644 lines
110 KiB
XML
644 lines
110 KiB
XML
<document id="512494B278F9F33FF0FEF52597E7E21C" ID-CLB-Dataset="45844" ID-DOI="10.11646/zootaxa.3794.4.2" ID-GBIF-Dataset="859cd600-edb9-4a72-bb65-dee901b7204c" ID-ISSN="1175-5326" ID-Zenodo-Dep="227062" ID-ZooBank="14828072-E311-4D78-89CE-49C8ABC91953" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460416709585" checkinUser="plazi" docAuthor="Rossi, Ilana, Fontoura, Marcela, Amaral, Silvana & Leal-Zanchet, Ana M." docDate="2014" docId="334787DB0B319410FF6BFE31F274FB8B" docLanguage="en" docName="zt03794p524.pdf" docOrigin="Zootaxa 3794 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cratera steffeni Rossi, Fontoura, Amaral & Leal-Zanchet, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="523" masterDocId="CF7EFFA30B309419FFFCFF89F739F90E" masterDocTitle="A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil" masterLastPageNumber="524" masterPageNumber="514" pageNumber="515" updateTime="1698328936979" updateUser="ExternalLinkService">
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<mods:title id="3C2F9C1E1FDB0B17EEEC3B101F8D1E4E">A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil</mods:title>
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<mods:namePart id="8AA6B3AC692A496AFF4F095639205FC4">Rossi, Ilana</mods:namePart>
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<mods:namePart id="0D4601514DB10A42B7CA1DE7F809FEA0">Fontoura, Marcela</mods:namePart>
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<mods:namePart id="E57DDABD36A7FCF0314FC3F2F3D3002F">Amaral, Silvana</mods:namePart>
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<mods:name id="2FA4FFB06E5305850EECFDA344C2BE97" type="personal">
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<mods:namePart id="632D5717BDAC3BF00B9B3BE783E70E28">Leal-Zanchet, Ana M.</mods:namePart>
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<mods:title id="878AE1C9DC37B90403B2696B3B9C42F3">Zootaxa</mods:title>
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<mods:date id="4D5E67974025A68700A503C593B73FE4">2014</mods:date>
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<mods:number id="FCF29E5817FC62584F58624433403EE9">3794</mods:number>
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<treatment id="334787DB0B319410FF6BFE31F274FB8B" ID-DOI="http://doi.org/10.5281/zenodo.5622586" ID-GBIF-Taxon="119425351" ID-Zenodo-Dep="5622586" LSID="urn:lsid:plazi:treatment:334787DB0B319410FF6BFE31F274FB8B" httpUri="http://treatment.plazi.org/id/334787DB0B319410FF6BFE31F274FB8B" lastPageId="9" lastPageNumber="523" pageId="1" pageNumber="515">
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<subSubSection id="F3F465460B319418FF6BFE31F530FB18" pageId="1" pageNumber="515" type="nomenclature">
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<paragraph id="BB5136CD0B319418FF6BFE31F695F8DE" blockId="1.[151,428,439,464]" box="[151,428,439,464]" pageId="1" pageNumber="515">
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<heading id="E01981A10B319418FF6BFE31F695F8DE" bold="true" box="[151,428,439,464]" fontSize="10" level="2" pageId="1" pageNumber="515" reason="2">
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<emphasis id="899AEADF0B319418FF6BFE31F695F8DE" bold="true" box="[151,428,439,464]" pageId="1" pageNumber="515">
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<taxonomicName id="7CEE4D4E0B319418FF6BFE31F672F8DE" ID-CoL="Z8R5" box="[151,331,439,464]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="1" pageNumber="515" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
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<emphasis id="899AEADF0B319418FF6BFE31F672F8DE" bold="true" box="[151,331,439,464]" italics="true" pageId="1" pageNumber="515">Cratera steffeni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="92A957A40B319418FEAFFE3EF695F8DE" box="[339,428,439,464]" pageId="1" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BB5136CD0B319418FF6BFD89F530FB18" blockId="1.[151,668,512,626]" box="[151,521,512,534]" pageId="1" pageNumber="515">
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<taxonomicName id="7CEE4D4E0B319418FF6BFD89F7C2FB1B" box="[151,251,512,533]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="515" phylum="Platyhelminthes" rank="genus">
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||
<emphasis id="899AEADF0B319418FF6BFD89F7C2FB1B" box="[151,251,512,533]" italics="true" pageId="1" pageNumber="515">Geoplana</emphasis>
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||
</taxonomicName>
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sp. 3:
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<bibRefCitation id="DF7F4B3C0B319418FEC2FD89F530FB18" author="Baptista" box="[318,521,512,534]" pageId="1" pageNumber="515" refString="Baptista, V. A., Matos, L. B., Fick, I. A. & Leal-Zanchet, A. M. (2006) Composicao de comunidades de planarias terrestres (Platyhelminthes, Tricladida, Terricola) do Parque Nacional dos Aparados da Serra, Brasil. Iheringia, 96, 293 - 297. http: // dx. doi. org / 10.1590 / s 0073 - 47212006000300004" type="journal article" year="2006">
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Baptista
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<emphasis id="899AEADF0B319418FE6BFD88F6FDFB1B" box="[407,452,512,534]" italics="true" pageId="1" pageNumber="515">et al</emphasis>
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||
., 2006
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</bibRefCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3F465460B319418FF6BFD96F504FB7C" pageId="1" pageNumber="515" type="reference_group">
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<paragraph id="BB5136CD0B319418FF6BFD96F504FB7C" blockId="1.[151,668,512,626]" pageId="1" pageNumber="515">
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<taxonomicName id="7CEE4D4E0B319418FF6BFD96F7C2FB3A" box="[151,251,543,564]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="515" phylum="Platyhelminthes" rank="genus">
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<emphasis id="899AEADF0B319418FF6BFD96F7C2FB3A" box="[151,251,543,564]" italics="true" pageId="1" pageNumber="515">Geoplana</emphasis>
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||
</taxonomicName>
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||
sp. 3:
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<bibRefCitation id="DF7F4B3C0B319418FEC2FD97F5A5FB3A" author="Fick" box="[318,668,542,564]" pageId="1" pageNumber="515" refString="Fick, I. A., Leal-Zanchet, A. M. & Vieira, E. M. (2006) Community structure of land flatworms (Platyhelminthes, Terricola): comparisons between Araucaria and Atlantic forest in Southern Brazil. Invertebrate Biolology, 125 (4), 306 - 313. http: // dx. doi. org / 10.1111 / j. 1744 - 7410.2006.00062. x" type="journal article" year="2006">Fick, Leal-Zanchet & Vieira, 2006</bibRefCitation>
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||
<taxonomicName id="7CEE4D4E0B319418FF6BFDB4F7C2FB5C" box="[151,251,573,594]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="515" phylum="Platyhelminthes" rank="genus">
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||
<emphasis id="899AEADF0B319418FF6BFDB4F7C2FB5C" box="[151,251,573,594]" italics="true" pageId="1" pageNumber="515">Geoplana</emphasis>
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||
</taxonomicName>
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||
sp. 9:
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||
<bibRefCitation id="DF7F4B3C0B319418FEC2FDB4F541FB5D" author="Leal-Zanchet" box="[318,632,573,595]" pageId="1" pageNumber="515" refString="Leal-Zanchet, A. M. & Baptista, V. A. (2009) Planarias terrestres (Platyhelminthes, Tricladida) em remanescentes de Floresta com Araucaria. In: Fonseca, C. R., Souza, A. F., Leal-Zanchet, A. M., Dutra, T., Backes, A. & Ganade, G. (Eds.), Floresta com Araucaria: Ecologia, Conservacao e Desenvolvimento Sustentavel. Holos, Ribeirao Preto, pp. 199 - 207." type="book chapter" year="2009">Leal-Zanchet & Baptista, 2009</bibRefCitation>
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||
<taxonomicName id="7CEE4D4E0B319418FF6BFDD5F7C2FB7F" box="[151,251,604,625]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="515" phylum="Platyhelminthes" rank="genus">
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||
<emphasis id="899AEADF0B319418FF6BFDD5F7C2FB7F" box="[151,251,604,625]" italics="true" pageId="1" pageNumber="515">Geoplana</emphasis>
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||
</taxonomicName>
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||
sp. 7:
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||
<bibRefCitation id="DF7F4B3C0B319418FEC2FDD5F504FB7C" author="Leal-Zanchet" box="[318,573,604,626]" pageId="1" pageNumber="515" refString="Leal-Zanchet, A. M., Baptista, V. A., Campos, L. M. & Raffo, J. F. (2011) Spatial and temporal patterns of land flatworm assemblages in Brazilian Araucaria forests. Invertebrate Biology, 130, 25 - 33. http: // dx. doi. org / 10.1111 / j. 1744 - 7410.2010.00215. x" type="journal article" year="2011">
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||
Leal-Zanchet
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<emphasis id="899AEADF0B319418FE37FDD5F6C7FB7F" box="[459,510,604,625]" italics="true" pageId="1" pageNumber="515">et al.</emphasis>
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||
, 2011
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||
</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3F465460B319418FF6BFD17F409FBD5" pageId="1" pageNumber="515" type="etymology">
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<paragraph id="BB5136CD0B319418FF6BFD17F409FBD5" blockId="1.[151,1437,670,2027]" pageId="1" pageNumber="515">
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<emphasis id="899AEADF0B319418FF6BFD17F62FFBB9" bold="true" box="[151,278,670,695]" pageId="1" pageNumber="515">Etymology</emphasis>
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||
. The specific name is in honor of the naturalist MSc. Clemente José Steffen S.J. (
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<emphasis id="899AEADF0B319418FB5BFD16F205FBB8" box="[1191,1340,671,694]" italics="true" pageId="1" pageNumber="515">in memoriam</emphasis>
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) and his relevant contribution to the ethnobotany in southern
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<collectingCountry id="C3F9765D0B319418FD18FD4BF415FBD5" box="[740,812,706,731]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3F465460B319418FF3BFD6FF4C9FDA1" pageId="1" pageNumber="515" type="materials_examined">
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<paragraph id="BB5136CD0B319418FF3BFD6FF221FD85" blockId="1.[151,1437,670,2027]" pageId="1" pageNumber="515">
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<emphasis id="899AEADF0B319418FF3BFD6FF640FBF1" bold="true" box="[199,377,742,767]" pageId="1" pageNumber="515">
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<typeStatus id="6455886F0B319418FF3BFD6FF63BFBF1" box="[199,258,742,767]" pageId="1" pageNumber="515">Type</typeStatus>
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material.
