivier 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="176" masterDocId="0E672D24FFAFC41BFFB6FFCBFFD42E76" masterDocTitle="Duvalius (Neoduvalius) lohaji n. sp., a new remarkable subterranean taxon of the isotopic Trechini lineage from Dinaric karst, Bosnia and Herzegovina (Coleoptera Carabidae: Trechinae)" masterLastPageNumber="192" masterPageNumber="173" pageNumber="176" updateTime="1698933310644" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="00CAB9A6A88742018FCCDAA11D7C115F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="99371F2B20A97366E361D5531A3D974B">Duvalius (Neoduvalius) lohaji n. sp., a new remarkable subterranean taxon of the isotopic Trechini lineage from Dinaric karst, Bosnia and Herzegovina (Coleoptera Carabidae: Trechinae)</mods:title>
</mods:titleInfo>
<mods:name id="8FFB19DD9E2EDB03212C79A72ADBE342" type="personal">
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<mods:namePart id="5640369E7BE7FFADCD7A78B7E39FE7D3">Quéinnec, Eric</mods:namePart>
<mods:affiliation id="AAD8328C859419877782B1282296A6C3">Institut de Systématique, Evolution, Biodiversité (ISYEB), Sorbonne Université, Muséum National d’Histoire Naturelle, CNRS, EPHE, Bâtiment A, 7, Quai Saint-Bernard, 75005 Paris, France.</mods:affiliation>
</mods:name>
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<mods:role id="CE971D8F33B246F4AA70A91DC3A21B4A">
<mods:roleTerm id="AADDD5945808B877ACAF39ADE40BF12A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAC200BCAFC58CA9A51BF6F3FD789749">Ollivier, Eric</mods:namePart>
<mods:affiliation id="D2113B9A0539E00AA34F7C6DA6101EC0">105, Avenue du Mont Gaillard, 76620 Le Havre, France.</mods:affiliation>
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<mods:title id="64B12B5853869B5478975E49DD184C70">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="58DF2286C92F356202B18C29DC3C6981">
<mods:date id="02E28778D387D0997BE57D7C7830A545">2021</mods:date>
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<mods:number id="BDB2ACC18928AA676A366CEA33D4540E">2021-03-12</mods:number>
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<mods:detail id="A95801267E25F5E4AB3770834A279A59" type="volume">
<mods:number id="A1407A4FAA6945157769B796D0C27B4E">4942</mods:number>
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<mods:number id="0569E63486340994C03184D342DC65F7">2</mods:number>
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<mods:classification id="C182FE71441E45096BBBCABF69954172">journal article</mods:classification>
<mods:identifier id="A51A722877C29FB6C20E361210873875" type="CLB-Dataset">7606</mods:identifier>
<mods:identifier id="80DAB152A16D136AA6EB16CA72E76B5E" type="DOI">10.11646/zootaxa.4942.2.2</mods:identifier>
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<mods:identifier id="068530628B5327698762F98D036CFA56" type="ISSN">1175-5326</mods:identifier>
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<treatment id="F25E555CFFACC41FFF21F950FBFE2F68" ID-DOI="http://doi.org/10.5281/zenodo.4638974" ID-GBIF-Taxon="180685391" ID-Zenodo-Dep="4638974" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68" httpUri="http://treatment.plazi.org/id/F25E555CFFACC41FFF21F950FBFE2F68" lastPageId="4" lastPageNumber="176" pageId="3" pageNumber="176">
<subSubSection id="32EDB7C1FFACC418FF21F950FCBE28C0" box="[151,874,1691,1718]" pageId="3" pageNumber="176" type="nomenclature">
<paragraph id="7A48E44AFFACC418FF21F950FCBE28C0" blockId="3.[151,874,1691,1754]" box="[151,874,1691,1718]" pageId="3" pageNumber="176">
