180 lines
17 KiB
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180 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro & Hirowatari, Toshiya" docDate="2014" docId="EDD8C19D1355AB068A14149B1E01E5E9" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus triguttatus Reitter 1874" docType="treatment" docVersion="3" lastPageNumber="27" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="25" updateTime="1668158244412" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshida, Takahiro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hirowatari, Toshiya</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>403</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
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<mods:identifier type="ZBK">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052612" LSID="urn:lsid:plazi:treatment:EDD8C19D1355AB068A14149B1E01E5E9" httpUri="http://treatment.plazi.org/id/EDD8C19D1355AB068A14149B1E01E5E9" lastPageId="12" lastPageNumber="27" pageId="10" pageNumber="25">
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<subSubSection pageId="10" pageNumber="25" type="nomenclature">
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<paragraph pageId="10" pageNumber="25">
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<taxonomicName LSID="http://species-id.net/wiki/Psammoecus_triguttatus" authority="Reitter, 1874" authorityName="Reitter" authorityYear="1874" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus Reitter, 1874</taxonomicName>
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Figs 1D, 6and 13
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<normalizedToken originalValue="J–L">J-L</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="reference_group">
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<paragraph pageId="10" pageNumber="25">
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<taxonomicName genus="Psamoecus" lsidName="Psamoecus triguttatus" pageId="10" pageNumber="25" rank="species" species="triguttatus">Psamoecus triguttatus</taxonomicName>
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[sic.] Reitter, 1874: 524. Type locality: Japan; Type deposition: the Natural History Museum, London; Type examined. (misspelling)
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</paragraph>
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<paragraph pageId="10" pageNumber="25">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttata" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="triguttata">Psammoecus triguttata</taxonomicName>
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[sic.]:
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<bibRefCitation author="Hisamatsu, S" journalOrPublisher="Transaction of the Shikoku Entomological Society" pageId="27" pageNumber="42" pagination="15 - 17" title="Some clavicorn beetles from Nepal (Coleoptera)." volume="16" year="1982">Hisamatsu 1982</bibRefCitation>
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: 16.
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</paragraph>
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<paragraph pageId="10" pageNumber="25">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
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:
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<bibRefCitation author="Hetschko, A" journalOrPublisher="W. Junk, Berlin" pageId="27" pageNumber="42" pagination="1 - 90" title="Coleopterorum Catalogus Pars 109" year="1930">Hetschko 1930</bibRefCitation>
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: 82. (catalogue) -
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<bibRefCitation author="Kamiya, H" journalOrPublisher="Tsukushi no Konchu" pageId="27" pageNumber="42" pagination="15 - 18" title="How to identify flat bark beetle (Nihonsan hiratamushi no miwake-kata)." volume="6" year="1961">Kamiya 1961</bibRefCitation>
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: 18, pl. 5. -
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<bibRefCitation author="Sasaji, H" editor="Kurosawa, Y" journalOrPublisher="Hoikusha Publishing, Osaka" pageId="28" pageNumber="43" pagination="202 - 205" title="The Coleoptera of Japan in Color" year="1985">Sasaji 1985</bibRefCitation>
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: 204, fig. 33 in pl. 32. -
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<bibRefCitation author="Halstead, DGH" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="42" pagination="496 - 501" title="Silvanidae." volumeTitle="Catalogue of Palaearctic Coleoptera" year="2007">Halstead et al. 2007</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="27" pageId="10" pageNumber="25" type="diagnosis">
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<paragraph pageId="10" pageNumber="25">Diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="25">
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This species is closely similar to
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus labyrinthicus" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="labyrinthicus">Psammoecus labyrinthicus</taxonomicName>
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sp. n. Morphological differences among these species were stated in diagnosis of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="10" pageNumber="25">Description. Body length. 2.32-2.93 mm (n=23).</paragraph>
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<paragraph lastPageId="11" lastPageNumber="26" pageId="10" pageNumber="25">
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Coloration (Fig. 1D). Head and pronotum yellowish-brown to reddish-brown. Elytra yellowish-brown with variable dark maculae: round ones at half, oval ones on the posterior half to posterior 1/4 of elytral suture, sometimes connected with maculae at
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<pageBreakToken pageId="11" pageNumber="26" start="start">half</pageBreakToken>
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, darkened around end of elytra. Antennae reddish-brown basally, blackish-brown from 8th to 10th antennomeres, 11th (apex) comparatively bright.
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</paragraph>
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<paragraph pageId="11" pageNumber="26">Head (Fig. 6A, B, C). Triangular, HW/HL 1.46-1.90; IE/HL 0.92-1.20. Temples slightly expanded behind eyes, narrowed toward posterior. Eyes large, prominent. Dorsal surface with relatively dense punctuation. Antennae 1.06-1.38 mm, moderately long; covered with considerable medium length or long semi-erect setae and short pubescence on each antennomere; approximate ratio of lectotype as follows: 2.7: 1.0: 1.1: 1.2: 1.3: 1.3: 1.3: 1.1: 1.1: 1.1: 2.0 (Fig. 6A).</paragraph>
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<paragraph pageId="11" pageNumber="26">Pronotum (Fig. 6B, C). Broad, PW/PL 1.31-1.50. Dorsal surface with relatively dense punctuation. Pubescence composed of relatively fine short or medium length setae, a long seta on each tooth on lateral margins and anterior and posterior angles. Each anterior angle with four or five small teeth; each lateral margin with four teeth, tooth I small, tooth II longer than tooth I, teeth III and IV relatively narrow and of almost identical size, longer than tooth II, tooth V smallest or absent; each posterior angle with a few very small protuberances of variable shape.</paragraph>
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<paragraph pageId="11" pageNumber="26">Elytra (Fig. 6E). Elongate-oval, EW/BL 0.39-0.45. Rows of punctures narrower than interstices. Pubescence composed of many semi-erect setae of medium length; several long setae in a row around anterolateral margins.</paragraph>
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<paragraph pageId="12" pageNumber="27">
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<pageBreakToken pageId="12" pageNumber="27" start="start">9</pageBreakToken>
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th abdominal sternite (Fig. 13J). Strut Y-shaped, cut at anterior 2/5. Lateral sclerites curved inward, relatively elongate.