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||
</emphasis>
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||
<typeStatus id="6455886F0B319418FE78FD6FF6C8FBF1" box="[388,497,742,767]" pageId="1" pageNumber="515" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="DDFFAE080B319418FE03FD6EF565FBF1" box="[511,604,743,767]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="515">MZUSP</collectionCode>
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<collectionCode id="DDFFAE080B319418FD94FD6FF5B4FBF1" box="[616,653,742,767]" pageId="1" pageNumber="515">PL</collectionCode>
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.1557:
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<emphasis id="899AEADF0B319418FD26FD6FF5C5FBF0" box="[730,764,742,766]" italics="true" pageId="1" pageNumber="515">leg</emphasis>
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||
. I. Fick,
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<date id="CF50100D0B319418FC92FD6EF315FBF1" box="[878,1068,742,767]" pageId="1" pageNumber="515" value="2000-03-31">31. March 2000</date>
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, Cambará do Sul, RS,
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<collectingCountry id="C3F9765D0B319418FAC3FD6FF2BAFBF1" box="[1343,1411,742,767]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
– anterior tip: transverse sections on 28 slides; anterior region at the level of the ovaries: sagittal sections on 33 slides; anterior region at the level of the anteriormost testes: sagittal sections on 83 slides; pre-pharyngeal region: transverse sections on 18 slides; pharynx: sagittal sections on 29 slides; copulatory apparatus: sagittal sections on 34 slides;
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<typeStatus id="6455886F0B319418FEF6FCFEF645FA81" box="[266,380,887,911]" pageId="1" pageNumber="515" type="paratype">Paratypes</typeStatus>
|
||
: MZU
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||
<collectionCode id="DDFFAE080B319418FE31FCFFF6C8FA81" box="[461,497,886,911]" pageId="1" pageNumber="515">PL</collectionCode>
|
||
.00176:
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||
<emphasis id="899AEADF0B319418FDB6FCFFF555FA80" box="[586,620,886,910]" italics="true" pageId="1" pageNumber="515">leg</emphasis>
|
||
. I. Fick, 0
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||
<date id="CF50100D0B319418FD1FFCFEF4ABFA81" box="[739,914,887,911]" pageId="1" pageNumber="515" value="2001-08-07">7. August 2001</date>
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||
, Cambará do Sul, RS,
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<collectingCountry id="C3F9765D0B319418FB68FCFFF3E1FA81" box="[1172,1240,886,911]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
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||
– anterior region on the level of the ovaries: horizontal sections on 19 slides; pre-pharyngeal region: transverse sections on 16 slides; pharynx: sagittal sections on 38 slides; copulatory apparatus: horizontal sections on 16 slides; MZU
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||
<collectionCode id="DDFFAE080B319418FAE1FC36F278FAD8" box="[1309,1345,959,982]" pageId="1" pageNumber="515">PL</collectionCode>
|
||
. 00177:
|
||
<emphasis id="899AEADF0B319418FF6BFC6AF780FAF5" box="[151,185,995,1019]" italics="true" pageId="1" pageNumber="515">leg</emphasis>
|
||
. I. Fick,
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||
<date id="CF50100D0B319418FEE0FC6AF6E9FAF5" box="[284,464,994,1019]" pageId="1" pageNumber="515" value="2002-03-24">24. March 2002</date>
|
||
, Cambará do Sul, RS,
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<collectingCountry id="C3F9765D0B319418FD36FC6BF437FAF5" box="[714,782,994,1019]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
– preserved on clove oil; MZU
|
||
<collectionCode id="DDFFAE080B319418FB8EFC6AF3AFFAF4" box="[1138,1174,995,1018]" pageId="1" pageNumber="515">PL</collectionCode>
|
||
. 00178:
|
||
<emphasis id="899AEADF0B319418FB08FC6AF22FFAF5" box="[1268,1302,995,1019]" italics="true" pageId="1" pageNumber="515">leg</emphasis>
|
||
. I. Fick,
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||
<date id="CF50100D0B319418FA85FC6AF62DFD11" pageId="1" pageNumber="515" value="2002-04-21">21. April 2002</date>
|
||
, Cambará do Sul, RS,
|
||
<collectingCountry id="C3F9765D0B319418FDEEFB8FF56FFD11" box="[530,598,1030,1055]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
– anterior tip: transverse sections on 22 slides; anterior region on the level of the ovaries: sagittal sections on 23 slides; pre-pharyngeal region: transverse sections on 10 slides; pharynx: sagittal sections on 24 slides; copulatory apparatus: sagittal sections on 34 slides; MZU
|
||
<collectionCode id="DDFFAE080B319418FB88FBC6F3A1FD68" box="[1140,1176,1103,1126]" pageId="1" pageNumber="515">PL</collectionCode>
|
||
. 00179:
|
||
<emphasis id="899AEADF0B319418FB0BFBC7F220FD68" box="[1271,1305,1102,1126]" italics="true" pageId="1" pageNumber="515">leg</emphasis>
|
||
. J. Bencke, 0
|
||
<date id="CF50100D0B319418FF56FBFAF643FD85" box="[170,378,1138,1163]" pageId="1" pageNumber="515" value="2002-09-01">1. September 2002</date>
|
||
, Cambará do Sul, RS,
|
||
<collectingCountry id="C3F9765D0B319418FD84FBFBF585FD85" box="[632,700,1138,1163]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
– copulatory apparatus: sagittal sections on 20 slides.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B319418FF3BFB1FF4C9FDA1" blockId="1.[151,1437,670,2027]" box="[199,1008,1174,1199]" pageId="1" pageNumber="515">
|
||
<emphasis id="899AEADF0B319418FF3BFB1FF651FDA1" bold="true" box="[199,360,1174,1199]" pageId="1" pageNumber="515">Type-locality.</emphasis>
|
||
Cambará do Sul, state of Rio Grande do Sul (RS),
|
||
<collectingCountry id="C3F9765D0B319418FC59FB1FF4D4FDA1" box="[933,1005,1174,1199]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3F465460B319418FF3BFB33F459FDDD" box="[199,864,1210,1235]" pageId="1" pageNumber="515" type="distribution">
|
||
<paragraph id="BB5136CD0B319418FF3BFB33F459FDDD" blockId="1.[151,1437,670,2027]" box="[199,864,1210,1235]" pageId="1" pageNumber="515">
|
||
<emphasis id="899AEADF0B319418FF3BFB33F659FDDD" bold="true" box="[199,352,1210,1235]" pageId="1" pageNumber="515">Distribution.</emphasis>
|
||
Rio Grande do Sul (Cambará do Sul),
|
||
<collectingCountry id="C3F9765D0B319418FCE8FB33F465FDDD" box="[788,860,1210,1235]" name="Brazil" pageId="1" pageNumber="515">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3F465460B319418FF3BFB57F61AFF19" pageId="1" pageNumber="515" type="diagnosis">
|
||
<paragraph id="BB5136CD0B319418FF3BFB57F61AFF19" blockId="1.[151,1437,670,2027]" pageId="1" pageNumber="515">
|
||
<emphasis id="899AEADF0B319418FF3BFB57F678FDF9" bold="true" box="[199,321,1246,1271]" pageId="1" pageNumber="515">Diagnosis.</emphasis>
|
||
Body elongate with parallel margins, anterior tip rounded and posterior tip pointed. Live specimens with a broad orange band, bordered by two black paramarginal stripes on the dorsal surface. Eyes dorsal, but restrict to a lateral band on each side of the body, without clear halos; conspicuous glandular margin with four
|
||
<typeStatus id="6455886F0B319418FA9DFAAEF2A5FC31" box="[1377,1436,1319,1343]" pageId="1" pageNumber="515">types</typeStatus>
|
||
of secretory cells; mc:h, 9%–10%; pharynx bell-form; esophagus short; anteriormost testes posterior to ovaries; sperm ducts opening laterally into the proximal portion of the prostatic vesicle; prostatic vesicle extrabulbar, unpaired, with ample proximal portion and tubular distal portion; ovovitelline ducts emerging dorso-laterally from the posterior half of ovaries, and ascending laterally to the female atrium; common ovovitelline duct short; vagina long, dorso-anteriorly curved; female and male atria with almost the same length; no folds separating male and female atria.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3F465460B319410FF3BF9ABF6B4FB19" lastPageId="9" lastPageNumber="523" pageId="1" pageNumber="515" type="description">
|
||
<paragraph id="BB5136CD0B319418FF3BF9ABF237FF8D" blockId="1.[151,1437,670,2027]" pageId="1" pageNumber="515">
|
||
<emphasis id="899AEADF0B319418FF3BF9ABF660FF35" bold="true" box="[199,345,1570,1595]" pageId="1" pageNumber="515">Description.</emphasis>
|
||
External morphology. Body elongate with parallel margins, anterior tip rounded and posterior tip pointed. When creeping, maximal length reaches
|
||
<quantity id="7C169B280B319418FD37F9CFF425FF50" box="[715,796,1606,1631]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="515" unit="mm" value="68.0">68 mm</quantity>
|
||
in the
|
||
<typeStatus id="6455886F0B319418FC91F9CFF4F6FF51" box="[877,975,1606,1631]" pageId="1" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
(
|
||
<tableCitation id="F66C03760B319418FC23F9CFF30CFF51" box="[991,1077,1606,1631]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="1" pageNumber="515" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
). Mouth distance from anterior tip, 68%; gonopore distance from anterior tip, 86.5% relative to body length in the
|
||
<typeStatus id="6455886F0B319418FBC4F9E3F3A3FF8D" box="[1080,1178,1642,1667]" pageId="1" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
(
|
||
<tableCitation id="F66C03760B319418FB55F9E3F3C5FF8D" box="[1193,1276,1642,1667]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="1" pageNumber="515" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B319418FF3BF907F633FE71" blockId="1.[151,1437,670,2027]" pageId="1" pageNumber="515">
|
||
Alive, dorsum with orange pigmentation constituting a broad band, bordered by two black paramarginal stripes. Under the stereomicroscope, pale-yellow dorsal ground-colour visible on body margins. Ventral surface pale yellow. After fixation, the dorsal band becomes greyish and the paramarginal stripes, brown (
|
||
<figureCitation id="23D52A480B319418FB07F95FF25BFFE1" box="[1275,1378,1750,1775]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1909,1932]" captionTargetBox="[187,1350,219,1863]" captionTargetId="figure@3.[168,1373,195,1863]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cratera steffeni sp. nov.: (1 – 2) colour pattern of a fixed specimen (paratype MZU PL. 00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL. 00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1 – 3) or to the top (Figs. 4 – 5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes." httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="1" pageNumber="515">Figs 1–2</figureCitation>
|
||
); the ventral surface remains pale yellow. Dorsal band with maximum width of
|
||
<quantity id="7C169B280B319418FC23F973F301FE1C" box="[991,1080,1786,1811]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="515" unit="mm" value="2.6">2.6 mm</quantity>
|
||
, corresponding to 44% of body width in the
|
||
<typeStatus id="6455886F0B319418FEC8F897F6A3FE39" box="[308,410,1822,1847]" pageId="1" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
. Paramarginal stripes begin close to the anterior tip and end next to the posterior tip, converging towards both extremities of the body (
|
||
<figureCitation id="23D52A480B319418FD1BF8CBF414FE55" box="[743,813,1858,1883]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1909,1932]" captionTargetBox="[187,1350,219,1863]" captionTargetId="figure@3.[168,1373,195,1863]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cratera steffeni sp. nov.: (1 – 2) colour pattern of a fixed specimen (paratype MZU PL. 00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL. 00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1 – 3) or to the top (Figs. 4 – 5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes." httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="1" pageNumber="515">Fig. 1</figureCitation>
|
||
); they are
|
||
<quantity id="7C169B280B319418FC49F8CAF329FE54" box="[949,1040,1859,1883]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="515" unit="mm" value="0.7">0.7 mm</quantity>
|
||
wide (12% of body width in the
|
||
<typeStatus id="6455886F0B319418FF6BF8EFF7C7FE71" box="[151,254,1894,1919]" pageId="1" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B31941BFF3BF802F4E3F9F6" blockId="1.[151,1437,670,2027]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="516" pageId="1" pageNumber="515">
|
||
Eyes, initially uniserial, surround anterior tip (
|
||
<figureCitation id="23D52A480B319418FD21F803F414FEAD" box="[733,813,1930,1955]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1909,1932]" captionTargetBox="[187,1350,219,1863]" captionTargetId="figure@3.[168,1373,195,1863]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cratera steffeni sp. nov.: (1 – 2) colour pattern of a fixed specimen (paratype MZU PL. 00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL. 00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1 – 3) or to the top (Figs. 4 – 5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes." httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="1" pageNumber="515">Figs. 3</figureCitation>
|
||
̶4). After the first millimeter, eyes become pluriserial and form a lateral band on each side of the dorsal surface. Three to four eyes occur on the same transversal level, being more numerous in the median third of the body (
|
||
<figureCitation id="23D52A480B319418FCF8F85BF471FEE5" box="[772,840,2002,2027]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1909,1932]" captionTargetBox="[187,1350,219,1863]" captionTargetId="figure@3.[168,1373,195,1863]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cratera steffeni sp. nov.: (1 – 2) colour pattern of a fixed specimen (paratype MZU PL. 00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL. 00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1 – 3) or to the top (Figs. 4 – 5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes." httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="1" pageNumber="515">Fig. 5</figureCitation>
|
||
), occupying a band of about
|
||
<quantity id="7C169B280B319418FB6FF85AF3D0FEE4" box="[1171,1257,2003,2027]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="515" unit="mm" value="0.7">0.7 mm</quantity>
|
||
on each side of the body (12.5% of body width). They become less numerous and uniserial towards posterior tip (
|
||
<figureCitation id="23D52A480B32941BFAFBFF1EF277F9BE" box="[1287,1358,151,176]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1909,1932]" captionTargetBox="[187,1350,219,1863]" captionTargetId="figure@3.[168,1373,195,1863]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cratera steffeni sp. nov.: (1 – 2) colour pattern of a fixed specimen (paratype MZU PL. 00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL. 00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1 – 3) or to the top (Figs. 4 – 5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes." httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="2" pageNumber="516">Fig. 3</figureCitation>
|
||
). Eyes monolobated; they are small in the first millimeter of the body (about 19 µm in diameter). After that, they are mixed with larger eyes (30µm–36 µm in diameter). Clear halos are absent.