<heading id="21005326FFACC418FF21F950FCBE28C0" bold="true" box="[151,874,1691,1718]" fontSize="11" level="1" pageId="3" pageNumber="176" reason="1">
<emphasis id="48833858FFACC418FF21F950FCBE28C0" bold="true" box="[151,874,1691,1718]" pageId="3" pageNumber="176">
<taxonomicName id="BDF79FC9FFACC418FF21F950FCF528C0" ID-CoL="8MG8D" authority="Queinnec and Ollivier" authorityName="Queinnec and Ollivier" authorityYear="2021" box="[151,801,1691,1718]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="176" phylum="Arthropoda" rank="species" species="lohaji" status="sp. nov." subGenus="Neoduvalius">
<emphasis id="48833858FFACC418FF21F950FDD028C0" bold="true" box="[151,516,1691,1718]" italics="true" pageId="3" pageNumber="176">Duvalius (Neoduvalius) lohaji</emphasis>
Quéinnec and Ollivier
</taxonomicName>
<taxonomicNameLabel id="53B08523FFACC418FC9EF957FCBE28C0" box="[808,874,1692,1718]" pageId="3" pageNumber="176" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32EDB7C1FFACC418FF21F90BFEA628AC" box="[151,370,1728,1754]" pageId="3" pageNumber="176" type="description">
<paragraph id="7A48E44AFFACC418FF21F90BFEA628AC" blockId="3.[151,874,1691,1754]" box="[151,370,1728,1754]" pageId="3" pageNumber="176">
(
<figureCitation id="E2CCF8CFFFACC418FF29F90BFF3B28AC" box="[159,239,1728,1754]" captionStart="FIGURE 2" captionStartId="4.[151,250,1973,1997]" captionTargetBox="[193,1394,319,1941]" captionTargetId="figure-101@4.[183,1404,309,1950]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of the male holotype of Duvalius (Neoduvalius) lohaji n. sp., from “Mračna Pećina, Livno, Prolog Planina, Bosnia and Herzegovina”. Dorsal view (A) and position of the main dorsal setae (B). Scale bar: 1.5mm." figureDoi="http://doi.org/10.5281/zenodo.4600430" httpUri="https://zenodo.org/record/4600430/files/figure.png" pageId="3" pageNumber="176">Figs. 2</figureCitation>
,
<figureCitation id="E2CCF8CFFFACC418FF4DF90BFECB28AC" box="[251,287,1728,1754]" captionStart="FIGURE 3" captionStartId="5.[151,250,1104,1128]" captionTargetBox="[160,1426,190,1072]" captionTargetId="figure-19@5.[151,1436,181,1080]" captionTargetPageId="5" captionText="FIGURE 3. Comparative right elytral morphology of different specialized genera belonging to the Duvalius lineage. Female paratype of Duvalius (Neoduvalius) lohaji n. sp. from “Mračna Pećina, Livno, Prolog Planina, Bosnia and Herzegovina” (A); Male of Agostinia launi (Gestro, 1892) from “Grotta superiore delle Camosciere, Val Pesio, Italy” (B); Male of Duvaliaphaenops raffaldianus (Lemaire, 1981) from “Grotte de Malpertus (= de Peira-Cava), Le Moulinet, Alpes-Maritimes, France” (C); Male of Trichaphaenops gounellei (Bedel, 1880) from “Grotte du Brudour, Bouvante, Drôme, France” (D); Scale bars: 0.75 mm. Black arrowheads indicate relative positions of the setae belonging to the humeral umbilicate series; in all genera of the Duvalius lineage the humeral setae are generally mutually equidistant, that is also the case in Duvaliaphaenops (C) and correspond to a plesiomorphic aggregated state, while in few genera, e.g. Agostinia (B) and Trichaphaenops (D) the first, second and third setae are more closely related than the third and fourth one (red arrowheads) (Jeannel, 1928). Among the genera Anophthalmus sp. (not shown) and in D. (N.) lohaji n. sp. (A), the position of the first humeral seta (= red arrowhead) is quite variable but is usually positioned anteriorly and is more or less shifted inward along the 7th stria. Among Duvalius lineage, disaggregated conditions are probably convergent and independently derived. White arrowheads indicate the relative position of the discal and preapical setae; the anterior one is located at the level of the second (Agostinia, B), the third (D. (N.) lohaji n. sp., A, and Duvaliaphaenops, C) or the fourth (Trichaphaenops, D) humeral seta." figureDoi="http://doi.org/10.5281/zenodo.4600432" httpUri="https://zenodo.org/record/4600432/files/figure.png" pageId="3" pageNumber="176">3A</figureCitation>
