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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.255.4173" ID-GBIF-Dataset="0859187d-19f5-47fc-af47-7a030cabc09f" ID-PMC="PMC3560860" ID-Pensoft-Pub="1313-2970-255-73" ID-PubMed="23459597" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-255-73" ModsDocOrigin="ZooKeys 255" ModsDocTitle="Beronaphaenops paphlagonicus, a new anophthalmous genus and species of Trechini (Coleoptera, Carabidae) from Turkey" checkinTime="1451247804851" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav V." docDate="2012" docId="D75B40A4F7DF5BA859DEC08530DB2D7C" docLanguage="en" docName="ZooKeys 255: 73-92" docOrigin="ZooKeys 255" docSource="http://dx.doi.org/10.3897/zookeys.255.4173" docTitle="Beronaphaenops paphlagonicus Gueorguiev, 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="87" masterDocId="A767FFEAFFE18D660434FFC5660E6561" masterDocTitle="Beronaphaenops paphlagonicus, a new anophthalmous genus and species of Trechini (Coleoptera, Carabidae) from Turkey" masterLastPageNumber="92" masterPageNumber="73" pageNumber="85" updateTime="1668155107587" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Beronaphaenops paphlagonicus, a new anophthalmous genus and species of Trechini (Coleoptera, Carabidae) from Turkey</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>255</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>73</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.255.4173</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.255.4173</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-255-73</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152039562" LSID="urn:lsid:plazi:treatment:D75B40A4F7DF5BA859DEC08530DB2D7C" httpUri="http://treatment.plazi.org/id/D75B40A4F7DF5BA859DEC08530DB2D7C" lastPageId="15" lastPageNumber="87" pageId="12" pageNumber="85">
<subSubSection pageId="12" pageNumber="85" type="nomenclature">
<paragraph pageId="12" pageNumber="85">
<taxonomicName LSID="urn:lsid:zoobank.org:act:E9DD4767-61C1-4347-BE9A-E505D072B8C3" class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops paphlagonicus" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="paphlagonicus">Beronaphaenops paphlagonicus</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="85">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="85" type="type material">
<paragraph pageId="12" pageNumber="85">Type material.</paragraph>
<paragraph pageId="12" pageNumber="85">
Holotype ♂, labelled: &quot;Turkey, Pinarbasi Distr., Milli Park
<normalizedToken originalValue="Kűre">Kűre</normalizedToken>
<normalizedToken originalValue="Dağlari">Daglari</normalizedToken>
, cave
<normalizedToken originalValue="Eşek">Esek</normalizedToken>
<normalizedToken originalValue="Çukuru">Cukuru</normalizedToken>
<normalizedToken originalValue="Mağarası">Magarasi</normalizedToken>
2, 14.VII.2008, P. Beron leg.&quot; [typeset] / &quot;Holotype
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
det. 2011&quot; [red typeset] (NMNHS); paratypes 1 ♂, 3 ♀♀, labelled: &quot;Turkey, Pinarbasi Distr., Milli Park
<normalizedToken originalValue="Kűre">Kűre</normalizedToken>
<normalizedToken originalValue="Dağlari">Daglari</normalizedToken>
, cave
<normalizedToken originalValue="Eşek">Esek</normalizedToken>
<normalizedToken originalValue="Çukuru">Cukuru</normalizedToken>
<normalizedToken originalValue="Mağarası">Magarasi</normalizedToken>
2, 14.VII.2008, P. Beron leg.&quot; [typeset] / &quot;Paratype
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. B.