|
||
</paragraph>
|
||
<caption id="EF9166450B32941BFF6BFEA1F52BF891" ID-Table-UUID="EF9166450B32941BFF6BFEA1F52BF891" httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="2" pageNumber="516" targetBox="[156,1387,442,1228]" targetIsTable="true" targetPageId="2">
|
||
<paragraph id="BB5136CD0B32941BFF6BFEA1F52BF891" blockId="2.[151,1436,295,415]" pageId="2" pageNumber="516">
|
||
<emphasis id="899AEADF0B32941BFF6BFEA1F634F831" bold="true" box="[151,269,296,319]" pageId="2" pageNumber="516">TABLE 1.</emphasis>
|
||
Measurements, in mm, of specimens of
|
||
<taxonomicName id="7CEE4D4E0B32941BFD2AFEA1F4B8F831" box="[726,897,295,319]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="2" pageNumber="516" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B32941BFD2AFEA1F4B8F831" box="[726,897,295,319]" italics="true" pageId="2" pageNumber="516">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A957A40B32941BFC77FEA1F4DDF831" box="[907,996,296,319]" pageId="2" pageNumber="516" rank="species">
|
||
<emphasis id="899AEADF0B32941BFC77FEA1F4DDF831" bold="true" box="[907,996,296,319]" pageId="2" pageNumber="516">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
–: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BB5136CD0B32941BFDA2FE33F21EFDC2" pageId="2" pageNumber="516">
|
||
<table id="C9EEC46D0B326BE6FF60FE33F252FDC2" box="[156,1387,442,1228]" gridcols="6" gridrows="18" pageId="2" pageNumber="516">
|
||
<tr id="05DE348F0B326BE6FF60FE33F252FB00" box="[156,1387,442,526]" gridrow="0" pageId="2" pageNumber="516" rowspan-0="1">
|
||
<th id="460F5DF30B326BE6FDA2FE33F585FB00" box="[606,700,442,526]" gridcol="1" gridrow="0" pageId="2" pageNumber="516">Holotype MZUSP PL. 1557</th>
|
||
<th id="460F5DF30B326BE6FCEEFE33F44CFB00" box="[786,885,442,526]" gridcol="2" gridrow="0" pageId="2" pageNumber="516">Paratype MZU PL.00176</th>
|
||
<th id="460F5DF30B326BE6FC4CFE33F32AFB00" box="[944,1043,442,526]" gridcol="3" gridrow="0" pageId="2" pageNumber="516">Paratype MZU PL.00177</th>
|
||
<th id="460F5DF30B326BE6FBADFE33F3F9FB00" box="[1105,1216,442,526]" gridcol="4" gridrow="0" pageId="2" pageNumber="516">Paratype MZU PL.00178</th>
|
||
<th id="460F5DF30B326BE6FAF4FE33F252FB00" box="[1288,1387,442,526]" gridcol="5" gridrow="0" pageId="2" pageNumber="516">Paratype MZU PL.00179</th>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FDA8F252FB39" box="[156,1387,545,567]" gridrow="1" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FDA8F6F5FB39" box="[156,460,545,567]" gridcol="0" gridrow="1" pageId="2" pageNumber="516">Maximum length in extension</th>
|
||
<td id="460F5DF30B326BE6FDA2FDA8F585FB39" box="[606,700,545,567]" gridcol="1" gridrow="1" pageId="2" pageNumber="516">68</td>
|
||
<td id="460F5DF30B326BE6FCEEFDA8F44CFB39" box="[786,885,545,567]" gridcol="2" gridrow="1" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FC4CFDA8F32AFB39" box="[944,1043,545,567]" gridcol="3" gridrow="1" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFDA8F3F9FB39" box="[1105,1216,545,567]" gridcol="4" gridrow="1" pageId="2" pageNumber="516">50</td>
|
||
<td id="460F5DF30B326BE6FAF4FDA8F252FB39" box="[1288,1387,545,567]" gridcol="5" gridrow="1" pageId="2" pageNumber="516">60</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FDC3F252FB6E" box="[156,1387,586,608]" gridrow="2" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FDC3F6F5FB6E" box="[156,460,586,608]" gridcol="0" gridrow="2" pageId="2" pageNumber="516">Maximum width in extension</th>
|
||
<td id="460F5DF30B326BE6FDA2FDC3F585FB6E" box="[606,700,586,608]" gridcol="1" gridrow="2" pageId="2" pageNumber="516">3</td>
|
||
<td id="460F5DF30B326BE6FCEEFDC3F44CFB6E" box="[786,885,586,608]" gridcol="2" gridrow="2" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FC4CFDC3F32AFB6E" box="[944,1043,586,608]" gridcol="3" gridrow="2" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFDC3F3F9FB6E" box="[1105,1216,586,608]" gridcol="4" gridrow="2" pageId="2" pageNumber="516">3</td>
|
||
<td id="460F5DF30B326BE6FAF4FDC3F252FB6E" box="[1288,1387,586,608]" gridcol="5" gridrow="2" pageId="2" pageNumber="516">4</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FDFDF252FB84" box="[156,1387,628,650]" gridrow="3" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FDFDF6F5FB84" box="[156,460,628,650]" gridcol="0" gridrow="3" pageId="2" pageNumber="516">Length at rest</th>
|
||
<td id="460F5DF30B326BE6FDA2FDFDF585FB84" box="[606,700,628,650]" gridcol="1" gridrow="3" pageId="2" pageNumber="516">38</td>
|
||
<td id="460F5DF30B326BE6FCEEFDFDF44CFB84" box="[786,885,628,650]" gridcol="2" gridrow="3" pageId="2" pageNumber="516">42.5</td>
|
||
<td id="460F5DF30B326BE6FC4CFDFDF32AFB84" box="[944,1043,628,650]" gridcol="3" gridrow="3" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFDFDF3F9FB84" box="[1105,1216,628,650]" gridcol="4" gridrow="3" pageId="2" pageNumber="516">35</td>
|
||
<td id="460F5DF30B326BE6FAF4FDFDF252FB84" box="[1288,1387,628,650]" gridcol="5" gridrow="3" pageId="2" pageNumber="516">20</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FD14F252FBBD" box="[156,1387,669,691]" gridrow="4" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FD14F6F5FBBD" box="[156,460,669,691]" gridcol="0" gridrow="4" pageId="2" pageNumber="516">Width at rest</th>
|
||
<td id="460F5DF30B326BE6FDA2FD14F585FBBD" box="[606,700,669,691]" gridcol="1" gridrow="4" pageId="2" pageNumber="516">9</td>
|
||
<td id="460F5DF30B326BE6FCEEFD14F44CFBBD" box="[786,885,669,691]" gridcol="2" gridrow="4" pageId="2" pageNumber="516">6.5</td>
|
||
<td id="460F5DF30B326BE6FC4CFD14F32AFBBD" box="[944,1043,669,691]" gridcol="3" gridrow="4" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFD14F3F9FBBD" box="[1105,1216,669,691]" gridcol="4" gridrow="4" pageId="2" pageNumber="516">5</td>
|
||
<td id="460F5DF30B326BE6FAF4FD14F252FBBD" box="[1288,1387,669,691]" gridcol="5" gridrow="4" pageId="2" pageNumber="516">5</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FD4FF252FBD2" box="[156,1387,710,732]" gridrow="5" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FD4FF6F5FBD2" box="[156,460,710,732]" gridcol="0" gridrow="5" pageId="2" pageNumber="516">Length*</th>
|
||
<td id="460F5DF30B326BE6FDA2FD4FF585FBD2" box="[606,700,710,732]" gridcol="1" gridrow="5" pageId="2" pageNumber="516">44</td>
|
||
<td id="460F5DF30B326BE6FCEEFD4FF44CFBD2" box="[786,885,710,732]" gridcol="2" gridrow="5" pageId="2" pageNumber="516">42.5</td>
|
||
<td id="460F5DF30B326BE6FC4CFD4FF32AFBD2" box="[944,1043,710,732]" gridcol="3" gridrow="5" pageId="2" pageNumber="516">40</td>
|
||
<td id="460F5DF30B326BE6FBADFD4FF3F9FBD2" box="[1105,1216,710,732]" gridcol="4" gridrow="5" pageId="2" pageNumber="516">38</td>
|
||
<td id="460F5DF30B326BE6FAF4FD4FF252FBD2" box="[1288,1387,710,732]" gridcol="5" gridrow="5" pageId="2" pageNumber="516">46</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FD79F252FA08" box="[156,1387,752,774]" gridrow="6" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FD79F6F5FA08" box="[156,460,752,774]" gridcol="0" gridrow="6" pageId="2" pageNumber="516">Width*</th>
|
||
<td id="460F5DF30B326BE6FDA2FD79F585FA08" box="[606,700,752,774]" gridcol="1" gridrow="6" pageId="2" pageNumber="516">6</td>
|
||
<td id="460F5DF30B326BE6FCEEFD79F44CFA08" box="[786,885,752,774]" gridcol="2" gridrow="6" pageId="2" pageNumber="516">6.5</td>
|
||
<td id="460F5DF30B326BE6FC4CFD79F32AFA08" box="[944,1043,752,774]" gridcol="3" gridrow="6" pageId="2" pageNumber="516">5.5</td>
|
||
<td id="460F5DF30B326BE6FBADFD79F3F9FA08" box="[1105,1216,752,774]" gridcol="4" gridrow="6" pageId="2" pageNumber="516">4.5</td>
|
||
<td id="460F5DF30B326BE6FAF4FD79F252FA08" box="[1288,1387,752,774]" gridcol="5" gridrow="6" pageId="2" pageNumber="516">5</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FC90F252FA21" box="[156,1387,793,815]" gridrow="7" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FC90F6F5FA21" box="[156,460,793,815]" gridcol="0" gridrow="7" pageId="2" pageNumber="516">DM*</th>