,
<figureCitation id="E2CCF8CFFFACC418FE9DF90BFEEF28AC" box="[299,315,1728,1754]" captionStart="FIGURES 4" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[264,1329,200,1904]" captionTargetId="figure-31@6.[244,1343,181,1919]" captionTargetPageId="6" captionText="FIGURES 4. Aedeagus of the holotype of Duvalius (Neoduvalius) lohaji n. sp.—Left lateral view (A);—Ventral view (B) with details of internal sac. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.4600434" httpUri="https://zenodo.org/record/4600434/files/figure.png" pageId="3" pageNumber="176">4</figureCitation>
,
<figureCitation id="E2CCF8CFFFACC418FEF1F90BFEBF28AC" box="[327,363,1728,1754]" captionStart="FIGURE 7" captionStartId="10.[151,250,1875,1899]" captionTargetBox="[322,1266,181,1851]" captionTargetId="figure-31@10.[322,1266,181,1851]" captionTargetPageId="10" captionText="FIGURE 7. Comparative head and pronotal morphologies (dorsal view) of divergent genera belonging to the Duvalius lineage. Duvalius (Neoduvalius) lohaji n. sp. from “Mračna Pećina, Livno, PrologPlanina, Bosnia and Herzegovina” (A); Trichaphaenops gounellei (Bedel, 1880) from “Grotte du Brudour, Bouvante, Drôme, France” (B); Agostinia launi (Gestro, 1892) from “Grotta superiore delle Camosciere, Val Pesio, Italy” (C); Arctaphaenops gaisbergeri Fischhuber, 1983 from “Ober̂sterreich, Ĥllengebirge, Hochlecken-Grossĥhle 1567/29, Austria” (D); Scale bars: 0.75 mm (A, B and C) and 0.4 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.4600440" httpUri="https://zenodo.org/record/4600440/files/figure.png" pageId="3" pageNumber="176">7A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="32EDB7C1FFACC41FFF21F8C3FBFE2F68" lastPageId="4" lastPageNumber="177" pageId="3" pageNumber="176" type="materials_examined">
<paragraph id="7A48E44AFFACC418FF21F8C3FD63291C" blockId="3.[151,1437,1800,2043]" pageId="3" pageNumber="176">
<materialsCitation id="CA9FEE17FFACC418FF21F8C3FD67291C" ID-GBIF-Occurrence="3061000301" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" location="Federation" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" pageId="3" pageNumber="176" specimenCount="1" typeStatus="holotype">
<emphasis id="48833858FFACC418FF21F8C3FEE62954" bold="true" box="[151,306,1800,1826]" pageId="3" pageNumber="176">
<typeStatus id="A54C5AE8FFACC418FF21F8C3FF062954" box="[151,210,1800,1826]" pageId="3" pageNumber="176">Type</typeStatus>
locality
</emphasis>
:
<collectingCountry id="02E0A4DAFFACC418FEF5F8C3FD892954" box="[323,605,1800,1826]" name="Bosnia and Herzegovina" pageId="3" pageNumber="176">Bosnia and Herzegovina</collectingCountry>
,
<location id="7F28B291FFACC418FDDCF8C3FD362954" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68:7F28B291FFACC418FDDCF8C3FD362954" box="[618,738,1800,1826]" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" name="Federation" pageId="3" pageNumber="176">Federation</location>
of
<collectingCountry id="02E0A4DAFFACC418FCBDF8C3FBF12954" box="[779,1061,1800,1826]" name="Bosnia and Herzegovina" pageId="3" pageNumber="176">Bosnia and Herzegovina</collectingCountry>
,