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
det. 2011&quot; [red typeset] (MHNG; NMNHS).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="85" type="diagnosis">
<paragraph pageId="12" pageNumber="85">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="85">
A medium sized trechine, included in
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n., with pubescent integument, elongated head, complete frontal grooves, labrum serrate on the anterior margin, labium fused, mentum tooth large, long, and deeply bifid at the tip, pronotum short and narrow, without posterolateral setae, and with pointed hind angles, elytra with superficial striae, one discal pore in stria 3, umbilicate series not aggregate, aedeagus with straight apex and poorly differentiated basal bulb, apical stylomere short, broadened basally, with two large ensiform setae on tergal position.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85" type="description">
<paragraph pageId="12" pageNumber="85">Description.</paragraph>
<paragraph pageId="12" pageNumber="85">Medium sized, overall length (from the apex of longer mandible to the apices of elytra): 5.72-5.95 (mean 5.87) mm; maximum width: 1.88-1.98 (mean 1.91) mm. General shape as in Fig. 1. Integument thin, translucent, covered by short, rather sparse and uniform pubescence, tibiae and tarsi densely pubescent. Microsculpture distinct on the head and elytra, very fine, almost indistinct on the pronotum, consisting of isodiametric and transverse meshes on the head and isodiametric ones on the elytra. Colour reddish-brown on the head and pronotum to yellowish on the elytra and appendages.</paragraph>
<paragraph lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85">
Head rather elongate, 1.49-1.59 (mean 1.56) times as long as wide, 1.6-1.74 (mean 1.68) times as long as the pronotum, slightly narrow than the pronotum: 0.88
<pageBreakToken pageId="13" pageNumber="86" start="start">-</pageBreakToken>
0.93 (mean 0.91) times as wide as pronotum, clearly constricted at the neck; frontal grooves complete, extending onward onto the clypeus, deep along the apical three-fourth of the head where outwardly sinuate, somewhat shallow at the level of posterior supraorbital seta, and again deeper downhill along the collar constriction; genae convex. Antennae filiform, very long and slender, 0.63-0.65 times as long as overall body length, posteriad not exceeding middle of elytra.
</paragraph>
<paragraph pageId="13" pageNumber="86">Pronotum cordate, 1.15-1.21 (mean 1.19) times as wide as long, constricted towards the base. Anterior margin 1.24-1.27 times as wide as the posterior margin, 0.78-0.83 times wide as the pronotal maximum width, gradually concave towards the middle, the fore angles prominent, round. Posterior margin 0.63-0.65 times wide as the pronotal maximum width, concave inwards as the hind angles projecting back. Sides convex, round anteriad, straight, convergent posteriorly, hardly sinuate before the acute hind angles, which slightly projecting outside and turned upwards. Lateral margins grooved throughout, less anteriorly, more posteriorly, their extremities reflexed upward throughout, less anteriorly, more posteriorly. Anterolateral setae present, with length exceeding two-thirds of the pronotal side length, situated at the place of maximal pronotum width, posterolateral setae absent. Disc smooth, slightly convex, midline well impressed, not reaching both the basal and apical margins; apical transverse impression less distinct, continuous, laterally reaching the anterior margin, basal transverse impression sharply impressed, continuous, laterally merging into the small, but distinct basal foveae. Scutellum small, triangular.</paragraph>
<paragraph pageId="13" pageNumber="86">Elytra subovate, 1.67-1.78 times longer than wide, 3.28-3.55 times as long as the pronotal maximum length, and 1.6-1.72 times wider than the pronotal maximum width. Sides narrow and slanting at the basis, becoming wider posteriorly, with maximal width at the third-fourth, round at the apex; lateral grooves reflexed moderately upward, becoming flat at the apical sixth; shoulders obtuse though still marked, basal border absent. Disc convex; striae superficial, more or less obliterated at the sides and apically, scutellar stria absent, apical recurrent one present; intervals flat, each one with single longitudinal row of hairs in the middle; stria 3 with single discal pore before the middle; apical triangle of pores complete; umbilicate series of elytra consisting of 8 pores, not aggregate.</paragraph>
<paragraph pageId="13" pageNumber="86">Legs long and slender; protibia straight, densely pubescent, without longitudinal groove on the external face; tarsi thin, densely pubescent, segments 1 and 2 in the males strongly dilated and inwardly denticulate at the apices.</paragraph>
<paragraph pageId="13" pageNumber="86">Male genitalia: as in Figs 13-15 (see also the genus description).</paragraph>
<paragraph pageId="13" pageNumber="86">Female genitalia: as in Fig. 16 (see also the genus description).</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="88" pageId="13" pageNumber="86" type="etymology">
<paragraph pageId="13" pageNumber="86">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="86">
The specific epithet is a noun in the nominative singular in apposition. It honors Paphlagonia, an ancient country on the
<normalizedToken originalValue="todays">today's</normalizedToken>
Kastamonu Province of Turkey.