|
||
<td id="460F5DF30B326BE6FDA2FC90F585FA21" box="[606,700,793,815]" gridcol="1" gridrow="7" pageId="2" pageNumber="516">30 (68)</td>
|
||
<td id="460F5DF30B326BE6FCEEFC90F44CFA21" box="[786,885,793,815]" gridcol="2" gridrow="7" pageId="2" pageNumber="516">30.5 (72)</td>
|
||
<td id="460F5DF30B326BE6FC4CFC90F32AFA21" box="[944,1043,793,815]" gridcol="3" gridrow="7" pageId="2" pageNumber="516">30 (75)</td>
|
||
<td id="460F5DF30B326BE6FBADFC90F3F9FA21" box="[1105,1216,793,815]" gridcol="4" gridrow="7" pageId="2" pageNumber="516">27.5 (72.5)</td>
|
||
<td id="460F5DF30B326BE6FAF4FC90F252FA21" box="[1288,1387,793,815]" gridcol="5" gridrow="7" pageId="2" pageNumber="516">30 (65)</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FCCBF252FA56" box="[156,1387,834,856]" gridrow="8" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FCCBF6F5FA56" box="[156,460,834,856]" gridcol="0" gridrow="8" pageId="2" pageNumber="516">DG*</th>
|
||
<td id="460F5DF30B326BE6FDA2FCCBF585FA56" box="[606,700,834,856]" gridcol="1" gridrow="8" pageId="2" pageNumber="516">38 (86.5)</td>
|
||
<td id="460F5DF30B326BE6FCEEFCCBF44CFA56" box="[786,885,834,856]" gridcol="2" gridrow="8" pageId="2" pageNumber="516">36.5 (86)</td>
|
||
<td id="460F5DF30B326BE6FC4CFCCBF32AFA56" box="[944,1043,834,856]" gridcol="3" gridrow="8" pageId="2" pageNumber="516">36 (90)</td>
|
||
<td id="460F5DF30B326BE6FBADFCCBF3F9FA56" box="[1105,1216,834,856]" gridcol="4" gridrow="8" pageId="2" pageNumber="516">33.5 (88)</td>
|
||
<td id="460F5DF30B326BE6FAF4FCCBF252FA56" box="[1288,1387,834,856]" gridcol="5" gridrow="8" pageId="2" pageNumber="516">38 (82.5)</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FCE5F252FA8C" box="[156,1387,876,898]" gridrow="9" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FCE5F6F5FA8C" box="[156,460,876,898]" gridcol="0" gridrow="9" pageId="2" pageNumber="516">DMG*</th>
|
||
<td id="460F5DF30B326BE6FDA2FCE5F585FA8C" box="[606,700,876,898]" gridcol="1" gridrow="9" pageId="2" pageNumber="516">8</td>
|
||
<td id="460F5DF30B326BE6FCEEFCE5F44CFA8C" box="[786,885,876,898]" gridcol="2" gridrow="9" pageId="2" pageNumber="516">6</td>
|
||
<td id="460F5DF30B326BE6FC4CFCE5F32AFA8C" box="[944,1043,876,898]" gridcol="3" gridrow="9" pageId="2" pageNumber="516">6</td>
|
||
<td id="460F5DF30B326BE6FBADFCE5F3F9FA8C" box="[1105,1216,876,898]" gridcol="4" gridrow="9" pageId="2" pageNumber="516">6</td>
|
||
<td id="460F5DF30B326BE6FAF4FCE5F252FA8C" box="[1288,1387,876,898]" gridcol="5" gridrow="9" pageId="2" pageNumber="516">8</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FC1CF252FAA5" box="[156,1387,917,939]" gridrow="10" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FC1CF6F5FAA5" box="[156,460,917,939]" gridcol="0" gridrow="10" pageId="2" pageNumber="516">DPVP*</th>
|
||
<td id="460F5DF30B326BE6FDA2FC1CF585FAA5" box="[606,700,917,939]" gridcol="1" gridrow="10" pageId="2" pageNumber="516">4</td>
|
||
<td id="460F5DF30B326BE6FCEEFC1CF44CFAA5" box="[786,885,917,939]" gridcol="2" gridrow="10" pageId="2" pageNumber="516">3</td>
|
||
<td id="460F5DF30B326BE6FC4CFC1CF32AFAA5" box="[944,1043,917,939]" gridcol="3" gridrow="10" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFC1CF3F9FAA5" box="[1105,1216,917,939]" gridcol="4" gridrow="10" pageId="2" pageNumber="516">4</td>
|
||
<td id="460F5DF30B326BE6FAF4FC1CF252FAA5" box="[1288,1387,917,939]" gridcol="5" gridrow="10" pageId="2" pageNumber="516">–</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FC37F252FADA" box="[156,1387,958,980]" gridrow="11" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FC37F6F5FADA" box="[156,460,958,980]" gridcol="0" gridrow="11" pageId="2" pageNumber="516">Creeping sole %</th>
|
||
<td id="460F5DF30B326BE6FDA2FC37F585FADA" box="[606,700,958,980]" gridcol="1" gridrow="11" pageId="2" pageNumber="516">100</td>
|
||
<td id="460F5DF30B326BE6FCEEFC37F44CFADA" box="[786,885,958,980]" gridcol="2" gridrow="11" pageId="2" pageNumber="516">100</td>
|
||
<td id="460F5DF30B326BE6FC4CFC37F32AFADA" box="[944,1043,958,980]" gridcol="3" gridrow="11" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFC37F3F9FADA" box="[1105,1216,958,980]" gridcol="4" gridrow="11" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FAF4FC37F252FADA" box="[1288,1387,958,980]" gridcol="5" gridrow="11" pageId="2" pageNumber="516">–</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FC61F252FAF0" box="[156,1387,1000,1022]" gridrow="12" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FC61F6F5FAF0" box="[156,460,1000,1022]" gridcol="0" gridrow="12" pageId="2" pageNumber="516">Ovaries</th>
|
||
<td id="460F5DF30B326BE6FDA2FC61F585FAF0" box="[606,700,1000,1022]" gridcol="1" gridrow="12" pageId="2" pageNumber="516">7.5 (17)</td>
|
||
<td id="460F5DF30B326BE6FCEEFC61F44CFAF0" box="[786,885,1000,1022]" gridcol="2" gridrow="12" pageId="2" pageNumber="516">8 (19)</td>
|
||
<td id="460F5DF30B326BE6FC4CFC61F32AFAF0" box="[944,1043,1000,1022]" gridcol="3" gridrow="12" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFC61F3F9FAF0" box="[1105,1216,1000,1022]" gridcol="4" gridrow="12" pageId="2" pageNumber="516">10 (26)</td>
|
||
<td id="460F5DF30B326BE6FAF4FC61F252FAF0" box="[1288,1387,1000,1022]" gridcol="5" gridrow="12" pageId="2" pageNumber="516">–</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FB98F252FD29" box="[156,1387,1041,1063]" gridrow="13" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FB98F6F5FD29" box="[156,460,1041,1063]" gridcol="0" gridrow="13" pageId="2" pageNumber="516">Anteriormost testes</th>
|
||
<td id="460F5DF30B326BE6FDA2FB98F585FD29" box="[606,700,1041,1063]" gridcol="1" gridrow="13" pageId="2" pageNumber="516">12 (27)</td>
|
||
<td id="460F5DF30B326BE6FCEEFB98F44CFD29" box="[786,885,1041,1063]" gridcol="2" gridrow="13" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FC4CFB98F32AFD29" box="[944,1043,1041,1063]" gridcol="3" gridrow="13" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFB98F3F9FD29" box="[1105,1216,1041,1063]" gridcol="4" gridrow="13" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FAF4FB98F252FD29" box="[1288,1387,1041,1063]" gridcol="5" gridrow="13" pageId="2" pageNumber="516">–</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FBB3F252FD5E" box="[156,1387,1082,1104]" gridrow="14" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FBB3F6F5FD5E" box="[156,460,1082,1104]" gridcol="0" gridrow="14" pageId="2" pageNumber="516">Posteriormost testes</th>
|
||
<td id="460F5DF30B326BE6FDA2FBB3F585FD5E" box="[606,700,1082,1104]" gridcol="1" gridrow="14" pageId="2" pageNumber="516">26 (59)</td>
|
||
<td id="460F5DF30B326BE6FCEEFBB3F44CFD5E" box="[786,885,1082,1104]" gridcol="2" gridrow="14" pageId="2" pageNumber="516">26 (61)</td>
|
||
<td id="460F5DF30B326BE6FC4CFBB3F32AFD5E" box="[944,1043,1082,1104]" gridcol="3" gridrow="14" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFBB3F3F9FD5E" box="[1105,1216,1082,1104]" gridcol="4" gridrow="14" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FAF4FBB3F252FD5E" box="[1288,1387,1082,1104]" gridcol="5" gridrow="14" pageId="2" pageNumber="516">–</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FBEDF252FD74" box="[156,1387,1124,1146]" gridrow="15" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FBEDF6F5FD74" box="[156,460,1124,1146]" gridcol="0" gridrow="15" pageId="2" pageNumber="516">Prostatic vesicle</th>
|
||
<td id="460F5DF30B326BE6FDA2FBEDF585FD74" box="[606,700,1124,1146]" gridcol="1" gridrow="15" pageId="2" pageNumber="516">0.7</td>
|
||
<td id="460F5DF30B326BE6FCEEFBEDF44CFD74" box="[786,885,1124,1146]" gridcol="2" gridrow="15" pageId="2" pageNumber="516">0.6</td>