<location id="7F28B291FFACC418FB84F8C3FAFB2955" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68:7F28B291FFACC418FB84F8C3FAFB2955" box="[1074,1327,1800,1827]" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" name="Herzeg-Bosnia county" pageId="3" pageNumber="176">Herzeg-Bosnia county</location>
(
<emphasis id="48833858FFACC418FAF4F8C2FEBC2930" italics="true" pageId="3" pageNumber="176">Hercegbosanska županija</emphasis>
) (n°10),
<location id="7F28B291FFACC418FE65F8E7FDC22930" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68:7F28B291FFACC418FE65F8E7FDC22930" box="[467,534,1836,1862]" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" name="Livno" pageId="3" pageNumber="176">Livno</location>
municipality,
<location id="7F28B291FFACC418FD0CF8E7FCB72930" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68:7F28B291FFACC418FD0CF8E7FCB72930" box="[698,867,1836,1862]" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" name="Dinara Planina" pageId="3" pageNumber="176">Dinara Planina</location>
(Dinaric Alps),
<location id="7F28B291FFACC418FBA8F8E7FBA22930" LSID="urn:lsid:plazi:treatment:F25E555CFFACC41FFF21F950FBFE2F68:7F28B291FFACC418FBA8F8E7FBA22930" box="[1054,1142,1836,1862]" country="Bosnia and Herzegovina" county="Mracna Pecina" latitude="43.829445" longLatPrecision="19" longitude="16.743889" municipality="Prolog mountain" name="Troglav" pageId="3" pageNumber="176">Troglav</location>
massif,
<collectingMunicipality id="9A2C7E30FFACC418FB6EF8E7FA4C2930" box="[1240,1432,1836,1862]" pageId="3" pageNumber="176">Prolog mountain</collectingMunicipality>
,
<collectingCounty id="93299CC6FFACC418FF21F89BFEEB291C" box="[151,319,1872,1898]" pageId="3" pageNumber="176">Mračna Pećina</collectingCounty>
(cave),
<geoCoordinate id="1FC3828DFFACC418FE2FF89BFDF6291C" box="[409,546,1872,1898]" degrees="43" direction="north" minutes="49" orientation="latitude" pageId="3" pageNumber="176" precision="15" seconds="46" value="43.829445">N43°49’46”</geoCoordinate>
<geoCoordinate id="1FC3828DFFACC418FD9FF89BFD67291C" box="[553,691,1872,1898]" degrees="16" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="176" precision="15" seconds="38" value="16.743889">E16°44’38”</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="7A48E44AFFACC41FFF71F8BFFBFE2F68" blockId="3.[151,1437,1800,2043]" lastBlockId="4.[151,1437,151,286]" lastPageId="4" lastPageNumber="177" pageId="3" pageNumber="176">
<emphasis id="48833858FFACC418FF71F8BFFEBD29F8" bold="true" box="[199,361,1908,1934]" pageId="3" pageNumber="176">Type material</emphasis>
:
<materialsCitation id="CA9FEE17FFACC418FEC0F8BFFCD429C4" ID-GBIF-Occurrence="3061000303" collectingDate="2017-08" collectingDateMax="2018-06" collectingDateMin="2017-08" collectorName="Ollivier" pageId="3" pageNumber="176" specimenCount="1" typeStatus="holotype">
<typeStatus id="A54C5AE8FFACC418FEC0F8BFFE0829F8" box="[374,476,1908,1934]" pageId="3" pageNumber="176" type="holotype">Holotype</typeStatus>
male labelled as follows: “Mračna Pećina / Prolog—Dinara Planina - BiH /
<date id="0E49C28AFFACC418FAB8F8BEFAAB29F8" box="[1294,1407,1908,1935]" pageId="3" pageNumber="176" value="2017-08">
<collectingDate id="1E0D3B62FFACC418FAB8F8BEFAAB29F8" box="[1294,1407,1908,1935]" pageId="3" pageNumber="176" value="2017-08">VIII. 2017</collectingDate>
</date>
—
<date id="0E49C28AFFACC418FF21F852FF2C29C4" box="[151,248,1944,1971]" pageId="3" pageNumber="176" value="2018-06">
<collectingDate id="1E0D3B62FFACC418FF21F852FF2C29C4" box="[151,248,1944,1971]" pageId="3" pageNumber="176" value="2018-06">VI. 2018</collectingDate>
</date>