</paragraph>
<paragraph lastPageId="14" lastPageNumber="87" pageId="13" pageNumber="86">
Distribution. Up to now the new species is known only from its type locality, the cave
<normalizedToken originalValue="Eşek">Esek</normalizedToken>
<normalizedToken originalValue="Çukuru">Cukuru</normalizedToken>
<normalizedToken originalValue="Mağarası">Magarasi</normalizedToken>
2 (Fig. 17). The cave is situated in the
<normalizedToken originalValue="Küre">Kuere</normalizedToken>
<normalizedToken originalValue="Dağları">Daglari</normalizedToken>
Milli
<normalizedToken originalValue="Parkı">Parki</normalizedToken>
, a Turkish national park which includes a forested, well protected karstic area in the
<normalizedToken originalValue="Küre">Kuere</normalizedToken>
Mountain. The most important distinctive feature of this karstic zone is
<pageBreakToken pageId="14" pageNumber="87" start="start">that</pageBreakToken>
it harbors old growth natural beech and fir forests, endemic plant species, forest ecosystems rich in biological diversity, as a result of being in the humid climatic zone. The climate in the inner part of the area is harsh.
</paragraph>
<caption pageId="14" pageNumber="87">
<paragraph pageId="14" pageNumber="87">
Figure 1.
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. Photo of habitus, holotype. Scale bar: 1 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="87">
<paragraph pageId="14" pageNumber="87">
Figures 2-7.
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. 2 Labrum and clypeus in dorsal aspect, holotype. 3 Left mandible in dorsal aspect, paratype ♀ 4 Right mandible in dorsal aspect, paratype ♀5 Labium in ventral aspect, paratype ♀ 6 Apical part of ligula in ventral aspect, paratype ♀ 7 Left maxilla in ventral aspect, paratype ♀. Scale bar 2, 3, 4, 5, 7= 0.3 mm; 6: 0.1 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="87">
<paragraph pageId="14" pageNumber="87">
Figures 8-12.
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. 8 Pronotum, holotype 9 Anterior part of left elytron, paratype ♀ 10 Posterior part of left elytron, paratype ♀ 11 Male protarsus in dorsal aspect, paratype ♂ 12 Female protarsus in dorsolateral aspect, paratype ♀. Scale bar 8: 1 mm; 9, 10: 0.5 mm; 11, 12: 0.3mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="87">
<paragraph pageId="14" pageNumber="87">
Figures 13-16. 13
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n. Aedeagus in left lateral aspect, holotype. 14 Aedeagus in right lateral aspect, holotype 15 Aedeagus in dorsal aspect, holotype16 Left ovipositor in ventral aspect, paratype ♀. Scale bar 13, 14, 15: 0.2 mm; 16: 0.1 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="87">
<paragraph pageId="14" pageNumber="87">
Figure 17. Administrative map of Turkey with black arrow and dot marking approximately the type locality of
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
gen. n. paphlagonicus sp. n.
</paragraph>
</caption>
<subSection lastPageId="15" lastPageNumber="88" pageId="14" pageNumber="87" type="key to the anatolian genera of trechini">
<paragraph pageId="14" pageNumber="87">
Key to the Anatolian genera of
<taxonomicName lsidName="" pageId="14" pageNumber="87" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
</paragraph>
<paragraph lastPageId="15" lastPageNumber="88" pageId="14" pageNumber="87">
<table lastPageId="15" lastPageNumber="88" pageId="14" pageNumber="87">
<tr pageId="14" pageNumber="87">
<td colspan="1" pageId="14" pageNumber="87" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Thalassophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thalassophilus" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Thalassophilus</taxonomicName>
</td>
</tr>
<tr pageId="14" pageNumber="87">
<td colspan="1" pageId="14" pageNumber="87" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Neoblemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoblemus" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Neoblemus</taxonomicName>
</td>
</tr>
<tr pageId="14" pageNumber="87">
<td colspan="1" pageId="14" pageNumber="87" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Perileptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perileptus" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Perileptus</taxonomicName>
</td>
</tr>
<tr pageId="14" pageNumber="87">
<td colspan="1" pageId="14" pageNumber="87" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Trechus</taxonomicName>
</td>
</tr>
<tr pageId="14" pageNumber="87">
<td colspan="1" pageId="14" pageNumber="87" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Trechoblemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechoblemus" order="Coleoptera" pageId="14" pageNumber="87" phylum="Arthropoda" rank="genus">Trechoblemus</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duvalius" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="15" pageNumber="88" start="start">Duvalius</pageBreakToken>
</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Anillidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillidius" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Anillidius</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Pontodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontodytes" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Pontodytes</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Troglocimmerites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglocimmerites" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Troglocimmerites</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Kosswigia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kosswigia" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Kosswigia</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Sbordoniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sbordoniella" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Sbordoniella</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="88">
<td colspan="1" pageId="15" pageNumber="88" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Beronaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Beronaphaenops" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">Beronaphaenops</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>