|
||
<td id="460F5DF30B326BE6FC4CFBEDF32AFD74" box="[944,1043,1124,1146]" gridcol="3" gridrow="15" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFBEDF3F9FD74" box="[1105,1216,1124,1146]" gridcol="4" gridrow="15" pageId="2" pageNumber="516">0.4</td>
|
||
<td id="460F5DF30B326BE6FAF4FBEDF252FD74" box="[1288,1387,1124,1146]" gridcol="5" gridrow="15" pageId="2" pageNumber="516">0.7</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FB04F252FDAD" box="[156,1387,1165,1187]" gridrow="16" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FB04F6F5FDAD" box="[156,460,1165,1187]" gridcol="0" gridrow="16" pageId="2" pageNumber="516">Male atrium</th>
|
||
<td id="460F5DF30B326BE6FDA2FB04F585FDAD" box="[606,700,1165,1187]" gridcol="1" gridrow="16" pageId="2" pageNumber="516">0.8</td>
|
||
<td id="460F5DF30B326BE6FCEEFB04F44CFDAD" box="[786,885,1165,1187]" gridcol="2" gridrow="16" pageId="2" pageNumber="516">0.7</td>
|
||
<td id="460F5DF30B326BE6FC4CFB04F32AFDAD" box="[944,1043,1165,1187]" gridcol="3" gridrow="16" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFB04F3F9FDAD" box="[1105,1216,1165,1187]" gridcol="4" gridrow="16" pageId="2" pageNumber="516">0.6</td>
|
||
<td id="460F5DF30B326BE6FAF4FB04F252FDAD" box="[1288,1387,1165,1187]" gridcol="5" gridrow="16" pageId="2" pageNumber="516">0.9</td>
|
||
</tr>
|
||
<tr id="05DE348F0B326BE6FF60FB3FF252FDC2" box="[156,1387,1206,1228]" gridrow="17" pageId="2" pageNumber="516">
|
||
<th id="460F5DF30B326BE6FF60FB3FF6F5FDC2" box="[156,460,1206,1228]" gridcol="0" gridrow="17" pageId="2" pageNumber="516">Female atrium</th>
|
||
<td id="460F5DF30B326BE6FDA2FB3FF585FDC2" box="[606,700,1206,1228]" gridcol="1" gridrow="17" pageId="2" pageNumber="516">0.8</td>
|
||
<td id="460F5DF30B326BE6FCEEFB3FF44CFDC2" box="[786,885,1206,1228]" gridcol="2" gridrow="17" pageId="2" pageNumber="516">0.8</td>
|
||
<td id="460F5DF30B326BE6FC4CFB3FF32AFDC2" box="[944,1043,1206,1228]" gridcol="3" gridrow="17" pageId="2" pageNumber="516">–</td>
|
||
<td id="460F5DF30B326BE6FBADFB3FF3F9FDC2" box="[1105,1216,1206,1228]" gridcol="4" gridrow="17" pageId="2" pageNumber="516">0.4</td>
|
||
<td id="460F5DF30B326BE6FAF4FB3FF252FDC2" box="[1288,1387,1206,1228]" gridcol="5" gridrow="17" pageId="2" pageNumber="516">0.4</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B32941BFF3BFA98F21DFC98" blockId="2.[151,1437,1297,1970]" pageId="2" pageNumber="516">
|
||
<emphasis id="899AEADF0B32941BFF3BFA98F6EAFC24" bold="true" box="[199,467,1297,1322]" pageId="2" pageNumber="516">Internal morphology.</emphasis>
|
||
Sensory organs, epidermis and body musculatures. Sensory pits, as simple invaginations, about 25 µm to 30 µm deep, contouring anterior tip, distributed ventromarginally in a single row (
|
||
<figureCitation id="23D52A480B32941BFF63FAD0F7C9FC7C" box="[159,240,1369,1394]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="2" pageNumber="516">Figs. 6</figureCitation>
|
||
̶7). They occur initially at intervals of about 24 µm, posteriorly becoming gradually sparser until they disappear approximately at
|
||
<quantity id="7C169B280B32941BFE31FAF4F537FC9B" box="[461,526,1405,1430]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="516" unit="mm" value="6.0">6 mm</quantity>
|
||
from anterior tip (approximately 14% of body length in the
|
||
<typeStatus id="6455886F0B32941BFB4EFAF4F220FC98" box="[1202,1305,1405,1430]" pageId="2" pageNumber="516" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B32941BFF3BFA28F4D0FFF0" blockId="2.[151,1437,1297,1970]" pageId="2" pageNumber="516">
|
||
Creeping sole occupying the whole body width in the
|
||
<typeStatus id="6455886F0B32941BFCAEFA28F48DFCB4" box="[850,948,1441,1466]" pageId="2" pageNumber="516" type="holotype">holotype</typeStatus>
|
||
(
|
||
<tableCitation id="F66C03760B32941BFC34FA28F318FCB4" box="[968,1057,1441,1466]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="2" pageNumber="516" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
). Three
|
||
<typeStatus id="6455886F0B32941BFB77FA2BF3FFFCB4" box="[1163,1222,1442,1466]" pageId="2" pageNumber="516">types</typeStatus>
|
||
of secretory cells discharge through dorsal epidermis and body margins of the pre-pharyngeal region: numerous rhabditogen cells with xanthophil secretion (rhammites) and cells with amorphous cyanophil secretion, as well as scarcer cells with coarse granular xanthophil secretion. Creeping sole receives the secretion from abundant coarse granular cyanophil glands and few rhabditogen cells with small, xanthophil rhabdites. Glandular margin (
|
||
<figureCitation id="23D52A480B32941BFB9EF9B8F3F4FF44" box="[1122,1229,1585,1610]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="2" pageNumber="516">Figs. 8–9</figureCitation>
|
||
) receiving at least four
|
||
<typeStatus id="6455886F0B32941BFF30F9DFF63EFF60" box="[204,263,1622,1646]" pageId="2" pageNumber="516">types</typeStatus>
|
||
of granular secretion: abundant glands with xanthophil coarse granules as well as numerous glands with ill-stained fine granules and scarcer xanthophil and cyanophil glands with fine granules. On anterior tip, numerous glands with cyanophil coarse granular secretion and scarcer cells with coarse granular xanthophil secretion open through the creeping sole, whereas few rhabditogen cells with small, xanthophil rhabdites and cells with amorphous cyanophil secretion open through the whole surface of the body.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B32941BFF3BF883F518FEBC" blockId="2.[151,1437,1297,1970]" pageId="2" pageNumber="516">
|
||
Cutaneous musculature with the usual three layers (circular, oblique and longitudinal layers), longitudinal layer with thick bundles (
|
||
<figureCitation id="23D52A480B32941BFE49F8A4F53AFE48" box="[437,515,1837,1862]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="2" pageNumber="516">Figs. 6</figureCitation>
|
||
̶11,
|
||
<tableCitation id="F66C03760B32941BFDC6F8A4F5A9FE48" box="[570,656,1837,1862]" captionStart="TABLE 2" captionStartId="6.[151,239,152,175]" captionTargetBox="[159,1243,266,515]" captionTargetPageId="6" captionText="TABLE 2. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc: h index) of specimens of Cratera steffeni sp. nov." httpUri="http://table.plazi.org/id/EF9166450B36941FFF6BFF11F517F9E1" pageId="2" pageNumber="516" tableUuid="EF9166450B36941FFF6BFF11F517F9E1">Table 2</tableCitation>
|
||
). Cutaneous musculature thinner in the pre-pharyngeal region than in the anterior region of the body, but gradually diminishing towards anterior tip. Musculature higher medially, becoming progressively lower towards body margins; ventral musculature higher than dorsal at the sagittal plane. Mc:h 9% in the
|
||
<typeStatus id="6455886F0B32941BFEB7F810F694FEBC" box="[331,429,1945,1970]" pageId="2" pageNumber="516" type="holotype">holotype</typeStatus>
|
||
(
|
||
<tableCitation id="F66C03760B32941BFE40F810F52AFEBC" box="[444,531,1945,1970]" captionStart="TABLE 2" captionStartId="6.[151,239,152,175]" captionTargetBox="[159,1243,266,515]" captionTargetPageId="6" captionText="TABLE 2. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc: h index) of specimens of Cratera steffeni sp. nov." httpUri="http://table.plazi.org/id/EF9166450B36941FFF6BFF11F517F9E1" pageId="2" pageNumber="516" tableUuid="EF9166450B36941FFF6BFF11F517F9E1">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="EF9166450B33941AFF6BF8FCF3D2FEE6" httpUri="https://zenodo.org/record/227063/files/figure.png" pageId="3" pageNumber="517" targetBox="[187,1350,219,1863]" targetPageId="3">
|
||
<paragraph id="BB5136CD0B33941AFF6BF8FCF3D2FEE6" blockId="3.[151,1436,1909,2024]" pageId="3" pageNumber="517">
|
||
<emphasis id="899AEADF0B33941AFF6BF8FCF60DFE85" bold="true" box="[151,308,1909,1932]" pageId="3" pageNumber="517">FIGURES 1–5</emphasis>
|
||
.