/ Quéinnec and
<collectorName id="D702819CFFACC418FE1FF853FE2A29C4" box="[425,510,1944,1970]" pageId="3" pageNumber="176">Ollivier</collectorName>
leg.” (white label, p) / “
</materialsCitation>
<materialsCitation id="CA9FEE17FFACC418FCB6F853FDCE29A0" ID-GBIF-Occurrence="3061000304" collectingDate="2016-06" collectingDateMax="2017-06" collectingDateMin="2016-06" collectorName="Queinnec &amp; Ollivier" pageId="3" pageNumber="176" specimenCount="2" typeStatus="holotype">
<typeStatus id="A54C5AE8FFACC418FCB6F853FCBD29C4" box="[768,873,1944,1970]" pageId="3" pageNumber="176" type="holotype">Holotype</typeStatus>
<emphasis id="48833858FFACC418FCD9F853FAC529C4" box="[879,1297,1944,1970]" italics="true" pageId="3" pageNumber="176">
<taxonomicName id="BDF79FC9FFACC418FCD9F853FB6E29C4" authority="Quéinnec &amp; Ollivier, 2021" authorityName="Quéinnec &amp; Ollivier" authorityYear="2021" box="[879,1210,1944,1970]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="176" phylum="Arthropoda" rank="species" species="lohaji" status="sp. nov." subGenus="Neoduvalius">Duvalius (Neoduvalius) lohaji</taxonomicName>
<taxonomicNameLabel id="53B08523FFACC418FB76F852FAC529C4" box="[1216,1297,1945,1970]" pageId="3" pageNumber="176" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
<collectorName id="D702819CFFACC418FAA1F853FA5429C4" box="[1303,1408,1944,1970]" pageId="3" pageNumber="176">Quéinnec</collectorName>
&amp;
<collectorName id="D702819CFFACC418FF21F877FF3829A0" box="[151,236,1980,2006]" pageId="3" pageNumber="176">Ollivier</collectorName>
des.” (red label, p), in cEQ;
</materialsCitation>
<materialsCitation id="CA9FEE17FFACC418FD96F876FE46298C" ID-GBIF-Occurrence="3061000305" collectingDate="2016-06" collectingDateMax="2017-06" collectingDateMin="2016-06" collectorName="Queinnec &amp; Ollivier" pageId="3" pageNumber="176" specimenCount="2" typeStatus="paratype">
<typeStatus id="A54C5AE8FFACC418FD96F876FD5B29A1" box="[544,655,1981,2007]" pageId="3" pageNumber="176" type="paratype">Paratypes</typeStatus>
: two females, with same general data as holotype, but one labelled: “
<date id="0E49C28AFFACC418FAC1F877FF04298C" pageId="3" pageNumber="176" value="2016-06">
<collectingDate id="1E0D3B62FFACC418FAC1F877FF04298C" pageId="3" pageNumber="176" value="2016-06">VI. 2016</collectingDate>
</date>
—
<date id="0E49C28AFFACC418FF5BF82BFE98298C" box="[237,332,2016,2042]" pageId="3" pageNumber="176" value="2017-06">
<collectingDate id="1E0D3B62FFACC418FF5BF82BFE98298C" box="[237,332,2016,2042]" pageId="3" pageNumber="176" value="2017-06">VI. 2017</collectingDate>
</date>
” and “
</materialsCitation>
<materialsCitation id="CA9FEE17FFACC41FFE24F82BFEDB2E8F" ID-GBIF-Occurrence="3061000302" collectingDate="2008-08-13" collectingDateMax="2016-06-12" collectingDateMin="2008-08-13" collectorName="Queinnec &amp; Ollivier" country="Bosnia and Herzegovina" lastPageId="4" lastPageNumber="177" pageId="3" pageNumber="176" specimenCount="1" typeStatus="paratype">
<typeStatus id="A54C5AE8FFACC418FE24F82BFE21298C" box="[402,501,2016,2042]" pageId="3" pageNumber="176" type="paratype">Paratype</typeStatus>
<emphasis id="48833858FFACC418FE4CF82BFC41298C" box="[506,917,2016,2042]" italics="true" pageId="3" pageNumber="176">
<taxonomicName id="BDF79FC9FFACC418FE4CF82BFC97298C" authorityName="Quéinnec &amp; Ollivier" authorityYear="2021" box="[506,835,2016,2042]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="176" phylum="Arthropoda" rank="species" species="lohaji" subGenus="Neoduvalius">Duvalius (Neoduvalius) lohaji</taxonomicName>
nov sp.
