treatments-xml/data/21/F2/EC/21F2EC0875F45665932126BCA413C50A.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="21F2EC0875F45665932126BCA413C50A" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila bidentata Pall-Gergely &amp; Jochum 2023, sp. nov." docType="treatment" docUuid="485E89E4-1CF2-48E2-B0BC-2EDE8440F451" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:485E89E4-1CF2-48E2-B0BC-2EDE8440F451" httpUri="http://treatment.plazi.org/id/21F2EC0875F45665932126BCA413C50A" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/485E89E4-1CF2-48E2-B0BC-2EDE8440F451" authority="Páll-Gergely &amp; Jochum" authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata" status="sp. nov.">
Angustopila bidentata 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Jochum
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Angustopila bidentata Pall-Gergely &amp; Jochum, sp. nov. (holotype, MNHN-IM- 2014 - 6411). Apertural (A), ventral (B), apical (C) and lateral (D) sides of the shell; sculpture on the protoconch (E), aperture (F); ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure63" httpUri="https://binary.pensoft.net/fig/808872" pageId="0" pageNumber="1">Fig. 63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Laos • 1 empty shell (H: 0.88 mm, D: 0.8 mm); South-Central Laos, Khammouane Province, ca. 35 km ENE of Thakhek (Muang Khammouane), ca. 7 km WNW of Mahaxai, on and under rocks in dry secondary forest under S exposed cliff (locality code: 4L07); 
<geoCoordinate degrees="17" direction="north" minutes="26.74" orientation="latitude" precision="9" value="17.445667">17°26.74'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="08.36" orientation="longitude" precision="9" value="105.139336">105°08.36'E</geoCoordinate>
; 25 Nov. 2007; A. Abdou &amp; I.V. Muratov leg.; MNHN-IM-2014-6411.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Laos • 3 shells; same data as for holotype; MNHN-IM-2014-6412 • 2 shells; South-Central Laos, Khammouane Province, ca. 15 km NE of Thakhek (Muang Khammouane), ca. 12.5 km SE of Ban Nase, on and under rocks in dry secondary forest near large flooded cave under W exposed cliff (locality code: 7L07); 
<geoCoordinate degrees="17" direction="north" minutes="30.55" orientation="latitude" precision="9" value="17.509167">17°30.55'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="53.44" orientation="longitude" precision="9" value="104.89066">104°53.44'E</geoCoordinate>
; 130 m a.s.l.; 28 Nov. 2007; A. Abdou &amp; I.V. Muratov leg.; MNHN-IM-2014-6410 • 12 shells (some are damaged); South-Central Laos, Khammouane Province, ca. 37 km ENE of Thakhek (Muang Khammouane), ca. 4.5 km WNW of Mahaxai, on and under rocks in dry secondary forest under E exposed cliff (locality code: 3L07); 
<geoCoordinate degrees="17" direction="north" minutes="25.96" orientation="latitude" precision="9" value="17.432667">17°25.96'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="09.67" orientation="longitude" precision="9" value="105.16116">105°09.67'E</geoCoordinate>
; 150 m a.s.l.; 25 Nov. 2007; A. Abdou &amp; I.V. Muratov leg.; MNHN-IM-2014-6409 • 10 adult shells; Khammouane Province, 17.5 km from centre of Thakhek towards Mahaxay, Tham Nang Aen (cave) (locality code: 2019/96); 
<geoCoordinate degrees="17" direction="north" minutes="26.65" orientation="latitude" precision="9" value="17.444166">17°26.65'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="57.02" orientation="longitude" precision="9" value="104.95033">104°57.02'E</geoCoordinate>
; 190 m a.s.l.; 27 Sep. 2019; A. Hunyadi leg.; coll. HA • 5 shells; South-Central Laos, Khammouane Province, ca. 10.5 km E of Thakhek (Muang Khammouane), on and under rocks, cave deposits, in secondary forest under entrance and in large cave on exposed NE steep slope (locality code: 25L07); 
<geoCoordinate degrees="17" direction="north" minutes="24.34" orientation="latitude" precision="9" value="17.405666">17°24.34'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="54.89" orientation="longitude" precision="9" value="104.91483">104°54.89'E</geoCoordinate>
; 160 m a.s.l.; 9 Dec. 2007; A. Abdou &amp; I.V. Muratov leg.; MNHN-IM-2014-6408 • 1 damaged shell; Khammouane Province, Tham Nam 
<normalizedToken originalValue="Dôn">Don</normalizedToken>
Cave, sediment deposited by temporary side rivulet at passage entrance (locality code: JG2A); 
<geoCoordinate degrees="17" direction="north" minutes="33.82" orientation="latitude" precision="9" value="17.563667">17°33.82'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="52.30" orientation="longitude" precision="9" value="104.871666">104°52.30'E</geoCoordinate>
; 160 m a.s.l.; 11 Feb. 2017; J. Grego leg.; coll. JG • 1 shell; Khammouane Province, Tham Nam 
<normalizedToken originalValue="Dôn">Don</normalizedToken>
Cave, Earthquake Dome, sand sediments at bank of river in cave (locality code: JG2B); 