|
||
<taxonomicName id="7CEE4D4E0B33941AFEBEF8FEF6DBFE85" box="[322,482,1910,1932]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="517" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B33941AFEBEF8FEF6DBFE85" box="[322,482,1910,1932]" italics="true" pageId="3" pageNumber="517">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A957A40B33941AFE15F8FEF503FE82" box="[489,570,1911,1932]" pageId="3" pageNumber="517" rank="species">
|
||
<emphasis id="899AEADF0B33941AFE15F8FEF503FE82" bold="true" box="[489,570,1911,1932]" pageId="3" pageNumber="517">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: (1–2) colour pattern of a fixed specimen (paratype MZU PL.00178) in dorsal view; (3) eyes pattern of a fixed specimen (paratype MZU PL.00177) in dorsal view; (4) detail of eyes pattern in the anterior extremity of the holotype; (5) detail of the median third of the body of the holotype. Anterior tip to the left (Figs. 1–3) or to the top (Figs. 4–5). (ca) copulatory apparatus; (db) dorsal band; (e) eyes; (ph) pharynx; (ps) paramarginal stripes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EF9166450B34941DFF6BF968F5DEFEE3" httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="4" pageNumber="518" targetBox="[183,1387,193,1730]" targetPageId="4">
|
||
<paragraph id="BB5136CD0B34941DFF6BF968F5DEFEE3" blockId="4.[151,1436,1761,2029]" pageId="4" pageNumber="518">
|
||
<emphasis id="899AEADF0B34941DFF6BF968F673FFF9" bold="true" box="[151,330,1761,1784]" pageId="4" pageNumber="518">FIGURES 6–13.</emphasis>
|
||
<taxonomicName id="7CEE4D4E0B34941DFEAEF96BF6CCFFF9" box="[338,501,1762,1783]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="4" pageNumber="518" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B34941DFEAEF96BF6CCFFF9" box="[338,501,1762,1783]" italics="true" pageId="4" pageNumber="518">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="899AEADF0B34941DFE01F96BF561FFF9" bold="true" box="[509,600,1762,1783]" pageId="4" pageNumber="518">
|
||
<taxonomicNameLabel id="92A957A40B34941DFE01F96BF56AFFF9" box="[509,595,1762,1783]" pageId="4" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EF9166450B35941CFF6BF94CF55AFE58" httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="5" pageNumber="519" targetBox="[168,1436,193,1712]" targetPageId="5">
|
||
<paragraph id="BB5136CD0B35941CFF6BF94CF55AFE58" blockId="5.[151,1436,1733,1878]" pageId="5" pageNumber="519">
|
||
<emphasis id="899AEADF0B35941CFF6BF94CF674FFD5" bold="true" box="[151,333,1733,1756]" pageId="5" pageNumber="519">FIGURES 14–15</emphasis>
|
||
.
|
||
<taxonomicName id="7CEE4D4E0B35941CFEA5F94FF6C0FFD5" box="[345,505,1734,1755]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="519" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B35941CFEA5F94FF6C0FFD5" box="[345,505,1734,1755]" italics="true" pageId="5" pageNumber="519">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A957A40B35941CFE03F94FF576FFD5" box="[511,591,1734,1755]" pageId="5" pageNumber="519" rank="species">
|
||
<emphasis id="899AEADF0B35941CFE03F94FF576FFD5" bold="true" box="[511,591,1734,1755]" pageId="5" pageNumber="519">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EF9166450B36941FFF6BFF11F517F9E1" ID-Table-UUID="EF9166450B36941FFF6BFF11F517F9E1" httpUri="http://table.plazi.org/id/EF9166450B36941FFF6BFF11F517F9E1" pageId="6" pageNumber="520" targetBox="[159,1243,266,515]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="BB5136CD0B36941FFF6BFF11F517F9E1" blockId="6.[151,1435,152,239]" pageId="6" pageNumber="520">
|
||
<emphasis id="899AEADF0B36941FFF6BFF11F632F9A1" bold="true" box="[151,267,152,175]" pageId="6" pageNumber="520">TABLE 2.</emphasis>
|
||
Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of
|
||
<taxonomicName id="7CEE4D4E0B36941FFED5FF51F6E8F9E1" box="[297,465,215,239]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="6" pageNumber="520" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B36941FFED5FF51F6E8F9E1" box="[297,465,215,239]" italics="true" pageId="6" pageNumber="520">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A957A40B36941FFE24FF51F517F9E1" box="[472,558,216,239]" pageId="6" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BB5136CD0B36941FFF63FE83F3BCFB0D" pageId="6" pageNumber="520">
|
||
<table id="C9EEC46D0B366BE6FF63FE83F3E2FB0D" box="[159,1243,266,515]" gridcols="4" gridrows="5" pageId="6" pageNumber="520">
|
||
<tr id="05DE348F0B366BE6FF63FE83F3E2F850" box="[159,1243,266,350]" gridrow="0" pageId="6" pageNumber="520">
|
||
<th id="460F5DF30B366BE6FF63FE83F653F850" box="[159,362,266,350]" gridcol="0" gridrow="0" pageId="6" pageNumber="520">Specimens</th>
|
||
<th id="460F5DF30B366BE6FDE5FE83F54EF850" box="[537,631,266,350]" gridcol="1" gridrow="0" pageId="6" pageNumber="520">Holotype MZUSP PL.1557</th>
|
||
<th id="460F5DF30B366BE6FCB4FE83F492F850" box="[840,939,266,350]" gridcol="2" gridrow="0" pageId="6" pageNumber="520">Paratype MZU PL.00176</th>
|
||
<th id="460F5DF30B366BE6FB84FE83F3E2F850" box="[1144,1243,266,350]" gridcol="3" gridrow="0" pageId="6" pageNumber="520">Paratype MZU PL.00178</th>
|
||
</tr>
|
||
<tr id="05DE348F0B366BE6FF63FEF8F3E2F889" box="[159,1243,369,391]" gridrow="1" pageId="6" pageNumber="520">
|
||
<th id="460F5DF30B366BE6FF63FEF8F653F889" box="[159,362,369,391]" gridcol="0" gridrow="1" pageId="6" pageNumber="520">Dorsal musculature</th>
|
||
<td id="460F5DF30B366BE6FDE5FEF8F54EF889" box="[537,631,369,391]" gridcol="1" gridrow="1" pageId="6" pageNumber="520">52</td>
|
||
<td id="460F5DF30B366BE6FCB4FEF8F492F889" box="[840,939,369,391]" gridcol="2" gridrow="1" pageId="6" pageNumber="520">56</td>
|
||
<td id="460F5DF30B366BE6FB84FEF8F3E2F889" box="[1144,1243,369,391]" gridcol="3" gridrow="1" pageId="6" pageNumber="520">39</td>
|
||
</tr>
|
||
<tr id="05DE348F0B366BE6FF63FE13F3E2F8BE" box="[159,1243,410,432]" gridrow="2" pageId="6" pageNumber="520">
|
||
<th id="460F5DF30B366BE6FF63FE13F653F8BE" box="[159,362,410,432]" gridcol="0" gridrow="2" pageId="6" pageNumber="520">Ventral musculature</th>
|
||
<td id="460F5DF30B366BE6FDE5FE13F54EF8BE" box="[537,631,410,432]" gridcol="1" gridrow="2" pageId="6" pageNumber="520">65</td>
|
||
<td id="460F5DF30B366BE6FCB4FE13F492F8BE" box="[840,939,410,432]" gridcol="2" gridrow="2" pageId="6" pageNumber="520">46.5</td>
|
||
<td id="460F5DF30B366BE6FB84FE13F3E2F8BE" box="[1144,1243,410,432]" gridcol="3" gridrow="2" pageId="6" pageNumber="520">56</td>
|
||
</tr>
|
||
<tr id="05DE348F0B366BE6FF63FE4DF3E2F8D4" box="[159,1243,452,474]" gridrow="3" pageId="6" pageNumber="520">
|
||
<th id="460F5DF30B366BE6FF63FE4DF653F8D4" box="[159,362,452,474]" gridcol="0" gridrow="3" pageId="6" pageNumber="520">Body height</th>
|
||
<td id="460F5DF30B366BE6FDE5FE4DF54EF8D4" box="[537,631,452,474]" gridcol="1" gridrow="3" pageId="6" pageNumber="520">1240</td>
|
||
<td id="460F5DF30B366BE6FCB4FE4DF492F8D4" box="[840,939,452,474]" gridcol="2" gridrow="3" pageId="6" pageNumber="520">1029</td>
|
||
<td id="460F5DF30B366BE6FB84FE4DF3E2F8D4" box="[1144,1243,452,474]" gridcol="3" gridrow="3" pageId="6" pageNumber="520">1060</td>
|
||
</tr>
|
||
<tr id="05DE348F0B366BE6FF63FE64F3E2FB0D" box="[159,1243,493,515]" gridrow="4" pageId="6" pageNumber="520">
|
||
<th id="460F5DF30B366BE6FF63FE64F653FB0D" box="[159,362,493,515]" gridcol="0" gridrow="4" pageId="6" pageNumber="520">Mc:h (%)</th>
|
||
<td id="460F5DF30B366BE6FDE5FE64F54EFB0D" box="[537,631,493,515]" gridcol="1" gridrow="4" pageId="6" pageNumber="520">9</td>
|
||
<td id="460F5DF30B366BE6FCB4FE64F492FB0D" box="[840,939,493,515]" gridcol="2" gridrow="4" pageId="6" pageNumber="520">10</td>
|
||
<td id="460F5DF30B366BE6FB84FE64F3E2FB0D" box="[1144,1243,493,515]" gridcol="3" gridrow="4" pageId="6" pageNumber="520">9</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B36941FFF3BFDCEF5B7FA1B" blockId="6.[151,1437,583,2013]" pageId="6" pageNumber="520">
|
||
Mesenchymal musculature (
|
||
<figureCitation id="23D52A480B36941FFDF6FDCEF562FB6E" box="[522,603,583,608]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="6" pageNumber="520">Figs. 6</figureCitation>
|
||
̶8, 10̶11) well developed, mainly composed of three layers: (1) dorsal subcutaneous, located mainly close to the cutaneous musculature, with oblique fibers variously oriented (about 8–12 fibers thick); (2) supra-intestinal transverse (about 18̶20 fibers thick); (3) sub-intestinal transverse (about 6̶8 fibers thick). In addition, there are scattered transverse subneural fibers, ventral subcutaneous oblique fibers, as well as numerous dorsoventral ones. On the anterior region of the body, the mesenchymal musculature is more developed than in the pre-pharyngeal region.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B36941FFF3BFC96F344FDE6" blockId="6.[151,1437,583,2013]" pageId="6" pageNumber="520">
|
||
Pharynx. Pharynx bell-form,
|
||
<quantity id="7C169B280B36941FFDF0FCA9F558FA39" box="[524,609,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="520" unit="mm" value="2.8">2.8 mm</quantity>
|
||
in length, with dorsal insertion located at the beginning of median third of pharyngeal pouch; mouth posterior to the dorsal insertion, in the median third of pharyngeal pouch (
|
||
<figureCitation id="23D52A480B36941FFACBFCCDF2B6FA52" box="[1335,1423,836,861]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="6" pageNumber="520">Fig. 12</figureCitation>
|
||