</emphasis>
<collectorName id="D702819CFFACC418FC2CF82BFBD7298C" box="[922,1027,2016,2042]" pageId="3" pageNumber="176">Quéinnec</collectorName>
&amp;
<collectorName id="D702819CFFACC418FB95F82BFBAC298C" box="[1059,1144,2016,2042]" pageId="3" pageNumber="176">Ollivier</collectorName>
des.” (red label, p) in cEO, cEQ; Remains of one specimen (a pair of elytra), same locality and collectors but “
<date id="0E49C28AFFABC41FFBA9FF5CFB7A2EC7" box="[1055,1198,151,177]" pageId="4" pageNumber="177" value="2016-06-12">
<collectingDate id="1E0D3B62FFABC41FFBA9FF5CFB7A2EC7" box="[1055,1198,151,177]" pageId="4" pageNumber="177" value="2016-06-12">12. VI. 2016</collectingDate>
</date>
”, in cEQ; One female labelled as follows: “
<collectingCountry id="02E0A4DAFFABC41FFECBFF70FE182EA3" box="[381,460,187,213]" name="Bosnia and Herzegovina" pageId="4" pageNumber="177">Bosnia</collectingCountry>
&amp; Hercegovina, Ođak, Livno, Dinara, Nevidna Voda,
<date id="0E49C28AFFABC41FFB95FF70FB7A2EA3" box="[1059,1198,187,213]" pageId="4" pageNumber="177" value="2008-08-13">
<collectingDate id="1E0D3B62FFABC41FFB95FF70FB7A2EA3" box="[1059,1198,187,213]" pageId="4" pageNumber="177" value="2008-08-13">13. 08. 2008</collectingDate>
</date>
,
<collectorName id="D702819CFFABC41FFB0CFF70FACC2EA3" box="[1210,1304,187,213]" pageId="4" pageNumber="177">J. Bedek</collectorName>
lgt.” (white label, p) / “
</materialsCitation>
<typeStatus id="A54C5AE8FFABC41FFEB9FF14FEA62E8F" box="[271,370,223,249]" pageId="4" pageNumber="177" type="paratype">Paratype</typeStatus>
<emphasis id="48833858FFABC41FFECFFF14FCCE2E8E" box="[377,794,223,249]" italics="true" pageId="4" pageNumber="177">
<taxonomicName id="BDF79FC9FFABC41FFECFFF14FD112E8F" authorityName="Quéinnec &amp; Ollivier" authorityYear="2021" box="[377,709,223,249]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="177" phylum="Arthropoda" rank="species" species="lohaji" subGenus="Neoduvalius">Duvalius (Neoduvalius) lohaji</taxonomicName>
nov sp.
</emphasis>
Quéinnec &amp; Ollivier des.” (red label, p) in cRL; (genitalia extracted, mounted in Euparal on transparent label and pinned under the specimen).
</paragraph>
</subSubSection>
</treatment>
</document>
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