<geoCoordinate degrees="17" direction="north" minutes="33.82" orientation="latitude" precision="9" value="17.563667">17°33.82'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="52.30" orientation="longitude" precision="9" value="104.871666">104°52.30'E</geoCoordinate>
; 160 m a.s.l.; 11 Feb. 2017; J. Grego leg.; coll. JG • 1 shell; Khammouane Province, Tham Pha Inh (cave), E Thakhek, at the base of limestone rocks (Locality code: La.7); 
<geoCoordinate degrees="17" direction="north" minutes="27.69" orientation="latitude" precision="9" value="17.4615">17°27.69'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="54.95" orientation="longitude" precision="9" value="104.91583">104°54.95'E</geoCoordinate>
; 180 m a.s.l.; Mar. 2010; A. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
leg.; coll. RE • 1 shell; Khammouane Province, 40 km from centre of Thakhek towards Mahaxay, 2.3 km southeast from Ban Na Coc, rock wall (locality code: 2019/95); 
<geoCoordinate degrees="17" direction="north" minutes="25.96" orientation="latitude" precision="9" value="17.432667">17°25.96'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="09.67" orientation="longitude" precision="9" value="105.16116">105°09.67'E</geoCoordinate>
; 155 m a.s.l.; 29 Sep. 2019; A. Hunyadi leg.; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2017-02-11" collectingDateMax="2019-09-27" collectingDateMin="2017-02-11" collectorName="J. Grego, Thakhek, Mahaxay, Tham Nang Aen, A. Hunyadi" country="Laos" elevation="160" latitude="17.444166" location="Tham Nam Don Cave" longLatPrecision="13" longitude="104.95033" specimenCount="1" stateProvince="Khammouane Province" typeStatus="paratype">
<collectingCountry name="Laos">Laos</collectingCountry>
• 1 damaged shell with missing aperture (identity questionable, not 
<typeStatus>paratype</typeStatus>
, much smaller than other shell from the same locality, see remarks); 
<collectingRegion country="Laos" name="Khammouan">Khammouane Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:21F2EC0875F45665932126BCA413C50A:FC1B989E77A7DEA9BA3E3B70D4D9A678" country="Laos" latitude="17.444166" longLatPrecision="13" longitude="104.95033" name="Tham Nam Don Cave" stateProvince="Khammouane Province">
Tham Nam 
<normalizedToken originalValue="Dôn">Don</normalizedToken>
Cave
</location>
, sediment deposited by temporary side rivulet at passage entrance (locality code: JG2A); 
<geoCoordinate degrees="17" direction="north" minutes="33.82" orientation="latitude" precision="9" value="17.563667">17°33.82'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="52.30" orientation="longitude" precision="9" value="104.871666">104°52.30'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2017-02-11">11 Feb. 2017</collectingDate>
; 
<collectorName>J. Grego</collectorName>
leg.; coll. JG • 6 j/b shells; 
<collectingRegion country="Laos" name="Khammouan">Khammouane Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.75" unit="km" value="17.5">17.5 km</quantity>
from centre of 
<collectorName>Thakhek</collectorName>
towards 
<collectorName>Mahaxay</collectorName>
, 
<collectorName>Tham Nang Aen</collectorName>
(cave) (locality code: 2019/96); 
<geoCoordinate degrees="17" direction="north" minutes="26.65" orientation="latitude" precision="9" value="17.444166">17°26.65'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="57.02" orientation="longitude" precision="9" value="104.95033">104°57.02'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-09-27">27 Sep. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small to large 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a very wide sinulus, an oblique aperture, a very narrow umbilicus, and strong parietal and upper palatal teeth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell small to large for the genus (mostly medium sized), higher than wide, or rarely slightly wider than high; pale grey, conical-globular, last or penultimate whorl widest from standard apertural view; protoconch consists of 1.5 whorls, with fine spiral striation on the entire protoconch (five spiral lines visible); teleoconch finely ornamented with irregularly spaced radial growth lines crossed by fine rows of regularly or irregularly spaced spiral striae (ca. 13-19 on body whorl from apertural view); on frontal and ventral surface of body whorl spiral and radial lines of comparable strength, or spiral ones dominant; whorls 4.5, slightly shouldered; aperture very strongly oblique to shell axis from lateral view; umbilicus, very narrow; aperture reniform, sinulus wide and strongly separated by parietal and palatal teeth; peristome slightly expanded, not reflected; mid-section comprising the parietal tooth sinuous and slightly protruding (in side view); aperture detached from the penultimate whorl forming a short, downward-directed tubus (lateral view); parietal callus strongly elevated, sharp; parietal tooth strongly developed and pointed towards the palatal tooth, which is situated high up on the palatal side; parietal and palatal teeth are directly opposite to each other.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.8-1.1, D = 0.75-0.95, H/D*100 = 98.8-115.4 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 8, from populations excluding JG2A), RUD = 18.7-25.8 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 10); H: 1.21, D = 1.09, H/D*100 = 111 (JG2A).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bidentata</emphasis>