). Esophagus short with folded wall; it is lined by cuboidal ciliated epithelium with insunk nuclei. The esophagus is coated with a thick circular subepithelial muscle layer, followed by longitudinal fibers, some of them mixed with the circular fibers (120 µm–180 µm thick). Esophagus: pharynx ratio, 10% in the
|
||
<typeStatus id="6455886F0B36941FFB6CFC26F3CFFAC6" box="[1168,1270,943,968]" pageId="6" pageNumber="520" type="holotype">holotype</typeStatus>
|
||
. Pharynx and pharyngeal lumen lined by ciliated cuboidal epithelium with insunk nuclei. Pharyngeal glands constituted by three secretory cell
|
||
<typeStatus id="6455886F0B36941FFEBCFC71F646FD1E" box="[320,383,1016,1040]" pageId="6" pageNumber="520">types</typeStatus>
|
||
: abundant secretory cells with fine granular erythrophil secretion; cells with amorphous cyanophil secretion; and cells with fine granular xanthophil secretion. Cell bodies of pharyngeal glands located in the mesenchyme, mainly anterior and laterally to pharynx. Pharyngeal outer musculature (30 µm – 40 µm thick) comprised of thin subepithelial layer of longitudinal muscles, followed by a thicker circular layer, mixed internally with numerous longitudinal fibers. Circular layer becomes as thin as longitudinal one towards pharyngeal tip. Inner pharyngeal musculature (100 µm–150 µm thick) comprises a thick circular subepithelial layer, mixed with longitudinal fibers. Inner musculature gradually becomes thinner towards pharyngeal tip.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B36941FFF3BFB7DF4A9FF5E" blockId="6.[151,1437,583,2013]" pageId="6" pageNumber="520">
|
||
Reproductive organs. Testes in one irregular row beneath the dorsal transverse mesenchymal muscles (
|
||
<figureCitation id="23D52A480B36941FFAB4FB7DF78FFC3E" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="6" pageNumber="520">Figs. 8, 10</figureCitation>
|
||
). They begin posteriorly to the ovaries and extend near to the root of the pharynx (
|
||
<tableCitation id="F66C03760B36941FFBAEFA9EF39CFC3E" box="[1106,1189,1303,1328]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="6" pageNumber="520" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
). Sperm ducts medial to ovovitelline ducts in pre-pharyngeal region. They form spermiducal vesicles posteriorly to pharynx. Distally, spermiducal vesicles enter laterally into the proximal expanded portion of the prostatic vesicle (
|
||
<figureCitation id="23D52A480B36941FFACFFAD6F2B6FC76" box="[1331,1423,1375,1400]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="6" pageNumber="520">Fig. 14</figureCitation>
|
||
). Extrabulbar prostatic vesicle, unpaired, located near to the common muscular coat, with ample proximal portion and tubular distal portion. The proximal portion shows a laterally expanded, T-shaped form (
|
||
<figureCitation id="23D52A480B36941FFB60FA2EF3C0FCCE" box="[1180,1273,1447,1472]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="6" pageNumber="520">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="23D52A480B36941FFAF8FA21F21BFCCE" box="[1284,1314,1448,1472]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1832,1855]" captionTargetBox="[151,1436,337,1810]" captionTargetId="figure@7.[151,1436,337,1811]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." httpUri="https://zenodo.org/record/227066/files/figure.png" pageId="6" pageNumber="520">17</figureCitation>
|
||
,
|
||
<figureCitation id="23D52A480B36941FFAD1FA21F273FCCE" box="[1325,1354,1448,1472]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="6" pageNumber="520">18</figureCitation>
|
||
) that is closer to the ventral epidermis than to the dorsal epidermis. Τhe prostatic vesicle penetrates the common muscular coat, becoming sinuous, and continues, inside the penis papilla, as an ejaculatory duct. Ejaculatory duct almost straight, opening through an expansion into the tip of the penis papilla (
|
||
<figureCitation id="23D52A480B36941FFC3BF99DF322FF22" box="[967,1051,1556,1581]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="6" pageNumber="520">Fig. 19</figureCitation>
|
||
). Male atrium unfolded, occupied by the conical and symmetrical penis papilla (
|
||
<tableCitation id="F66C03760B36941FFD65F9BEF5D4FF5E" box="[665,749,1591,1616]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="6" pageNumber="520" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
,
|
||
<figureCitation id="23D52A480B36941FFD07F9BEF46CFF5E" box="[763,853,1591,1616]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="6" pageNumber="520">Figs. 14</figureCitation>
|
||
̶17).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B36941FFF3BF9D5F602FE7E" blockId="6.[151,1437,583,2013]" pageId="6" pageNumber="520">
|
||
Lining epithelium of sperm ducts cuboidal and ciliated; thin muscularis (3 µm) mainly constituted of circular fibers. Prostatic vesicle lined with ciliated tall columnar or pseudo-stratified epithelium, receiving abundant fine granular erythrophil and amorphous cyanophil secretions, both from secretory cells with bodies lying in mesenchyme, mainly around vesicle. Muscularis of prostatic vesicle (about 40 µm–80 µm thick) comprises interwoven longitudinal, oblique and circular fibers (
|
||
<figureCitation id="23D52A480B36941FFD09F965F4B9FE0B" box="[757,896,1772,1797]" captionStart-0="FIGURES 16 – 17" captionStart-1="FIGURES 18 – 23" captionStartId-0="7.[151,264,1832,1855]" captionStartId-1="8.[151,264,1021,1044]" captionTargetBox-0="[151,1436,337,1810]" captionTargetBox-1="[151,1436,193,999]" captionTargetId-0="figure@7.[151,1436,337,1811]" captionTargetId-1="figure@8.[151,1436,193,1000]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." captionText-1="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri-0="https://zenodo.org/record/227066/files/figure.png" httpUri-1="https://zenodo.org/record/227067/files/figure.png" pageId="6" pageNumber="520">Figs. 16–18</figureCitation>
|
||
). Ejaculatory duct lined with ciliated columnar epithelium, receiving few openings from secretory cells with weakly cyanophil, amorphous secretion. These glands show intrapapillar cell bodies. Muscle coat of ejaculatory duct thin (5 µm–10 µm thick), mainly constituted of circular fibers.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B36941EFF3BF8F5F67FF813" blockId="6.[151,1437,583,2013]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="521" pageId="6" pageNumber="520">
|
||
Penis papilla lined with non-ciliated columnar epithelium with apical xanthophil layer. The epithelium becomes flat towards the tip of the papilla (
|
||
<figureCitation id="23D52A480B36941FFD5EF829F417FEB7" box="[674,814,1952,1977]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1832,1855]" captionTargetBox="[151,1436,337,1810]" captionTargetId="figure@7.[151,1436,337,1811]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." httpUri="https://zenodo.org/record/227066/files/figure.png" pageId="6" pageNumber="520">Figs. 16–17</figureCitation>
|
||
,
|
||
<figureCitation id="23D52A480B36941FFCC2F829F463FEB6" box="[830,858,1952,1976]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="6" pageNumber="520">19</figureCitation>
|
||
). Penis glands with fine granular xanthophil and erythrophil secretions as well as cyanophil amorphous secretion. They run longitudinally in the papilla, with numerous openings into the male atrium through its lining epithelium (
|
||
<figureCitation id="23D52A480B37941EFC33FF1EF36EF9BE" box="[975,1111,151,176]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1832,1855]" captionTargetBox="[151,1436,337,1810]" captionTargetId="figure@7.[151,1436,337,1811]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." httpUri="https://zenodo.org/record/227066/files/figure.png" pageId="7" pageNumber="521">Figs. 16–17</figureCitation>
|
||
). Erythrophil and cyanophil glands present cell bodies external to common muscle coat among fibers of this coat; xanthophil cells, intrapapillar cells bodies. Muscularis of the penis papilla (25 µm) composed of a subepithelial circular layer and a longitudinal subjacent layer.
|
||
</paragraph>
|
||
<caption id="EF9166450B37941EFF6BF8A1F2A5FE94" httpUri="https://zenodo.org/record/227066/files/figure.png" pageId="7" pageNumber="521" targetBox="[151,1436,337,1810]" targetPageId="7">
|
||
<paragraph id="BB5136CD0B37941EFF6BF8A1F2A5FE94" blockId="7.[151,1436,1832,1946]" pageId="7" pageNumber="521">
|
||
<emphasis id="899AEADF0B37941EFF6BF8A1F668FE33" bold="true" box="[151,337,1832,1855]" pageId="7" pageNumber="521">FIGURES 16–17</emphasis>
|
||
.
|
||
<taxonomicName id="7CEE4D4E0B37941EFE9DF8A0F53DFE30" box="[353,516,1833,1854]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="521" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B37941EFE9DF8A0F53DFE30" box="[353,516,1833,1854]" italics="true" pageId="7" pageNumber="521">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="899AEADF0B37941EFDF2F8A0F55BFE30" bold="true" box="[526,610,1833,1854]" pageId="7" pageNumber="521">
|
||
<taxonomicNameLabel id="92A957A40B37941EFDF2F8A0F55BFE30" box="[526,610,1833,1854]" pageId="7" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL.00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EF9166450B389411FF6BFC74F46BFDC5" httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="8" pageNumber="522" targetBox="[151,1436,193,999]" targetPageId="8">
|
||
<paragraph id="BB5136CD0B389411FF6BFC74F46BFDC5" blockId="8.[151,1436,1021,1227]" pageId="8" pageNumber="522">
|
||
<emphasis id="899AEADF0B389411FF6BFC74F676FD1D" bold="true" box="[151,335,1021,1044]" pageId="8" pageNumber="522">FIGURES 18–23</emphasis>
|
||
.