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</paragraph>
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<paragraph pageId="12" pageNumber="27">Aedeagus (Fig. 13K, L). Parameres club-shaped, wide around bases, posterior inner margins of wide portions a little prominent, punctuation on narrow portions relatively sparse, a little denser on wide portions, posterior half of inner margins of wide portions with many setae, narrow portions with several sparse setae, apex with a long seta. Posterior half of phallobase wider towards posterior margin, distance between posterior margin and deepest point of incision of margin of upper layer narrow, protuberances around anterior 1/4 pointed inwards, posterior margin of lower layer widely deeply incised. Penis relatively flat and thin, narrowed toward apex, punctuation around apex denser toward apex.</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="27" type="type series">
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<paragraph pageId="12" pageNumber="27">Type series.</paragraph>
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<paragraph pageId="12" pageNumber="27">Lectotype: male, Nagasaki, Nagasaki Prefecture, Japan, 1869, G. Lewis leg. (BMNH). Paralectotype: 1 female, same locality, 1869, G. Lewis leg. (BMNH). (here designated)</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="27" type="specimens examined">
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<paragraph pageId="12" pageNumber="27">Specimens examined.</paragraph>
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<paragraph pageId="12" pageNumber="27">
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JAPAM: [Tokyo Pref.] 1 male, Hatagaya, Shibuya-ku, 30
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1956, K. Tanaka leg. (EUMJ). [Gifu Pref.] 1 male, Tentoku,
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<normalizedToken originalValue="Toki-chô">Toki-cho</normalizedToken>
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, Mizunami City, 6
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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2010, K.
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<normalizedToken originalValue="Itô">Ito</normalizedToken>
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leg. (ELKU); 1 male, Konokure,
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<normalizedToken originalValue="Toki-chô">Toki-cho</normalizedToken>
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, Mizunami City, 16
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2011, K.
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<normalizedToken originalValue="Itô">Ito</normalizedToken>
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leg. (ELKU). [Kagoshima Pref.] 1 male, 4 females and 3exs., Sata Cape,
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<normalizedToken originalValue="Minamiôsumi">Minamiosumi</normalizedToken>
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Town, 6
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1968, K. Suga leg. (HUSE); 1 male, Nakanoshima Island, Toshima Village, 9
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1969, M.
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<normalizedToken originalValue="Satô">Sato</normalizedToken>
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leg. (EUMJ); 1 male, same locality, 26
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1965, S. Fukuda leg. (HUSE). [Okinawa Pref.] 1 male, Aha beach, Kunigami Village, Okinawa Island, 23
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<normalizedToken originalValue="–I–">-I-</normalizedToken>
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2008, Y. Hirano leg. (YHC); 2 males and 2 exs., Motobu Town, Okinawa Island, 19
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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2008, T. Yoshida leg. (ELKU); 1 male, Fusato,
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<normalizedToken originalValue="Nanjô">Nanjo</normalizedToken>
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City, Okinawa Island, 5
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1970, M. Takagi leg. (EUMJ); 1 male, Mt. Omoto-san, Ishigaki Island, 1
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1965, Y. Hori leg. (EUMJ); 1 male, Ushuku-no-mori, Iriomote Island, 26
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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1965, Y. Hori leg. (EUMJ).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="27" type="distribution">
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<paragraph pageId="12" pageNumber="27">Distribution.</paragraph>
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<paragraph pageId="12" pageNumber="27">JAPAN: Honshu, Kyushu, Nakanoshima (Tokara Islands), Okinawa, Ishigaki and Iriomote Islands.</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="27" type="remarks">
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<paragraph pageId="12" pageNumber="27">Remarks.</paragraph>
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<paragraph pageId="12" pageNumber="27">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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sp. 4 illustrated by
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano (2009)</bibRefCitation>
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and
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano (2010)</bibRefCitation>
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seems to be this species or
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus labyrinthicus" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="labyrinthicus">Psammoecus labyrinthicus</taxonomicName>
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sp. n. However, identification is difficult, because the specimen is a female. The Japanese name of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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sp. 4 given by
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano (2009)</bibRefCitation>
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and
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano (2010)</bibRefCitation>
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<normalizedToken originalValue="“Nise-mitsumon-semaru-hiratamushi”">"Nise-mitsumon-semaru-hiratamushi"</normalizedToken>
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is adopted as the Japanese name of this species in this paper.
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</paragraph>
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<paragraph pageId="12" pageNumber="27">
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<bibRefCitation author="Halstead, DGH" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="42" pagination="496 - 501" title="Silvanidae." volumeTitle="Catalogue of Palaearctic Coleoptera" year="2007">Halstead et al. (2007)</bibRefCitation>
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recorded distribution of this species from Russia (Far East), Korea, China and Japan. However, it is now clear that at least two closely similar species occur in Japan, so, past records of this species should be reconfirmed. In addition, two or more closely similar species were sometimes found in the same limited area such as Tentoku, Toki-cho, Mizunami City, Gifu Prefecture where
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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were collected. Thus, these species should be identified carefully.
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</paragraph>
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<paragraph pageId="12" pageNumber="27">Syntypes of this species consist of two specimens, one male and one female. The male specimen is designated as a lectotype, and the female specimen is designated as a paralectotype.</paragraph>
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</subSubSection>
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</treatment>
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</document> |