</taxonomicName>
sp. nov. can be distinguished from all other 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species by the presence of two apertural denticles on a semi-detached tuba and the strongly oblique, wing-shaped aperture formed by the very wide sinulus and a very narrow umbilicus. The latter two traits allow the distinction of this species from congeners even at the juvenile stage. The most similar species is probably 
<taxonomicName lsidName="A. dominikae" pageId="0" pageNumber="1" rank="species" species="dominikae">
<emphasis italics="true" pageId="0" pageNumber="1">A. dominikae</emphasis>
</taxonomicName>
, which has an oblique, oblong, and adnate aperture, and a more globular shell. See also under 
<taxonomicName lsidName="A. erawanica" pageId="0" pageNumber="1" rank="species" species="erawanica">
<emphasis italics="true" pageId="0" pageNumber="1">A. erawanica</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="A. huoyani" pageId="0" pageNumber="1" rank="species" species="huoyani">
<emphasis italics="true" pageId="0" pageNumber="1">A. huoyani</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="A. tamlod" pageId="0" pageNumber="1" rank="species" species="tamlod">
<emphasis italics="true" pageId="0" pageNumber="1">A. tamlod</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet 
<taxonomicName authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bidentata</emphasis>
</taxonomicName>
(Latin, meaning two-teeth) refers to the two strongly developed apertural teeth of this new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bidentata</emphasis>
</taxonomicName>
sp. nov. is known from Khammouane Province of Laos, in the limestone area east of Thakhek (Fig. 
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Distribution of Angustopila species in Laos." figureDoi="10.3897/zookeys.1147.93824.figure64" httpUri="https://binary.pensoft.net/fig/808873" pageId="0" pageNumber="1">64</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure63" httpUri="https://binary.pensoft.net/fig/808872" pageId="0" pageNumber="1" start="Figure 63" startId="F63">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 63.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Jochum" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bidentata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bidentata</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Jochum, sp. nov. (holotype, MNHN-IM-2014-6411). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and lateral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure64" httpUri="https://binary.pensoft.net/fig/808873" pageId="0" pageNumber="1" start="Figure 64" startId="F64">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 64.</emphasis>
Distribution of 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species in Laos.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species shows a considerable variability in terms of shell size and some conchological characters. The populations JG2B, 3L07, 4L07, and 7L07 are indistinguishable from each other. Shells of population 25L07 are slightly larger than those in the other populations and bear a more elongated (less rounded) aperture, denser spiral striae, and show stronger radial sculpture. The single shell of the sample JG2A is very large compared to the others, and has a rounded aperture, similar to shells of the four aforementioned populations. The single shell from locality JG2A was already cracked and was subsequently damaged during imaging. However, other than shell size, no obvious morphological differences warrant distinction of that large specimen from the remaining 
<taxonomicName lsidName="A. bidentata" pageId="0" pageNumber="1" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">A. bidentata</emphasis>
</taxonomicName>
sp. nov. shells. A single, toothless juvenile shell, similar in size to typical 
<taxonomicName lsidName="A. bidentata" pageId="0" pageNumber="1" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">A. bidentata</emphasis>
</taxonomicName>
sp. nov., was also found in the same locality (JG2A). If that shell was an 
<taxonomicName lsidName="A. bidentata" pageId="0" pageNumber="1" rank="species" species="bidentata">
<emphasis italics="true" pageId="0" pageNumber="1">A. bidentata</emphasis>
</taxonomicName>
sp. nov. individual, then the large shell of the same locality represents a distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>