|
||
<taxonomicName id="7CEE4D4E0B389411FEA0FC77F6C4FD1D" box="[348,509,1022,1043]" class="Turbellaria" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="522" phylum="Platyhelminthes" rank="species" species="steffeni" status="sp. nov.">
|
||
<emphasis id="899AEADF0B389411FEA0FC77F6C4FD1D" box="[348,509,1022,1043]" italics="true" pageId="8" pageNumber="522">Cratera steffeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A957A40B389411FDF8FC77F56CFD1D" box="[516,597,1022,1043]" pageId="8" pageNumber="522" rank="species">
|
||
<emphasis id="899AEADF0B389411FDF8FC77F56CFD1D" bold="true" box="[516,597,1022,1043]" pageId="8" pageNumber="522">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: (18) detail of prostatic vesicle of paratype MZU PL.00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL.00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19–20; 22–23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BB5136CD0B389411FF3BFB71F4D9FCCB" blockId="8.[151,1437,1272,2017]" pageId="8" pageNumber="522">
|
||
Epithelial lining of male atrium columnar (20 µm–30 µm), non-ciliated. Epithelial cells with xanthophil apical secretion, higher in the distal portion of the male atrium. Four
|
||
<typeStatus id="6455886F0B389411FC87FA94F48FFC3B" box="[891,950,1309,1333]" pageId="8" pageNumber="522">types</typeStatus>
|
||
of glands empty through this epithelium: abundant cells with fine granular, cyanophil secretion; cells with fine granular erythrophil secretion; cells with cyanophil amorphous secretion; cells with fine granular xanthophil secretion. Numerous necks of cyanophil glands concentrate their openings through the dorsal wall of the male atrium (
|
||
<figureCitation id="23D52A480B389411FC57FA01F4C6FCAF" box="[939,1023,1416,1441]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="8" pageNumber="522">Fig. 20</figureCitation>
|
||
). Muscularis of male atrium (15 µm) comprised of circular subepithelial fibers and subjacent longitudinal fibers.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B389411FF3BFA59F48DFFCF" blockId="8.[151,1437,1272,2017]" pageId="8" pageNumber="522">
|
||
Vitellaria, situated between intestinal branches, open into the ovovitelline ducts. Ovaries ovoid, measuring about 250 µm in diameter in the
|
||
<typeStatus id="6455886F0B389411FDE0FA7DF5B8FF03" box="[540,641,1524,1549]" pageId="8" pageNumber="522" type="holotype">holotype</typeStatus>
|
||
. Ovovitelline ducts emerge dorso-laterally from the posterior half of ovaries (
|
||
<figureCitation id="23D52A480B389411FF0AF991F67FFF3F" box="[246,326,1560,1585]" captionStart="FIGURES 6 – 13" captionStartId="4.[151,264,1761,1784]" captionTargetBox="[183,1387,193,1730]" captionTargetId="figure@4.[183,1388,193,1738]" captionTargetPageId="4" captionText="FIGURES 6 – 13. Cratera steffeni sp. nov., holotype: (6) anterior region of the body in transverse section; (7) detail of the anterior region of the body in transverse section; (8) pre-pharyngeal region in transverse section; (9) detail of body margin of the pre-pharyngeal region in transverse section; (10) detail of the dorsal surface of the pre-pharyngeal region in transverse section; (11) detail of the ventral surface of the pre-pharyngeal region in transverse section; (12) pharynx in sagittal section (anterior tip to the left); (13) ovary in sagittal section (anterior tip to the left). (de) dorsal epidermis; (di) dorsal insertion; (dm) dorsal cutaneous musculature; (e) eyes; (es) esophagus; (gm) glandular margin; (i) intestine; (im) internal musculature; (lu) pharyngeal lumen; (m) mouth; (mm) mesenchymal muscles; (n) nerve cord; (o) ovary; (om) outer musculature; (ov) ovovitelline duct; (pp) pharyngeal pouch; (sc) secretory cells; (sp) sensory pit; (t) testes; (v) vitellaria; (ve) ventral epidermis; (vi) ventral insertion; (vm) ventral cutaneous musculature." httpUri="https://zenodo.org/record/227064/files/figure.png" pageId="8" pageNumber="522">Fig. 13</figureCitation>
|
||
) and run posteriorly immediately above the nerve plate. Laterally to the female atrium, ovovitelline ducts ascend posteriorly and medially inclined, to unite dorsally to the ental part of female atrium, thus forming a short common glandular ovovitelline duct. Ental portion of female atrium presents a long, dorso-anteriorly curved diverticulum (vagina) (
|
||
<figureCitation id="23D52A480B389411FE66F90DF51DFF93" box="[410,548,1668,1693]" captionStart-0="FIGURES 14 – 15" captionStart-1="FIGURES 16 – 17" captionStart-2="FIGURES 18 – 23" captionStartId-0="5.[151,264,1733,1756]" captionStartId-1="7.[151,264,1832,1855]" captionStartId-2="8.[151,264,1021,1044]" captionTargetBox-0="[168,1436,193,1712]" captionTargetBox-1="[151,1436,337,1810]" captionTargetBox-2="[151,1436,193,999]" captionTargetId-0="figure@5.[151,1436,193,1712]" captionTargetId-1="figure@7.[151,1436,337,1811]" captionTargetId-2="figure@8.[151,1436,193,1000]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." captionText-1="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." captionText-2="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri-0="https://zenodo.org/record/227065/files/figure.png" httpUri-1="https://zenodo.org/record/227066/files/figure.png" httpUri-2="https://zenodo.org/record/227067/files/figure.png" pageId="8" pageNumber="522">Figs. 14–22</figureCitation>
|
||
). Female atrium funnel-shaped with folded walls (
|
||
<figureCitation id="23D52A480B389411FBA0F90DF38EFF93" box="[1116,1207,1668,1693]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="8" pageNumber="522">Figs. 14</figureCitation>
|
||
̶17). Female atrium length almost the same as male atrium length in the
|
||
<typeStatus id="6455886F0B389411FD22F921F479FFCF" box="[734,832,1704,1729]" pageId="8" pageNumber="522" type="holotype">holotype</typeStatus>
|
||
(
|
||
<tableCitation id="F66C03760B389411FCB3F921F49BFFCF" box="[847,930,1704,1729]" captionStart="TABLE 1" captionStartId="2.[151,239,296,319]" captionTargetBox="[156,1387,442,1228]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera steffeni sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/EF9166450B32941BFF6BFEA1F52BF891" pageId="8" pageNumber="522" tableUuid="EF9166450B32941BFF6BFEA1F52BF891">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B389411FF3BF945F41EFE5F" blockId="8.[151,1437,1272,2017]" pageId="8" pageNumber="522">
|
||
Paired ovovitelline ducts and common oviduct lined with ciliated, cuboidal to columnar epithelium, and coated with muscularis comprising mixed circular and longitudinal muscle fibers (5 Μm – 10 Μm thick). Abundant shell glands with xanthophil secretion empty into almost the whole ascending portion of paired ovovitelline ducts and common glandular ovovitelline duct (
|
||
<figureCitation id="23D52A480B389411FDC1F8B1F5FDFE5F" box="[573,708,1848,1873]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="8" pageNumber="522">Figs. 14, 15</figureCitation>
|
||
,
|
||
<figureCitation id="23D52A480B389411FD2CF8B0F422FE5F" box="[720,795,1849,1873]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="8" pageNumber="522">21–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B389410FF3BF8D5F545F9F6" blockId="8.[151,1437,1272,2017]" lastBlockId="9.[151,1436,151,645]" lastPageId="9" lastPageNumber="523" pageId="8" pageNumber="522">
|
||
Female atrium lined by tall columnar epithelium exhibiting irregular height and multilayered aspect in some places (
|
||
<figureCitation id="23D52A480B389411FF12F809F677FE97" box="[238,334,1920,1945]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1832,1855]" captionTargetBox="[151,1436,337,1810]" captionTargetId="figure@7.[151,1436,337,1811]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Cratera steffeni sp. nov.: (16) copulatory apparatus of the holotype in sagittal section; (17) copulatory apparatus of paratype MZU PL. 00176 in horizontal section. Anterior tip to the left. The arrow indicates the approximate position of the common muscle coat. (cmc) common muscle coat; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (va) vagina." httpUri="https://zenodo.org/record/227066/files/figure.png" pageId="8" pageNumber="522">Figs. 16</figureCitation>
|
||
,
|
||
<figureCitation id="23D52A480B389411FEA0F808F642FE97" box="[348,379,1921,1945]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1021,1044]" captionTargetBox="[151,1436,193,999]" captionTargetId="figure@8.[151,1436,193,1000]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Cratera steffeni sp. nov.: (18) detail of prostatic vesicle of paratype MZU PL. 00176 in horizontal section; (19) detail of penis papilla of the holotype in sagittal section; (20) detail of male atrium of the holotype in sagittal section; (21) detail of vagina and ovovitelline ducts in paratype MZU PL. 00176 in horizontal section; (22) detail of vagina and common ovovitelline duct of the holotype in sagittal section; (23) detail of female atrium of the holotype in sagittal section. Anterior tip to the left (Figs. 19 – 20; 22 – 23) or to the bottom (Figs. 18 and 21). (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (v) vitellaria; (va) vagina." httpUri="https://zenodo.org/record/227067/files/figure.png" pageId="8" pageNumber="522">23</figureCitation>
|
||
); with xanthophil apical surface. Vagina lined with columnar to pseudostratified epithelium, ciliated in its proximal portion. Vagina and female atrium receive abundant erythrophil and xanthophil granular secretions as well as cyanophil amorphous secretion. Cell bodies of cyanophil and erythrophil glands are located between fibers of the atrial stroma or external to the common muscle coat and those of xanthophil cells are internal to the common muscle coat. Muscularis of vagina and female atrium (20 µm thick) composed mainly of circular fibers mixed with some longitudinal fibers.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B399410FF3BFE8DF6E7F8A3" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="523">
|
||
Gonopore canal vertical at the sagittal plane. Male and female atria with ample communication, without separating folds (
|
||
<figureCitation id="23D52A480B399410FEA6FEAEF68FF84E" box="[346,438,295,320]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1733,1756]" captionTargetBox="[168,1436,193,1712]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Cratera steffeni sp. nov.: (14) sagittal composite reconstruction of the copulatory apparatus of the holotype; (15) diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL. 00176. Anterior tip to the left. (cmc) common muscle coat; (cov) common glandular ovovitelline duct; (ej) ejaculatory duct; (fa) female atrium; (go) gonopore; (ma) male atrium; (ov) ovovitelline ducts; (p) penis papilla; (pv) prostatic vesicle; (sc) secretory cells; (sg) shell glands; (sv) spermiducal vesicle; (va) vagina." httpUri="https://zenodo.org/record/227065/files/figure.png" pageId="9" pageNumber="523">Figs. 14</figureCitation>
|
||
̶17). Gonopore canal lined with ciliated columnar epithelium, receiving the openings of abundant rhabditogen cells and scarcer glands containing amorphous cyanophil secretion as well as glands with fine granular erythrophil secretion. Muscularis of gonopore canal comprised of circular subepithelial fibers and subjacent longitudinal fibers.
|
||
</paragraph>
|
||
<paragraph id="BB5136CD0B399410FF3BFE31F6B4FB19" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="523">Common muscle coat with circular, longitudinal and oblique fibers, thickest entally and along dorsal wall of male, thinner around female atrium. A stroma with sparse mixed muscle fibers separates the atrial muscularis and common muscle coat.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3F465460B399410FF3BFDADF274FB8B" pageId="9" pageNumber="523" type="discussion">
|
||
<paragraph id="BB5136CD0B399410FF3BFDADF274FB8B" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="523">
|
||
<emphasis id="899AEADF0B399410FF3BFDADF602FB33" bold="true" box="[199,315,548,573]" pageId="9" pageNumber="523">Remarks.</emphasis>
|
||
Vitellaria well-developed in the
|
||
<typeStatus id="6455886F0B399410FD57FDADF434FB33" box="[683,781,548,573]" pageId="9" pageNumber="523" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="6455886F0B399410FCB8FDACF496FB33" box="[836,943,549,573]" pageId="9" pageNumber="523" type="paratype">paratypes</typeStatus>
|
||
MZU PL.00176 e MZU PL.00179. The not fully mature
|
||
<typeStatus id="6455886F0B399410FED6FDC1F6B3FB6E" box="[298,394,584,608]" pageId="9" pageNumber="523" type="paratype">paratype</typeStatus>
|
||
MZU PL.00178 shows the dorsal insertion of the pharynx located at the end of the anterior third of the pharyngeal pouch, which is thus anteriad-shifted in relation to the
|
||
<typeStatus id="6455886F0B399410FBFDFDE5F35AFB8B" box="[1025,1123,620,645]" pageId="9" pageNumber="523" type="holotype">holotype</typeStatus>
|
||
and other
|
||
<typeStatus id="6455886F0B399410FB26FDE4F270FB8B" box="[1242,1353,621,645]" pageId="9" pageNumber="523" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |